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1.
In tropical regions where agricultural activities are limited, agroforestry is an alternative for both economic development and for the management and conservation of biodiversity. The potential role of different types of land use as reservoirs of dung beetle diversity in the wet tropical forest of the Pacific lowlands of Colombia is evaluated in three agroforestry systems that differ in canopy cover and the sowing density of Borojoa patinoi crops associated with timber forest. Although total species richness was similar among land use types, differences related to the decrease in the abundance and biomass of the species were remarkable, and reflected in turn by the diversity and structure of the guilds. The general pattern observed was one in which the structure of the dung beetle assemblage of B. patinoi growing below a diversified and permanent tree cover was similar to that of the primary and secondary forest. Beetle diversity in management systems with less tree cover or a high sowing density of B. patinoi was lower and very similar to that of abandoned agricultural fields. This suggests that B. patinoi agroforestry systems can be viewed as valuable instruments for biodiversity management and conservation in the wet tropical forests of the Pacific lowlands and not just as substitutes for forest, though we must be aware that structural changes in the beetle community may in turn affect the ecological processes regulated by these insects in the agrosystems under study.  相似文献   

2.
Small seasonally flooded forest ponds have received increased attention due to a growing recognition of their abundance in many landscapes, their importance as habitat for a variety of organisms, and the contributions they make to species and ecosystem diversity. There also is concern over potential negative effects of forest management in adjacent uplands on seasonal pond ecology. Several studies have examined invertebrate and songbird responses to upland harvest around seasonal ponds. Less attention has been given to examining how seasonal pond plant communities respond to adjacent forest harvesting. We studied the response of seasonal pond plant communities to adjacent upland timber harvests, assessing whether buffers around ponds (15.25 m uncut and partially cut) mitigated changes in species abundance and community composition, relative to changes in ponds that were clearcut to the pond margin. We addressed our objective using an operational-scale experiment in northern Minnesota, which included pre-harvest sampling, replicated treatments, and uncut controls. After treatment, changes in tree basal area and canopy openness in the pond basins reflected reductions in upland basal areas. Specifically, control ponds had significantly higher basal area and lower openness than ponds cut to their margins, while ponds with uncut buffers and partially cut buffers were intermediate. Changes in plant communities were evident in the ground layer and shrub/large regeneration layer. After treatment and over time, the control stands did not change significantly in ground layer structure or shrub/large regeneration layer composition. The three upland harvest treatments displayed increasingly greater deviation from their starting conditions and from the control along a gradient of increasing treatment intensity, from the buffer treatment to the partially cut buffer to the clearcut. The response in the ground layer was largely associated with increased sedge and grass cover, while the response in the shrub/large regeneration layer was associated with increases of Salix sp., Alnus incana, and Populus tremuloides. Our results indicate that adjacent upland timber harvest can lead to altered plant communities within seasonal ponds, at least temporarily. Moreover, uncut forest buffers (∼15.25 m) surrounding seasonal ponds can mitigate plant community changes to some degree. If seasonal ponds are an important resource on the management landscape and a high percentage of upland forest is in a recently cut condition at any given time, than use of harvest buffers around seasonal ponds may be an appropriate approach for mitigating short term alteration of pond plant communities.  相似文献   

3.
Development of understory vegetation has been influenced by the many densely stocked second-growth forest stands in North America, which have an extended stem exclusion successional stage. Understory composition and structure is important for ecosystem functioning, while also having social and economic value through the harvest of certain herb and shrub species. The potential for co-management of young and mature, managed and unmanaged stands for timber and non-timber forest products (NTFPs) was assessed in two separate replicated experiments. Experiment A examined pole-sized lodgepole pine (Pinus contorta) stands that had been pre-commercially thinned (PCT) to target densities of 250, 500, 1000, and 2000 stems/ha. Half of each of these four thinning units was repeatedly fertilized, resulting in eight experimental units. Experiment B examined six different stand types: young plantations, pole-sized lodgepole pine stands either PCT, PCT plus repeated fertilization, or unthinned, mature, and old growth. Fifty-four herb and shrub species were identified as potential NTFPs, with the responses of individual species, as well as mean total herb, shrub, berry-producing and overall total NTFPs being assessed. In Experiment A, mean total abundance (crown volume index) of NTFPs, as well as mean total herb NTFPs were significantly greater in fertilized than in unfertilized stands. Thinning treatments did not significantly affect NTFP volume, however, fertilization treatments produced five significant responses by individual species (Achillea millefolium, Epilobium angustifolium, Taraxacum officinale, Arctostaphylos uva-ursi, Rubus idaeus). In Experiment B, four of the six species responses that were significant had greater abundance in young, managed stands (young plantation, thinned, or thinned-fertilized) than in the unmanaged stands. Mean total NTFP volume and mean total herb NTFP volume also followed this pattern. A. uva-ursi, E. angustifolium, Lonicera involucrata, Sorbus sitchensis and Thalictrum occidentale all had significantly higher abundance in young, managed stands than in all other treatments. Results suggest that co-management for timber and NTFPs is possible in this ecosystem, with further research needed to evaluate livelihood values of these crops.  相似文献   

4.
In eastern Canada, boreal forests develop structural diversity in association with time since stand replacing fire. In some regions, this is associated with significant changes in the bryophyte community (Sphagnum moss invasion) and paludification (thick waterlogged forest floor development). The bryophyte community responds to opening of the canopy, and increasing moisture by replacement of slow growing species by faster growing Sphagnum spp. (e.g. magellanicum, fallax) that are dependent on constant hydration. Within a forest management context, partial harvest systems have been proposed as a strategy to maintain structural diversity, which is currently not accomplished with low retention systems. However, it is unknown whether these interventions will effectively accelerate community succession. The questions addressed in this study were: (1) is the composition of Sphagnum colonies in partially cut stands more similar to old-growth communities than in control, and low retention cut stands, (2) what aspects of harvest disturbance drive these changes, and (3) is the growth rate of Sphagnum capillifolium (an early successional shade tolerant species) different in partial versus low retention harvest systems? After harvest, Sphagnum patch size was reduced by 19.8% and 11.7% after low retention and partial harvest, respectively. While trends were not constant across three separate partial cut trials, the proportion of Sphagnum magellanicum, Sphagnum fallax and Sphagnum fuscum increased compared to controls and low retention 1–2 years after harvest. Models of percent Sphagnum cover indicated machinery track cover, percent cover of vascular plants, and patch depth were positive factors, while the influence of open canopy varied among species. Despite the inclusion of individual disturbance variables, the summary variable ‘treatment’ was significant in all models. Growth of S. capillifolium in partial cuts was intermediate to growth rates in control and low-retention cuts. Growth was positively influenced by slash cover and, contrary to the patch level, negatively influenced by track cover. These results indicate that partial harvest does represent an intermediate level of disturbance, as direct and indirect harvest effects were reduced, as was Sphagnum death. Change in composition 1 and 2 years after harvest indicates that partial harvests may effectively shift the bryophyte community towards an older community type and may thus be used to create landscape diversity. Long term trends and entire community compositions need to be assessed before this can be stated definitively. However, as paludified stands are less productive, the capacity of these partially harvested sites to produce merchantable timber is questioned.  相似文献   

5.
Ecological investigation of the distribution and composition of dung beetle species utilizing feces of wild mammals has rarely been attempted, especially in cool-temperate forests. In this study we recorded and evaluated the community structure of dung beetle species utilizing feces of Japanese monkeys (Macaca fuscata) by conducting a macro-scale inventory survey in the northeastern Shirakami Mountains, located in the northernmost part of mainland Japan. Moreover, to assess the effect of human-induced forest disturbance on species assemblages in this cool-temperate area, we also carried out field sampling using pitfall traps baited with monkey feces in primary forests, secondary forests, and coniferous plantations, during every season except winter. Our findings were: (1) 14 dung beetle species comprising eight dwellers and six tunnelers utilized monkey feces; (2) Aphodius eccoptus was the core species in dung beetle assemblages attracted to the feces during spring and summer; (3) the frequency of occurrence of every species was extremely high in spring; and (4) human-induced forest disturbance did not increase species richness or biomass in cool-temperate forests.  相似文献   

6.
Leaf nitrogen content (Nmass, %) and leaf mass per area (LMA, g m−2) are two important features that are closely linked to the photosynthetic performance of plants and, thus, the NPP of forest ecosystems. Forest management practices, such as burning and thinning, change stand structure and soil dynamics, which may result in changes in Nmass and LMA. The objective of this study was to understand how Nmass and LMA of seven canopy tree species/genus (Quercus alba, Q. coccinea, Q. prinus, Q. velutina, Carya spp., Acer rubrum, and Liriodendron tulipifera) responded to (i) thinning and/or burning treatments and to (ii) different landscape and soil properties in southern Ohio. We collected leaves from the top, and bottom, of five individuals of each taxa in each treatment unit. Leave traits (Nmass and LMA) were compared using analysis of variance followed by orthogonal contrasts. To further understand the factors that influence the canopy leaf traits, we used regression tree analysis (RTA) to examine how variations of LMA and Nmass were linked to thinning and/or burning treatments, soil, and landscape variables. Finally, we assessed the potential ramifications of changes in these traits on canopy carbon budgets using a PnET-Day model, which is a daily time-step canopy carbon exchange model. We found significant effects of thinning, burning, and their interactions on LMA at the bottom of the crown while none of the treatments showed significant effects on LMA at the top of the crown. Nmass responded significantly to only burning treatment. RTA results exhibited minor effects of landscape features and soil properties on Nmass and LMA. Interspecific differences accounted for most variations of both leaf traits. Sensitivity analysis of PnET-Day model suggested these subcanopy changes in LMA increased the annual net primary production (NPP) by 8%. In summary, our results suggest that forest management can substantially influence canopy leaf traits such as Nmass and LMA and that alteration of these traits can influence forest NPP. Given the role of forests as global carbon sinks, the potential influence of thinning and burning on canopy traits, and thus NPP, is an important consideration for forest management.  相似文献   

7.
In the Congo Basin where most timber species are light-demanding, the low logging intensities commonly implemented (1-2 trees harvested ha−1) do not provide sufficient canopy gaps to ensure species regeneration. The regeneration of light-demanding timber species may therefore benefit from more intensive logging, or from post-harvest treatments such as thinning by poison girdling that increases light penetration. Little is known of the impact of post-harvest treatments on the floristic composition of tropical moist forests. This study therefore aimed to assess the effects of low and high selective logging (?2.33 and 4.73 trees harvested ha−1, and ?4.96 and 9.16 m2 ha−1 of basal area removed (logging + damage), respectively) - followed or not by thinning (?21.14 trees thinned ha−1, and ?6.57 m2 ha−1 of basal area removed) - on the floristic composition of a tropical moist forest in the Central African Republic, from 7 to 23 years after logging.We analyzed abundance data for 110 tree genera recorded every year for 14 years in 25 one-hectare permanent subplots. We used multivariate analysis to detect floristic variations between treatments and we assessed changes in floristic composition throughout the period. We compared floristic composition recovery between thinned and unthinned subplots, using unlogged subplots as a reference characterizing the pre-logging floristic composition.Logging and thinning had little impact on the floristic composition of the subplots as quantified 7 to 23 years later, though they did increase the proportion of pioneer species. Surprisingly, additional thinning at both logging levels failed to further distance floristic composition from that of the unlogged subplots, though it did increase disturbance intensity. Floristic composition recovery appeared to be facilitated when thinning was associated with logging. Thinning seemed to favor the growth and survival of non-pioneer species, to the detriment of pioneer species. These non-pioneer species could either be non-pioneer light demanders or shade-bearers. One explanation for this is that thinning by tree-poison girdling increased light availability without causing major damage to the forest, and thus increased the growth and survival of advance regeneration. The resulting enhanced competition then reduced the survival of pioneer species.  相似文献   

8.
Currently, information about the effect of forest management on biodiversity of subtropical plantation forests in Asia is quite limited. In this study, we compared the spider community structures and guild compositions of subtropical Cryptomeria japonica plantation forests receiving different degree of thinning (0, 25 and 50 %) in central Taiwan. The ground spider diversities and environmental variables were sampled/measured once every 3 months for 1 year before thinning and 2 years after thinning. Results showed that before thinning spider compositions did not differ significantly among three plantation forest types. Two years after thinning, spider species and family compositions of three plantation forest types differed significantly. In all three plantation forest types, the spider composition differed from year to year, indicating existence of temporal variations in spider diversity. Ground hunters (increased 200–600 % in thinned forests), sheet web weavers (increased 50–300 % in thinned forests) and space web weavers (decreased 30–50 % in thinned forests) were the major contributors of the observed spider composition differences among plantation forests receiving different treatments. The stands receiving thinning treatments also had higher illumination, litter decomposition rate, temperature and understory vegetation density. Thinning treatments might have changed the structures of understory vegetation and canopy cover and consequently resulted in abundance and diversity changes of these guilds. Moreover, the heterogeneity in understory vegetation recovery rate and temporal variation of spider composition might further generate spider diversity variations in subtropical forests receiving different degree of thinning.  相似文献   

9.
10.
In order to better understand the structure and composition of forest plant communities, we aimed to predict the abundance of understory herbaceous species locally at the stand level and according to different environments. For this, we seeked to model species distributions of abundance at a regional scale in relationship with the local stand structure (canopy openness) and regional soil resources (soil pH).Floristic inventories, performed in different light and soil conditions located in 1202 records of north eastern France, were used to analyze the combined effect of canopy openness and soil pH on the abundance of 12 common western European forest species: Anemone nemorosa, Deschampsia flexuosa, Festuca altissima, Hedera helix, Lamium galeobdolon, Lonicera periclymenum, Molinia caerulea, Oxalis acetosella, Pteridium aquilinum, Rubus fruticosus, Rubus idaeus, and Vaccinium myrtillus. Ordinal regression models relating species abundance responses to their environment were developed.For most species (eight out of 12), distribution was significantly affected by canopy openness and soil pH. Differences among low-abundance (i.e. cover <25%) and high-abundance (i.e. cover >25%) responses were noted for 11 species along the canopy openness gradient and four species along the pH gradient. The present study quantifies optimal light and soil nutrient requirements for high-abundance responses and quantifies light and soil nutrients tolerance conditions for low-abundance responses. The combination of both factors highlights the pre-eminence of pH conditions occurrence and canopy openness for species abundance.The models developed by this study may be used to define canopy openness thresholds in function of soil characteristics to control the development of species during forest regeneration. The species-specific reactions on local canopy openness along a regional soil gradient illustrate the need for a species-specific management approach.  相似文献   

11.
Dry Douglas-fir (Pseudotsuga menziesii) forests offer a wide range of timber and non-timber values, which may benefit from a balanced timber harvest by variable retention systems with conservation of biodiversity. A major component of biodiversity are forest floor small mammal communities whose abundance and diversity serve as ecological indicators of significant change in forest structure and function from harvesting activities. This study was designed to test the hypotheses that abundance, reproduction, and survival of (i) the southern red-backed vole (Myodes gapperi, formerly Clethrionomys gapperi), will decline; (ii) the deer mouse (Peromyscus maniculatus), will be similar; and (iii) the meadow vole (Microtus pennsylvanicus) and northwestern chipmunk (Tamias amoenus), will increase, with decreasing levels of tree retention. Small mammal populations were live-trapped from 1994 to 1997 in replicated sites of uncut forest, 20% and 50% volume removal by single tree selection, 20%, 35%, and 50% patch cuts based on openings of 0.1–1.6 ha, and small 1.6 ha clearcuts in Douglas-fir forest near Kamloops, British Columbia, Canada. M. gapperi dominated the small mammal community, starting with an abundance of 74–98 animals/ha with mean values ranging from 33 to 51 animals/ha. In the two post-harvest years, abundance, reproduction, and survival of M. gapperi populations were consistently similar among uncut forest and the various levels of tree retention. Thereafter, M. gapperi was seldom found on the small clearcuts. M. pennsylvanicus, T. amoenus, and P. maniculatus occurred predominantly in clearcut sites. As with other types of forest disturbance, responses to our treatments were species-specific. The most striking result was the high abundance and productivity of M. gapperi populations in a dry forest ecosystem, a novel result for this bio-indicator species of closed-canopy forest conditions. At least with respect to small mammals, the retention systems studied seem to enable timber extraction and maintenance of mature forest habitat in these dry fir ecosystems.  相似文献   

12.

Context

Understanding the range of possible climate change impacts on forests and the interactions between them is vital to sustainable forest management.

Aims

We examine whether the combined influence of climate change and timber harvest will affect tree species distribution and productivity beyond predictions based on climate alone.

Methods

We used the landscape disturbance model LANDIS-II to simulate two climate and two harvest scenarios in 14,000 ha of managed watersheds.

Results

The elevated temperature led to a decline in the abundance of boreal species and a substantial increase in some temperate and pioneer species. Importantly, the interaction of climate change and timber harvest yielded changes in the distribution of some species that would not be expected based on climate alone. Conversely, some late-successional species exhibited resistance to climate-driven changes in their distribution. Climate change caused an increase in forest productivity when harvest was simulated, but a decrease in no-harvest scenarios. A time lag in forest response was likely responsible for this decrease in the absence of widespread mortality.

Conclusions

The finding that disturbance may drive the range expansion of early-successional broadleaved species and cause a decline of red spruce has implications for forest community associations, as well as for forest management where conifers are favoured for pulp production.  相似文献   

13.
Demand for alternative energy sources has led to increased interest in intensive biomass production. When applied across a broad spatial extent, intensive biomass production in forests, which support a large proportion of biodiversity, may alter species composition, nutrient cycling and subsequently biodiversity. Because forest thinning and fuels treatment thinning are viewed as possible wide-spread biomass harvest options, it is important to understand what is known about forest biodiversity response to these practices and what additional information is needed by forest managers and policymakers. Therefore, we summarized documented relationships between forest thinning treatments and forest biodiversity from 505 biodiversity effect sizes (incl. taxa and guild abundance and species richness measures) from 33 studies conducted across North America. We used meta-analysis to summarize biodiversity response by region, taxa and harvest treatments. Biodiversity responses included species richness, diversity, abundance of taxa or groups of species (guilds) and abundance of individual species for birds, mammals, reptiles, amphibians, and invertebrates. Forest thinning treatments had generally positive or neutral effects on diversity and abundance across all taxa, although thinning intensity and the type of thinning conducted may at least partially drive the magnitude of response. Our review highlights the need for more research to determine effects of thinning on amphibians and reptiles and manipulative experiments designed to test the effects of biomass removal on biodiversity.  相似文献   

14.
Boreal forest birds have adapted to changes caused by natural disturbances such as fire and this adaptation forms the basis for the Natural Disturbance Paradigm (NDP) underlying recent proposed changes in forest harvesting practices in western Canada. To date, this paradigm has been evaluated primarily at the stand level and within conventional harvesting systems. The potential for improvements in avian conservation at the landscape scale by adopting the NDP approach is largely unknown. We examined the effects of landscape-scale disturbances on forest bird communities by contrasting richness and abundance of birds in (1) 16 single-pass harvest sites with residual forest patches, (2) 29 multi-pass harvest sites with residuals; and (3) 15 salvage-logged post-fire sites with variable harvest intensity. We contrasted bird communities in these treatments with those in unsalvaged post-fire sites of similar age. Post-fire sites were used to provide a metric of the Natural Range of Variation (NRV) to be expected in bird communities. Sites were surveyed for avian community composition and abundance 1–5 years post-disturbance. Redundancy analysis indicated that bird communities differed from the NRV in all of the harvest treatments. However, single-pass harvests provided a somewhat better fit to NRV than did multi-pass harvesting. Avian community similarity was influenced by non-linear responses to area harvested, amount of residual retention, residual composition and pre-disturbance forest composition. An optimization routine created from a General Linear Model, suggests that community similarity to NRV can be maximized by using single-pass harvests over multi-pass harvests, harvesting 66–88% of the timber in the planning unit, and retaining 5–19% of the disturbance area as live residual patches, with 50% of harvests having at least 9% of the area in residuals.  相似文献   

15.
We compared avian communities among three timber harvesting treatments in 45-m wide even-age riparian management zones (RMZs) placed between upland clearcuts and along one side of first- or second-order streams in northern Minnesota, USA. The RMZs had three treatments: (1) unharvested, (2) intermediate residual basal area (RBA) (targeted goal 11.5 m2/ha, realized 16.0 m2/ha), and (3) low RBA (targeted goal 5.7 m2/ha, realized 8.7 m2/ha). Surveys were conducted one year pre-harvest and three consecutive years post-harvest. There was no change in species richness, diversity, or total abundance associated with harvest but there were shifts in the types of birds within the community. In particular, White-throated Sparrows (Zonotrichia albicollis) and Chestnut-sided Warblers (Dendroica pensylvanica) increased while Ovenbirds (Seiurus aurocapilla) and Red-eyed Vireos (Vireo olivaceus) decreased. The decline of avian species associated with mature forest in the partially harvested treatments relative to controls indicates that maintaining an unharvested RMZ adjacent to an upland harvest may aid in maintaining avian species associated mature forest in Minnesota for at least three years post-harvest. However, our observations do not reflect reproductive success, which is an area for future research.  相似文献   

16.
We studied the short-term effects of a catastrophic windstorm and subsequent salvage-logging and prescribed-burning fuel-reduction treatments on ground beetle (Coleoptera: Carabidae) assemblages in a sub-boreal forest in northeastern Minnesota, USA. During 2000–2003, 29,873 ground beetles represented by 71 species were caught in unbaited and baited pitfall traps in aspen/birch/conifer (ABC) and jack pine (JP) cover types. At the family level, both land-area treatment and cover type had significant effects on ground beetle trap catches, but there were no effects of pinenes and ethanol as baits. Six times more beetles were trapped in the burned forests than in the other land-area treatments; more beetles were caught in undisturbed than in wind-disturbed sites, and one-third more beetles were caught in the ABC than in the JP cover type. Thus, the windstorm generally reduced the activity-abundance of the beetles, but prescribed-burning increased it. Both salvaged and burned forest plots (especially in the ABC cover type) had the greatest species richness, diversity, and the most unique species assemblages. There was a highly similar ground beetle species composition (nearly 100%) between the ABC and JP burned forests, indicating that burning was a more primary driver of composition than cover type. At the species level, Pterostichus melanarius, an invasive ground beetle from Europe and a cover type generalist, was the most abundant beetle in the study (one-third of the total catch), and was caught in greatest numbers in burned forests. Removal of P. melanarius from the species composition analyses altered similarities among cover types and land-area treatments. Sphaeroderus nitidicollis brevoorti and Myas cyanescens were caught exclusively in the ABC and JP cover type, respectively; two rare pyrophilous species, Sericoda obsoleta and Sericoda quadripunctata, were only caught in burned sites; three forest species, Pterostichus coracinus, P. pensylvanicus, and Sphaeroderus lecontei, were caught more often in undisturbed JP sites; and two frequently trapped, open-habitat species, Agonum cupripenne and Poecilus l. lucublandus, were nearly absent from the undisturbed and wind-disturbed sites, as salvage-logging had a significant positive effect on their activity-abundance. Most species of Amara and Harpalus were trapped only in the salvaged or burned sites, indicating invasion of these disturbed sites by open-habitat species. We conclude that both the combined effect of fuel-reduction activities subsequent to the wind event and the numerical response of the invasive P. melanarius to habitat disturbances can alter the short-term succession of ground beetle assemblages in the sub-boreal forest.  相似文献   

17.
We assessed the effectiveness of alternative forest management strategies for maintaining American martens (Martes americana) in a sub-boreal landscape subject to an extensive mountain pine beetle (Dendroctonus ponderosae) outbreak, associated salvage logging, and climate change in north-central British Columbia, Canada. We structured the analysis in a Bayesian network (BN) meta-model that incorporated the results of spatially explicit modelling of landscape conditions (natural and logging disturbance, habitat quality, number of potential territories, and connectedness of territories) with analytical population modelling. The BN meta-model was then used to examine habitat and population size responses (adult females only are presented) to management scenarios, in the context of uncertainty of model parameters, management objectives, and climate change. Status quo management is dominantly clear-cutting with 3–20% of each harvest unit retained as mature patches, with reforestation by planting in the remainder. Management options we examined were: (1) the status quo, (2) varying the total annual timber harvest on the landscape (100%, 80% or 50% of current long-term sustained yield estimates), (3) the protection of understory trees during logging, and (4) 30–70% retention of overstory (partial cutting in distinct patches <1 ha in size) in each harvest unit, for 33% or 50% of the annual timber harvest. We found that marten habitat and population size declined substantively with the beetle outbreak and associated salvage cutting. The choice of management strategy then had a long-term effect on the potential for marten recovery after the beetle outbreak. Partial cutting scenarios had the greatest average long-term marten population levels, followed in order by reduced harvest rates, understory protection, and the status quo. Management scenarios with the best chance of meeting conservation goals without over-protecting habitat (and thus unnecessarily constraining timber management) varied with the population objective chosen. The choice of management strategy will depend on the weighting of marten outcomes against the economic desirability of timber harvest strategies, willingness to gamble on climate change, and the time-frame of interest.  相似文献   

18.
In conifer plantations, enhancement of species diversity has become an important management goal. Although thinning is a useful method to enhance diversity, determining optimum thinning intensities may be rather complicated because of potential trade-offs among a broad array of management goals (e.g., recovery of biodiversity, increasing individual tree sizes, increasing net primary production, saving management costs). To evaluate the optimum thinning intensity by analyzing these relationships, we conducted a thinning experiment with three different thinning intensities—unthinned, 33% thinned, and 67% thinned—in a Cryptomeria japonica plantation in 2003, and investigated the number, diameter at breast height (DBH), and diversity of hardwoods (height?>?1.5?m) in 2008, and the growth of conifers over five?years. In hardwoods, the number of individuals, number of species, mean DBH, and total basal area were greatest in the 67% thinned treatment, irrespective of successional status. However, Shannon’s diversity index did not differ among the three treatments due to a disproportionate increase with thinning intensity in the abundance of a mid-successional species, Cornus controversa. Diameter growth of conifers was also highest in the 67% thinned and lowest in the unthinned treatment, whereas the reverse was true for stand volume increment. These results suggest that intensive thinning is a reliable method to convert conifer plantations into conifer–hardwood mixed forests at canopy level much more quickly and consistently than weak thinning, although primary production is to some extent reduced. If forest managers prefer sustainable timber production of conifers rather than full recovery of diversity, weak thinning may be suitable.  相似文献   

19.
Thinning of young forest stands encourages development of the understory layer by increasing the levels of light and belowground resources. High-intensity thinning, with associated ground disturbance and high light levels, can lead to dominance by early seral species or by a few species of shrubs, while low-intensity thinning may not increase resource levels enough to encourage forest herbs. Changes in herb-layer abundance can be hard to detect because forest-floor herbs are often slow growing, but flowering increases rapidly in response to high resource levels. This study examined flowering of the understory herb community before and 5 years after low-intensity thinning. Flowering of 10 herb species was sampled in 62 nine-meter radius plots in six treatment units within the H.J. Andrews Experimental Forest. Thinning proved to be the most important determinant of the composition of the flowering assemblage (that is of which plants were flowering in a given plot). Old-growth species and release species (those specializing in large forest openings) showed higher numbers of flowering ramets following thinning. Release species also showed significant positive linear relationships with the plot-level degree of reduction in Douglas-fir (Pseudotsuga menziesii) basal area (BA), while old-growth species showed no significant relationships and forest generalist species showed significant negative linear relationships with reduction in BA. Plot-level reduction in BA explained little of the variation in numbers of flowering ramets per plot for most species except for the release species Hieracium albiflorum and the forest generalist Trillium ovatum. The overall lack of strong linear relationships between herb flowering and reduction in BA makes it difficult to predict optimal thinning intensities for these species. However, the results of this study suggest that even when low-intensity thinning does not significantly change herb abundance it could still influence the ecology of the understory herb community by increasing carbon allocation to sexual reproduction.  相似文献   

20.
Forest communities across the landscape of the Central Hardwood Forest Region are experiencing a transition from dominance by oak (Quercus) and hickory (Carya) to maple (Acer) driven largely by a prolonged period of fire suppression. In many cases, this shift in community composition, structure, and function is considered undesirable as oak-hickory forests are valued for timber, wildlife habitat, and natural heritage. Considerable management and restoration efforts target the restoration of oak-hickory forest communities, yet treatments have yielded varying degrees of success. In some cases, difficulties in meeting targets may be due to ecological thresholds created by complex vegetation-environment interactions that maintain the maple-dominated community state. We examined direct and indirect interactions among vegetation layers and environmental gradients for the mature, second-growth forest communities of the Ironton Unit of the Wayne National Forest (WNF) in southeastern Ohio. Using a stratified random approach, we identified 72 study communities with trees at least 70 years old and without evidence of recent disturbance. Within these communities, we sampled all overstory vegetation on two-four 500 m2 plots and recorded saplings and ground-flora species in nested sub-plots. At each plot, we also collected soil samples for physical and chemical analyses and recorded physiographic variables. Our first objective was to describe the Ironton forest landscape, where communities were likely transitioning from oak to maple. To identify such patterns, we used ordination analyses that relate species occurrence to implied environmental gradients. Our second objective was to use the relationships to develop a structural equation model (SEM) to quantify the strength of pathways among the canopy, sapling, and ground-flora vegetation layers and environmental factors (e.g., soil chemistry and physiographic position). Our results indicate that the forest landscape of the Ironton Unit of the WNF is at a transition point with communities dominated by either oak or maple, and a sapling layer dominated by maple. Maple may be most likely to replace oak and hickory in the canopies of communities at mid- and lower-slope positions with intermediate soil moisture. This transition will likely have cascading effects throughout the sapling and ground-flora layers, which SEM demonstrates are directly influenced by the canopy. We believe the simultaneous consideration of direct and indirect interactions shaping vegetation structure and composition using techniques such as SEM will advance understanding of the current transition from an oak-hickory to a maple-dominated forest landscape. This information will contribute to the continued improvement of appropriate forest management and ecosystem restoration techniques for the Central Hardwood Forest Region, including those designed to shift the dominance of forest communities from maple to oak.  相似文献   

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