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1.
Many organisms persist in fragmented habitat where movement between patches is essential for long-term demographic and genetic stability. In the absence of direct observation of movement, connectivity or isolation metrics are useful to characterize potential patch-level connectivity. However, multiple metrics exist at varying levels of complexity, and empirical data on species distribution are rarely used to compare performance of metrics. We compared 12 connectivity metrics of varying degrees of complexity to determine which metric best predicts the distribution of prairie dog colonies along an urban gradient of 385 isolated habitat patches in Denver, Colorado, USA. We found that a modified version of the incidence function model including area-weighting of patches and a cost-weighted distance surface best predicted occupancy, where we assumed roads were fairly impermeable to movement, and low-lying drainages provided dispersal corridors. We also found this result to be robust to a range of cost weight parameters. Our results suggest that metrics should incorporate both patch area and the composition of the surrounding matrix. These results provide guidance for improved landscape habitat modeling in fragmented landscapes and can help identify target habitat for conservation and management of prairie dogs in urban systems.  相似文献   

2.
Habitat isolation can affect the distribution and abundance of wildlife, but it is an ambiguous attribute to measure. Presumably, isolation is a characteristic of a habitat patch that reflects how spatially inaccessible it is to dispersing organisms. We identified four isolation metrics (nearest-neighbor distance, Voronoi polygons, proximity index, and habitat buffers) that were representative of the different families of metrics that are commonly used in the literature to measure patch isolation. Using simulated data, we evaluated the ability of each isolation metric to predict animal dispersal. We examined the simulated movement of organisms in two types of landscapes: an artificially-generated point-pattern landscapes where patch size and shape were consistent and only the arrangement of patches varied, and realistic landscapes derived from a geographic information system (GIS) of forest-vegetation maps where patch size, shape, and isolation were variable. We tested the performance of the four isolation metrics by examining the strength of the correlation between observed immigration rate in the simulations and each patch isolation metric. We also evaluated whether each isolation metric would perform consistently under varying conditions of patch size/shape, total amount of habitat in the landscape, and proximity of the patch to the landscape edge. The results indicate that a commonly-used distance-based metric, nearest-neighbor distance, did not adequately predict immigration rate when patch size and shape were variable. Area-informed isolation metrics, such as the amount of available habitat within a given radius of a patch, were most successful at predicting immigration. Overall, the use of area-informed metrics is advocated despite the limitation that these metrics require parameterization to reflect the movement capacity of the organism studied.This revised version was published online in May 2005 with corrections to the Cover Date.  相似文献   

3.
Habitat fragmentation is expected to disrupt dispersal, and thus we explored how patch metrics of landscape structure, such as percolation thresholds used to define landscape connectivity, corresponded with dispersal success on neutral landscapes. We simulated dispersal as either a purely random process (random direction and random step lengths) or as an area-limited random walk (random direction, but movement limited to an adjacent cell at each dispersal step) and quantified dispersal success for 1000 individuals on random and fractal landscape maps across a range of habitat abundance and fragmentation. Dispersal success increased with the number of cells a disperser could search (m), but poor dispersers (m<5) searching via area-limited dispersal on fractal landscapes were more successful at locating suitable habitat than random dispersers on either random or fractal landscapes. Dispersal success was enhanced on fractal landscapes relative to random ones because of the greater spatial contagion of habitat. Dispersal success decreased proportionate to habitat loss for poor dispersers (m=1) on random landscapes, but exhibited an abrupt threshold at low levels of habitat abundance (p<0.1) for area-limited dispersers (m<10) on fractal landscapes. Conventional metrics of patch structure, including percolation, did not exhibit threshold behavior in the region of the dispersal threshold. A lacunarity analysis of the gap structure of landscape patterns, however, revealed a strong threshold in the variability of gap sizes at low levels of habitat abundance (p<0.1) in fractal landscapes, the same region in which abrupt declines in dispersal success were observed. The interpatch distances or gaps across which dispersers must move in search of suitable habitat should influence dispersal success, and our results suggest that there is a critical gap-size structure to fractal landscapes that interferes with the ability of dispersers to locate suitable habitat when habitat is rare. We suggest that the gap structure of landscapes is a more important determinant of dispersal than patch structure, although both are ultimately required to predict the ecological consequences of habitat fragmentation.  相似文献   

4.
How should we measure landscape connectivity?   总被引:9,自引:0,他引:9  
The methods for measuring landscape connectivity have never been compared or tested for their responses to habitat fragmentation. We simulated movement, mortality and boundary reactions across a wide range of landscape structures to analyze the response of landscape connectivity measures to habitat fragmentation. Landscape connectivity was measured as either dispersal success or search time, based on immigration into all habitat patches in the landscape. Both measures indicated higher connectivity in more fragmented landscapes, a potential for problematic conclusions for conservation plans. We introduce cell immigration as a new measure for landscape connectivity. Cell immigration is the rate of immigration into equal-sized habitat cells in the landscape. It includes both within- and between-patch movement, and shows a negative response to habitat fragmentation. This complies with intuition and existing theoretical work. This method for measuring connectivity is highly robust to reductions in sample size (i.e., number of habitat cells included in the estimate), and we hypothesize that it therefore should be amenable to use in empirical studies. The connectivity measures were weakly correlated to each other and are therefore generally not comparable. We also tested immigration into a single patch as an index of connectivity by comparing it to cell immigration over the landscape. This is essentially a comparison between patch-scale and landscape-scale measurement, and revealed some potential for patch immigration to predict connectivity at the landscape scale. However, this relationship depends on the size of the single patch, the dispersal characteristics of the species, and the amount of habitat in the landscape. We conclude that the response of connectivity measures to habitat fragmentation should be understood before deriving conclusions for conservation management.  相似文献   

5.
Maintaining and restoring connectivity among high-quality habitat patches is recognized as an important goal for the conservation of animal populations. To provide an efficient measure of potential connectivity pathways in heterogeneous landscapes, least-cost route analysis has been combined with graph-theoretical techniques. In this study we use spatially explicit least-cost habitat graphs to examine how matrix quality and spatial configuration influence assessments of habitat connectivity. We generated artificial landscapes comprised of three landcover types ranked consistently from low to high quality: inhospitable matrix, hospitable matrix, and habitat. We controlled the area and degree of fragmentation of each landcover in a factorial experiment for a total of 20 combinations replicated 100 times. In each landscape we compared eight sets of relative landcover qualities (cost values of 1 for habitat, between 1.5 and 150 for hospitable matrix, and 3–10,000 for inhospitable matrix). We found that the spatial location of least-cost routes was sensitive to differences in relative cost values assigned to landcover types and that the degree of sensitivity depended on the spatial structure of the landscape. Highest sensitivity was found in landscapes with fragmented habitat and between 20 and 50% hospitable matrix; sensitivity decreased as habitat fragmentation decreased and the amount of hospitable matrix increased. As a means of coping with this sensitivity, we propose identifying multiple low-cost routes between pairs of habitat patches that collectively delineate probable movement zones. These probable movement zones account for uncertainty in least-cost routes and may be more robust to variation in landcover cost values.  相似文献   

6.
Land-use change is forcing many animal populations to inhabit forest patches in which different processes can threaten their survival. Some threatening processes are mainly related to forest patch characteristics, but others depend principally on the landscape spatial context. Thus, the impact of both patch and landscape spatial attributes needs to be assessed to have a better understanding of the habitat spatial attributes that constraint the maintenance of populations in fragmented landscapes. Here, we evaluated the relative effect of three patch-scale (i.e., patch size, shape, and isolation) and five landscape-scale metrics (i.e., forest cover, fragmentation, edge density, mean inter-patch isolation distance, and matrix permeability) on population composition and structure of black howler monkeys (Alouatta pigra) in the Lacandona rainforest, Mexico. We measured the landscape-scale metrics at two spatial scales: within 100 and 500 ha landscapes. Our findings revealed that howler monkeys were more strongly affected by local-scale metrics. Smaller and more isolated forest patches showed a lower number of individuals but at higher densities. Population density also tended to be positively associated to matrices with higher proportion of secondary forests and arboreal crops (i.e. with greater permeability), most probably because these matrices can offer supplementary foods. The immature-to-female ratio also increased with matrix permeability, shape complexity, and edge density; habitat characteristics that can increase landscape connectivity and sources availability. The prevention of habitat loss and isolation, and the increment of matrix permeability are therefore needed for the conservation of this endangered Neotropical mammal.  相似文献   

7.
Landscape connectivity can be viewed from two perspectives that could be considered as extremes of a gradient: functional connectivity (refers to how the behavior of a dispersing organism is affected by landscape structure and elements) and structural connectivity (depends on the spatial configuration of habitat patches in the landscape like vicinity or presence of barriers). Here we argue that dispersal behavior changes with landscape configuration stressing the evolutionary dimension that has often been ignored in landscape ecology. Our working hypothesis is that the functional grain of resource patches in the landscape is a crucial factor shaping individual movements, and therefore influencing landscape connectivity. Such changes are likely to occur on the short-term (some generations). We review empirical studies comparing dispersal behavior in landscapes differing in their fragmentation level, i.e., with variable resource grain. We show that behavioral variation affecting each of the three stages of the dispersal process (emigration, displacement or transfer in the matrix, and immigration) is indeed likely to occur according to selective pressures resulting from changes in the grain of the landscape (mortality or deferred costs). Accordingly, landscape connectivity results from the interaction between the dispersal behavior of individuals and the grain of each particular landscape. The existence of this interaction requires that connectivity estimates (being based on individual-based models, least cost distance algorithms, and structural connectivity metrics or even Euclidian distance) should be carefully evaluated for their applicability with respect to the required level of precision in species-specific and landscape information.  相似文献   

8.
Individual movement is a key process affecting the distribution of animals in heterogeneous landscapes. For specialist species in patchy habitat, a central issue is how dispersal distances are related to landscape structure. We compared dispersal distances for cactus bugs (Chelinidea vittiger) on two naturally fragmented landscapes (≤ 4% suitable habitat) with different matrix structures (i.e., vegetation height of nonsuitable habitat between suitable patches). Using mark-release-recapture studies, we determined that most transfers between cactus patches occurred during the mating season. Dispersal distances were reduced by > 50% on the landscape that had reduced structural connectivity due to relatively high matrix structure and low patch density. An experiment with detailed movement pathways demonstrated that greater matrix structure decreased mean step lengths, reduced directionality, and thus decreased net displacement by > 60%. However, habitat edges between two matrix elements that differed substantially in resistance to movement were completely permeable. Therefore, the difference in distributions of dispersal distances between the two landscapes mainly reflected the average resistance of matrix habitat and not the level of matrix heterogeneity per se. Our study highlights the merits of combining estimates of dispersal distances with insights on mechanisms from detailed movement pathways, and emphasizes the difficulty of treating dispersal distances of species as fixed traits independent of landscape structure.  相似文献   

9.
Applied ecology could benefit from new tools that identify potential movement pathways of invasive species, particularly where data are sparse. Cost surface analysis (CSA) estimates the permeability (friction) across a landscape and can be applied to dispersal modelling. Increasingly used in a diversity of applications, several fundamental assumptions that might influence the outputs of CSA (cost surfaces and least-cost pathways) have yet to be systematically examined. Thus, we explore two issues: the presumed relationship between habitat preferences and dispersal behaviour as well as the degree of landscape fragmentation through which an organism moves by modelling a total of 18 sensitivity and dispersal scenarios. We explored the effect of fragmentation by altering the friction values (generally assigned using expert opinion) associated with patch and linear features. We compared these sensitivity scenarios in two sites that differed in fragmentation. We also used eastern grey squirrels (Sciurus carolinensis) as an example invading species and compared diffusion models and two contrasting cost surface dispersal scenarios. The diffusion model underestimated spread because squirrels did not move randomly through the landscape. Despite contrasting assumptions regarding dispersal behaviour, the two cost surfaces were strikingly similar while the least-cost paths differed. Furthermore, while the cost surfaces were insensitive to changes in friction values for linear features, they were sensitive to assumptions made for patch features. Our results suggest that movement in fragmented landscapes may be more sensitive to assumptions regarding friction values than contiguous landscapes. Thus, the reliability of CSA may depend not only on the range of friction values used for patches but also the degree of contiguity in the landscape.  相似文献   

10.
Dispersal has been shown to be a key determinant of spatially structured populations. One crucial aspect is predicting patch accessibility: the probability rij of a certain patch j being reached by individuals starting at another patch i. Patch accessibility rij depends on both the landscape structure and the individuals’ dispersal behaviour. To investigate the effects of these factors on rij, we developed a simulation model focusing on animal dispersal. Our model analyses show that there is an important intrinsic effect of the interplay between landscape structure and dispersal behaviour on patch accessibility: the competition between patches for migrants. We derive a formula for patch accessibility. This formula is very simple because it just takes distances into account: not only the distance between start patch and target patch, but also between the start patch and all the other patches in the landscape. Despite its simplicity, the formula is able to cover effects such as the competition for migrants. The formula was found to have high predictive power for a variety of movement behaviours (random walk with various degrees of correlation, Archimedean spirals and loops) in any given landscape. The formula can be interpreted as a generic function for patch accessibility for further population dynamics analyses. It also delivers insights into the consequences of dispersal in fragmented landscapes.  相似文献   

11.
The landscape matrix is suggested to influence the effect of habitat fragmentation on species richness, but the generality of this prediction has not been tested. Here, we used data from 10 independent studies on butterfly species richness, where the matrix surrounding grassland patches was dominated by either forest or arable land to test if matrix land use influenced the response of species richness to patch area and connectivity. To account for the possibility that some of the observed species use the matrix as their main or complementary habitat, we analysed the effects on total species richness and on the richness of grassland specialist and non-specialist (generalists and specialists on other habitat types) butterflies separately. Specialists and non-specialists were defined separately for each dataset. Total species richness and the richness of grassland specialist butterflies were positively related to patch area and forest cover in the matrix, and negatively to patch isolation. The strength of the species-area relationship was modified by matrix land use and had a slope that decreased with increasing forest cover in the matrix. Potential mechanisms for the weaker effect of grassland fragmentation in forest-dominated landscapes are (1) that the forest matrix is more heterogeneous and contains more resources, (2) that small grassland patches in a matrix dominated by arable land suffer more from negative edge effects or (3) that the arable matrix constitutes a stronger barrier to dispersal between populations. Regardless of the mechanisms, our results show that there are general effects of matrix land use across landscapes and regions, and that landscape management that increases matrix quality can be a complement to habitat restoration and re-creation in fragmented landscapes.  相似文献   

12.

Context

Landscape-scale population dynamics are driven in part by movement within and dispersal among habitat patches. Predicting these processes requires information about how movement behavior varies among land cover types.

Objectives

We investigated how butterfly movement in a heterogeneous landscape varies within and between habitat and matrix land cover types, and the implications of these differences for within-patch residence times and among-patch connectivity.

Methods

We empirically measured movement behavior in the Baltimore checkerspot butterfly (Euphydryas phaeton) in three land cover classes that broadly constitute habitat and two classes that constitute matrix. We also measured habitat preference at boundaries. We predicted patch residence times and interpatch dispersal using movement parameters estimated separately for each habitat and matrix land cover subclass (5 categories), or for combined habitat and combined matrix land cover classes (2 categories). We evaluated the effects of including edge behavior on all metrics.

Results

Overall, movement was slower within habitat land cover types, and faster in matrix cover types. Butterflies at forest edges were biased to remain in open areas, and connectivity and patch residence times were most affected by behavior at structural edges. Differences in movement between matrix subclasses had a greater effect on predictions about connectivity than differences between habitat subclasses. Differences in movement among habitat subclasses had a greater effect on residence times.

Conclusions

Our findings highlight the importance of careful classification of movement and land cover in heterogeneous landscapes, and reveal how subtle differences in behavioral responses to land cover can affect landscape-scale outcomes.
  相似文献   

13.
Although the landscape matrix is increasingly incorporated into spatial-ecological population studies, little consideration has been given to the likely possibility that patch quality is confounded with the composition of the matrix surrounding each patch. For example, the nutritional quality of host-plant patches to an herbivore may be highly correlated with matrix composition, consequently obfuscating the importance of the matrix itself to interpatch dispersal. From a literature survey of the effects of the matrix on herbivore movement among host-plant patches, we found that 55% of the studies (6/11) failed to experimentally or statistically isolate the effects of the matrix from potential patch-quality effects on dispersal. Most studies consisted of mark-recapture experiments in natural landscapes where patch equality was not controlled or manipulated. Of the few studies that evaluated the relationship between matrix composition and patch quality, all of them (3/3) found that these two landscape factors covaried. These data suggest that in most matrix studies, effects of the matrix on dispersal may be wholly, or in part, due to underlying differences in patch quality. This revised version was published online in May 2005 with corrections to the Cover Date. This revised version was published online in July 2006 with corrections to the Cover Date.  相似文献   

14.
In spatial studies of populations, Euclidean distance is commonly used to measure the structural connectivity between habitat patches. The role of the matrix on patch connectivity is thereby ignored. However, the importance of the matrix for (dispersal) movement is increasingly being acknowledged. Our study compared the cost-distance measure with the Euclidean distance. The cost-distance is a simple GIS-calculated connectivity measure that incorporates the resistance of the landscape matrix to movement behaviour. We used presence-absence data from a field study on the Speckled wood butterfly in two Belgian landscapes. Logistic regression revealed that the cost-distance measure had a significantly better predictive power than the Euclidean distance. This result was consistent for all the six sets of different matrix resistance values. In our study the cost-distance proves to be a better connectivity measure than the Euclidean distance. This revised version was published online in July 2006 with corrections to the Cover Date.  相似文献   

15.
Habitat fragmentation, patch quality and landscape structure are important predictors for species richness. However, conservation strategies targeting single species mainly focus on habitat patches and neglect possible effects of the surrounding landscape. This project assesses the impact of management, habitat fragmentation and landscape structure at different spatial scales on the distribution of three endangered butterfly species, Boloria selene, Boloria titania and Brenthis ino. We selected 36 study sites in the Swiss Alps differing in (1) the proportion of suitable habitat (i.e., wetlands); (2) the proportion of potential dispersal barriers (forest) in the surrounding landscape; (3) altitude; (4) habitat area and (5) management (mowing versus grazing). Three surveys per study site were conducted during the adult flight period to estimate occurrence and density of each species. For the best disperser B. selene the probability of occurrence was positively related to increasing proportion of wetland on a large spatial scale (radius: 4,000 m), for the medium disperser B. ino on an intermediate spatial scale (2,000 m) and for the poorest disperser B. titania on a small spatial scale (1,000 m). Nearby forest did not negatively affect butterfly species distribution but instead enhanced the probability of occurrence and the population density of B. titania. The fen-specialist B. selene had a higher probability of occurrence and higher population densities on grazed compared to mown fens. The altitude of the habitat patches affected the occurrence of the three species and increasing habitat area enhanced the probability of occurrence of B. selene and B. ino. We conclude that, the surrounding landscape is of relevance for species distribution, but management and habitat fragmentation are often more important. We suggest that butterfly conservation should not focus only on a patch scale, but also on a landscape scale, taking into account species-specific dispersal abilities.  相似文献   

16.
Acknowledgment that the matrix matters in conserving wildlife in human-modified landscapes is increasing. However, the complex interactions of habitat loss, habitat fragmentation, habitat condition and land use have confounded attempts to disentangle the relative importance of properties of the landscape mosaic, including the matrix. To this end, we controlled for the amount of remnant forest habitat and the level of fragmentation to examine mammal species richness in human-modified landscapes of varying levels of matrix development intensity and patch attributes. We postulated seven alternative models of various patch habitat, landscape and matrix influences on mammal species richness and then tested these models using generalized linear mixed-effects models within an information theoretic framework. Matrix attributes were the most important determinants of terrestrial mammal species richness; matrix development intensity had a strong negative effect and vegetation structural complexity of the matrix had a strong positive effect. Distance to the nearest remnant forest habitat was relatively unimportant. Matrix habitat attributes are potentially a more important indicator of isolation of remnant forest patches than measures of distance to the nearest patch. We conclude that a structurally complex matrix within a human-modified landscape can provide supplementary habitat resources and increase the probability of movement across the landscape, thereby increasing mammal species richness in modified landscapes.  相似文献   

17.
Genetic analysis of landscape connectivity in tree populations   总被引:1,自引:0,他引:1  
Genetic connectivity in plant populations is determined by gene movement within and among populations. When populations become genetically isolated, they are at risk of loss of genetic diversity that is critical to the long-term survival of populations. Anthropogenic landscape change and habitat fragmentation have become so pervasive that they may threaten the genetic connectivity of many plant species. The theoretical consequences of such changes are generally understood, but it is not immediately apparent how concerned we should be for real organisms, distributed across real landscapes. Our goals here are to describe how one can study gene movement of both pollen and seeds in the context of changing landscapes and to explain what we have learned so far. In the first part, we will cover methods of describing pollen movement and then review evidence for the impact of fragmentation in terms of both the level of pollen flow into populations and the genetic diversity of the resulting progeny. In the second part, we will describe methods for contemporary seed movement, and describe findings about gene flow and genetic diversity resulting from seed movement. Evidence for pollen flow suggests high connectivity, but it appears that seed dispersal into fragments may create genetic bottlenecks due to limited seed sources. Future work should address the interaction of pollen and seed flow and attention needs to be paid to both gene flow and the diversity of the incoming gene pool. Moreover, if future work is to model the impact of changing landscapes on propagule movement, with all of its ensuing consequences for genetic connectivity and demographic processes, we will need an effective integration of population genetics and landscape ecology.  相似文献   

18.
The matrix is an important element of landscape mosaics that influences wildlife indirectly through its influence on habitat, and directly, if they live in or move through it. Therefore, to quantify and manage habitat quality for wildlife in modified landscapes, it is necessary to consider the characteristics of both patch and matrix elements of the whole landscape mosaic. To isolate matrix effects from the often simultaneous and confounding influence of patch and landscape characteristics, we identified nineteen 500 m radius landscapes in southeast Queensland, Australia with similar remnant forest patch attributes, habitat loss, and fragmentation, but exhibiting a marked gradient from rural through high-density suburban development of the matrix, quantified by a weighted road-length metric. We measured habitat disturbance, structure, and floristics in patch core, patch edge and matrix landscape elements to characterise how landscape habitat quality changes for small mammals. Correlation analyses identified that with increased matrix development intensity, human disturbance of core sites increased, predators and exotic plant species richness in matrix sites increased, and structural complexity (e.g. logs and stumps) in the matrix decreased. Ordination analyses showed landscape elements were most similar in habitat structure and floristics at low to moderate levels of matrix development, suggesting enhanced landscape habitat quality. Matrix development intensity was not, however, the greatest source of overall variation of habitat throughout landscapes. Many variables, such as landholder behaviour, complicate the relationship. For enhanced conservation outcomes the matrix needs to be managed to control disturbances and strategically plan for matrix habitat retention and restoration.  相似文献   

19.
Large-scale patterns of land use and fragmentation have been associatedwith the decline of many imperiled wildlife populations. Lesserprairie-chickens(Tympanuchus pallidicinctus) are restricted to thesouthernGreat Plains of North America, and their population and range have declined by> 90% over the past 100 years. Our objective was to examine scale-dependentrelationships between landscape structure and change and long-term populationtrends for lesser prairie-chicken populations in the southern Great Plains. Weused a geographic information system (GIS) to quantify landscape composition,pattern and change at multiple scales (extents) for fragmented agriculturallandscapes surrounding 10 lesser prairie-chicken leks. Trend analysis oflong-term population data was used to classify each population and landscape(declined, sustained). We analyzed metrics of landscape structure and changeusing a repeated measures analysis of variance to determine significant effects( = 0.10) between declining and sustained landscapes across multiplescales. Four metrics of landscape structure and change (landscape change index,percent cropland, increases in tree-dominated cover types, and changes in edgedensity) contained significant interactions between population status andscale,indicating different scaling effects on landscapes with declining and stablepopulations. Any single spatial scale that was evaluated would not have givencomplete results of the influences of landscape structure and change on lesserprairie-chicken populations. The smallest spatial scales (452, 905, and 1,810ha) predicted that changes in edge density and largest patch sizewere the only important variables, while large-scale analysis (7,238ha) suggested that the amount of cropland, increase in trees(mostly Juniperus virginiana), and general landscapechanges were most important. Changes in landscape structure over the pastseveral decades had stronger relationships with dynamics of lesserprairie-chicken populations than current landscape structure. Observed changessuggest that these local populations may be appropriately viewed from ametapopulation perspective and future conservation efforts should evaluateeffects of fragmentation on dispersal, colonization, and extinction patterns.This revised version was published online in May 2005 with corrections to the Cover Date.  相似文献   

20.
With return times between 20 and 100 years, ice storms are a primary disturbance type for temperate forests of eastern North America. Many studies have been conducted at the forest patch and plot scales to examine relations between damage and variables describing site, composition and structure. This paper presents results from a landscape scale study of fragmentation relations with damage in eastern Ontario forests. Data previously collected for two independent and spatially non-overlapping patch level damage studies were used. A Generalized Linear Model (GLM) was used to analyse relations between damage and fragmentation metrics representing patch isolation, edge density, and the relative size and distribution of patches in the landscape. The metrics were applied using spatial extents of 1 × 1 km and 4 × 4 km, following analyses of the variability of numbers of patches and of the lacunarity of forest patterns over a range of extents. The results showed that patch isolation, as measured by the mean Euclidean distance between patches (ENN) was significantly related to damage.  相似文献   

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