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1.
The effects of microhabitat (shrubs and herbs), plant litter, and seed burial on the regeneration of Liaodong oak (Quercus wutaishanica Mayr) and Chinese pine (Pinus tabulaeformis Carr.) were studied in three typical stands (Liaodong oak forest, Chinese pine plantation, and grassland) in the Loess Plateau, China. We monitored the establishment and growth of seedlings of these two woody species in sown experimental plots, in which shrubs and herbs, plant litter, and seed burial were manipulated. In the grassland, shrubs and herbs facilitated Liaodong oak establishment, with no effect on the establishment of Chinese pine. In the two forest stands, shrubs and herbs primarily had an inhibition effect on the establishment of these trees. The effects of plant litter were facilitation or inhibition, depending on the target species and the habitat. Seed burial had a positive effect on seedling establishment. In all three habitats, shrubs and herbs had inhibition effects on seedling growth of both tree species. Plant litter and seed burial did not influence seedling growth in either species. Liaodong oak and Chinese pine use different regeneration strategies during early stages of succession and similar strategies during late stages of succession.  相似文献   

2.
Otsamo  Riikka 《New Forests》2000,19(1):69-93
Natural regeneration of native tree species in the understoreys of fast-growing plantation tree species (Acacia mangium, Gmelina arborea and Paraserianthes falcataria) and in adjacent natural riverine forest and non-cultivated grassland areas were studied on Imperata cylindrica grassland sites in Riam Kiwa plantation area, South Kalimantan, Indonesia. Seedling and sapling densities as well as species composition and richness were assessed in a total of 35 ha by a systematic line plot survey.Floristic composition was similar in A. mangium and P. falcataria stands which differed distinctly from G. arborea stands and Imperata grassland area. Riverine forest was characterized by later successional species. In A. mangium and P. falcataria stands, a total of 63 species from 24 families were recorded. Seedling stock consisted merely of pioneer and secondary species, but also several fairly commonly occurring primary forest species. Euphorbiaceae was the most common family representing 42% of sapling and 29% of seedling densities. The most abundant regeneration occurred in A. mangium stands (4,706 seedling/sapling ha-1), followed by P. falcataria (2,242 ha-1) and G. arborea (554 ha-1). Seedling recruitment was significantly lower in plots dominated by grass vegetation than those dominated by no vegetation or shrub vegetation. Breast height diameter and basal area of canopy trees as well as coverage and height of ground vegetation did not explain seedling/sapling density. Results show that plantation establishment with suitable fast-growing tree species facilitates recruitment of a variety of native tree species. This provides potential for rapid restoration of forest ecosystem and for further development of the mixed stands of fast-growing exotics and naturally regenerated native species.  相似文献   

3.
To profile secondary succession after clearcutting a conifer plantation, we examined the composition of the soil seed bank and the process of vegetation recovery after clearcutting a mature Hinoki (Chamaecyparis obtusa) plantation in southwest Japan. We set up two study plots and collected 25 and 26 soil samples (each 20 cm × 30 cm × 5 cm depth), respectively, in April 1999, before clearcutting. Using the seedling emergence method, we detected 500–839 buried viable seeds m−2 in 42–44 species. Soil seed banks in both plots contained typical pioneer species but few late-successional species. After clearfelling the plantation in June 1999, we censused seedling establishment for 4 years and the composition of the vegetation for 6 years. More than half of the seedlings emerged in the second year, suggesting that the timing of clearcutting is possibly critical to seedling establishment. Nineteen plant species disappeared after clearcutting, 40 species were retained and 43 species newly emerged after clearcutting. Most of the newly emerged species were pioneer species which originated in the soil seed bank. Seven of the pioneer species, as typified by Rubus crataegifolius, showed clonal growth from the third year after clearcutting. Japanese serow browsed the seedlings of major pioneer species, possibly affecting the process of secondary succession. Late-successional species such as Abies firma declined in coverage or disappeared after clearcutting. Our results suggest that the soil seed bank makes a major contribution to vegetation recovery after clearcutting, but possibly not to restoration of the late-successional species.  相似文献   

4.
Abstract

Natural regeneration differences between oak and pine are common in the Qinling Mountains of central China. Thus, we examined whether this difference could be explained by seed dispersal. We investigated the seed dispersal of Quercus aliena var. acuteserrata Maxim. and Pinus tabulaeformis Carr. and the regeneration characteristics of both species were investigated in the pine-oak forest belt (oak stands, pine-oak stands, and pine stands) of Huoditang Forest in the Qinling Mountains in central China. We found that Q. aliena var. acuteserrata acorns were usually scatterhoarded, while seeds of P. tabulaeformis were generally consumed immediately. We found that 55% of the released acorns were transferred to the P. tabulaeformis stands. Moreover, Q. aliena var. acuteserrata regenerated successfully and was recruited in the three forest stands. However, the natural regeneration of P. tabulaeformis was very low in all stands because of high predation by small rodents. We propose, therefore, that the seed dispersal advantage of Q. aliena var. acuteserrata compared with P. tabulaeformis contributed greatly to the regeneration difference between these two species in the pine-oak forest belt of the Qinling Mountains in China.  相似文献   

5.
Senbeta  Feyera  Teketay  Demel  Näslund  Bert-Åke 《New Forests》2002,24(2):131-145
Regeneration of native woody species was studied in the plantations and the adjacent natural forest at Munessa-Shashemene Forest Project Area, Ethiopia. The aim of the study was to test the hypothesis that tree plantations foster regeneration of native woody species. A total of 60 plots, having 10 × 10 m area each, were studied in monoculture plantations of 4 exotic species (Cupressus lusitanica, Eucalyptus globulus, E. saligna, Pinus patula) and an adjacent natural forest. Ages of the plantations ranged between 9 and 28 years. Soil seed bank analysis was also undertaken from soil samples collected in each of the 60 plots to examine the similarity between the soil seed flora and aboveground vegetation. A total of 56 naturally regenerated woody species were recorded beneath all plantation stands with densities ranging between 2300 and 18650 individuals / ha in different stands. There was a significant difference among plantation stands with regard to understorey density (standard deviation: 4836 ± 1341). Vegetation diversity was assessed through analyses of floristic composition, species richness and abundance. Generally, seedling populations were the most abundant components of the regeneration in most of the plantation stands, forming 68 % of the total regeneration count in all stands. A total of 77 plant species represented by 44 herbs, 13 woody species, 8 grasses and 12 unidentified species were recorded in the soil seed bank from all stands. Similarity between the soil seed bank and aboveground flora was very low implying that the role of soil seed banks is negligible rather dispersal plays an important role in the process of regeneration. These results support the concept that forest plantations can foster the regeneration of native woody species, thereby increasing biological diversity, provided that there are seed sources in the vicinity of the plantations.  相似文献   

6.
An observational study of early seedling establishment (first 1–2 summers after emergence) was conducted in four ponderosa pine (Pinus ponderosa var. ponderosa Dougl. ex Laws.) stands east of the Cascade Range crest in central Oregon, USA. Newly emerged ponderosa pine seedlings were identified at the start of summer and were monitored through their second summer; a subsequent cohort of seedlings was identified and monitored for one summer. About 3% of the viable seed that was produced resulted in new seedlings. Mortality was substantial immediately following emergence. Most seedlings emerged without shading from understory vegetation, but few survived. Most seedlings alive after one year had emerged beneath live cover; by the end of two summers only shaded seedlings were alive. 63–85% of new seedlings initiated in clusters attributed to rodent caching, and seedlings originating in clusters remained a substantial proportion of the cohort (60%) after two seasons. Results suggest that understory vegetation in these types of stands facilitates the recruitment of ponderosa pine seedlings, most likely by buffering them against environmental stressors during the seedling establishment phase. They also reveal that seed caching rodents are highly active in seed redistribution and can exert a lasting influence on seedling recruitment.  相似文献   

7.
Regeneration by seeds for cork oak (Quercus suber) and companion oaks (holm oak Quercus ilex and downy oak Quercus pubescens) is likely to be poor in the fire-prone Maures massif (southern France) but the causes are poorly known. Our objective was to assess the effective recruitment for these three oak species and their temporal pattern of recruitment, in order to determine the main limitation factors and the regeneration window of each species. We studied oak recruits (height <3 m) in naturally regenerated populations according to a gradient of fire recurrence and in five main vegetation types including shrublands and mixed mature woodlands. Fire recurrence was the main explanatory factor of oak recruitment, either directly or through vegetation type and microsite characteristics. The results indicate nil to low recruitment for holm oak and downy oak in shrublands, especially those recurrently burned and dominated by Cistus species. Cork oak recruited better than the other oaks in medium and high shrublands dominated by Erica arborea. In contrast, recruitment was high for holm and downy oak in mixed oak stands and mixed pine-oak stands that have not burned for decades. Microsite conditions such as coverage by litter and shrubs influenced oak recruitment, whereas landscape configuration and stand basal area had no influence. Our results suggest that strategic shrub-clearing, oak planting and protection of mixed oak woodlands as seed sources would help maintaining oak populations in the woodland–shrubland mosaic.  相似文献   

8.
We investigated the seasonal patterns of soil phosphorus (P) fractions under five vegetation types – Ulmus macrocarpa savanna, grassland, Pinus sylvestris var. mongolica plantation, Pinus tabulaeformis plantation, and Populus simonii plantation – in the southeastern Keerqin Sandy Lands of China. The measured P fractions (0–20 cm depth) included: soil total P (TP), total organic and inorganic P (TPo and TPi), bicarbonate extractable organic and inorganic P (BPo and BPi), microbial biomass P (MBP), and in situ resin-adsorbed P (resin-P). Soil TP and TPo concentrations in the savanna and grassland were significantly lower in summer than in spring and autumn. However, they were relatively stable in three forest plantations. Soil labile P fractions showed a significant seasonal pattern under all vegetation types with the peak in summer, except soil MBP that was constant in the savanna and grassland and BPo that decreased over time in the savanna. This pattern of labile P fractions was attributed to a combination of seasonal climatic changes, low P availability, as well as the biological controls of soil P transformation in the study area. Litter decomposition played a key role in soil P availability. The monthly resin-P released from litter decomposition in summer was 2.6–7.4 times greater than in other seasons, and was 1.7–3.4 times of that in the 10 cm depth soil. Concentrations of soil P fractions were obviously affected by vegetation type. The savanna had the highest total P and MBP concentrations and the P. tabulaeformis plantation had the highest BPi and resin-P among all vegetation types. Among forest plantations, P. simonii plantation had the highest total P and MBP. These results suggest that U. macrocarpa savanna is the best system conserving soil nutrient (particularly P) stocks and microbial activity, followed by the grassland and P. simonii plantation, while the pine plantations are the worst.  相似文献   

9.
The purpose of this study was to examine the effect of habitat edges on the probability of acorn germination of three oak species (Quercus crassifolia Humb. and Bonpl., Quercus rugosa Née and Quercus laurina Humb. and Bonpl.). The effects of edge type (hard and soft), habitat type (grassland, edges and forest) and leaf litter cover (covered or uncovered acorns) on acorn germination was evaluated by the experimental establishment of acorns along transects crossing habitat edges. More acorns developed into seedlings in grasslands (38%) than in the forest edge (18%) or the forest interior (15%). In sites with soft edges, a higher number of seedlings emerged from acorns covered by litter compared with acorns sowed in the adjacent forest edge and forest interior (P < 0.05). In sites with hard edges, fewer seedlings emerged in the edge (14%) compared with the adjacent grassland (38%), and the adjacent forest (20%) presented intermediate values. However, in sites with soft edges significant differences in seedling emergence were recorded between the grassland (38%) and the forest (10%), whereas the edge presented intermediate values (23%). The effect of leaf litter cover on acorn germination was only significant in grasslands in sites with soft edges (P < 0.05). Acorns in grasslands received relatively little insect damage (10%) compared with the edge (19%) and the forest (30%, P < 0.05), emphasising the importance of acorn dispersal for successful oak establishment. The implications for regeneration of these oak species and the dynamics of montane oak forests in Mexico are discussed.  相似文献   

10.
Species composition and community structure of naturally regenerated trees in Larix kaempferi plantations and natural forests were compared in relation to altitudinal gradient. Fifty-nine L. kaempferi plantations and 26 natural forest stands including old-growth and secondary forests were selected from 1300 to 2000 m a.s.l. in Mt Kushigata of central Japan. Vegetation plots (10 × 10 m) were established in each stand (85 plots in total). Species composition differed significantly between forest types, particularly on high altitude plots. Some species were significantly biased to natural forests, while no species were biased to plantations. Although some species that responded significantly to altitude were common to both forest types (e.g., Acer ukurunduese, Castanea crenata, and Fraxinus lanuginose f. serrata), some differed between forest types (e.g., Fagus japonica, Prunus maximowiczii, and P. nipponica). Thus, plantation management altered species responses to altitude. Altitude should be considered when planning and carrying out forest ecological restoration in plantations, because species composition of naturally regenerated trees has already been changed by past plantation management. In particular, attention should be paid to ecological restoration at high altitude.  相似文献   

11.
对塞罕坝机械林场8个林分类型108个林班的地表森林可燃物组成、载量、含水率等进行了调查。结果表明落叶松成林和幼林、樟子松成林和幼林、针叶混交林、桦次生林等林分地表可燃物以枯落物为主,栎次生林和灌木林地表可燃物由枯落物与灌木组成,未成林地与草地可燃物以草本植物为主;地表可燃物载量依次为落叶松成林〉落叶松幼林〉针叶混交林〉樟子松成林〉樟子松未成林〉针阔混交林〉桦次生林〉樟子松幼林〉落叶松未成林〉灌木林〉栎次生林〉草地;草地与栎次生林的含水率最低,着火点也最低,火险危险程度最高;将现有林分按燃烧类型划分为5个类型20个亚类,全场地表可燃物总载量约为2．5×10^6t。  相似文献   

12.
Fruit size and seed moisture content were measured and seed production and dispersal were monitored to understand the seed biology of the Korean ash (Fraxinus rhynchophylla Hance) in two Korean ash forests. A combination of four ground treatments and four crown closure levels were created in a natural forest to understand the site conditions needed for seedling emergence in natural stands and to determine practical regeneration methods for direct seeding. The seed size of Korean ash increased until late May and its moisture content decreased rapidly in early November. Prolific seed bearing occurred every 3 yr. The distance of seed dispersal by wind was about 30 m, but more than 90% of the seeds dispersed within 10 m from the seed trees. Twenty-five seed trees per hectare provided for successful natural regeneration. The seedling emergence in the natural stands was best with scarification treatment and 25～50% of crown closure.  相似文献   

13.
Pinus tabulaeformis has been widely planted in order to conserve soil and water and improve the ecological environment in China. This study aimed to unravel how soil aggregates and soil carbon (C) stock stability of a P. tabulaeformis plantation change after 60 years of natural development and was performed in Vitex negundo var. heterophylla and Ziziphus jujuba var. spinosa shrub (shrub), a P. tabulaeformis forest (pine), and a coniferous broadleaf P. tabulaeformis mixed forest (pine-oak). Afforestation increased the stability of soil aggregates in the 0–10 cm soil layer but resulted in a decrease in the 10–20 cm soil layer. However, the presence of deciduous broadleaf species in the pine plantation improved the stability of soil aggregates. The total soil C stock was increased by afforestation, especially in the pine-oak forest, where it reached a significant level. The mineral soil C stock in the shrub stand was higher than that in pine and lower than that in pine-oak forests, but the C fractions had a different change. Afforestation increased the C fraction of macroaggregates in the 0–10 cm soil layer but decreased it in the 10–20 cm soil layer. This result suggested that afforestation could improve soil C stabilization in deeper soil. However, the pine-oak forest had a higher C fraction of macroaggregates than the pine forest in the 10–20 cm soil layer, indicating that soil C stabilization of the P. tabulaeformis plantation decreased when deciduous broadleaf species were present and thus formed the coniferous broadleaf mixed forest.  相似文献   

14.
We examined the regeneration and structure of mixed conifer forests under single-tree harvest management in western Bhutan. Sixteen 900 m2 (30 m × 30 m) plots were sampled at four Forest Management Units (FMUs; Chamgang, Gidakom, Paro-Zonglela, and Haa-East) representing the forest type, including half the plots in single-tree harvest stands and half in unlogged stands. In addition, we solicited information on traditional forest management practices from informants using survey questionnaires and collected tree species data from felling records from respective local forest offices. Rural timber demand is concentrated on the removal of straight and well-formed bluepine trees for beams, planks, and scaffolding. Single-tree harvest, however, has not significantly altered stand structures from unlogged stands. Similarly, tree regeneration is not different when comparing single-tree harvest and unlogged stands, except at Chamgang FMU, where seedling densities were generally higher in harvested stands than in unlogged stands. These results indicate that single-tree harvest is not detrimental to regeneration and utilization of mixed conifer forests in western Bhutan.  相似文献   

15.
The Huanglong Mountain forest zone is one of the major natural secondary forest zones in the southern Loess Plateau in Shaanxi Province, China. Since 1950, a mode of fully closed hillside afforestation (FHA) has been applied in the forest. On some special sites, the forest age exceeds 80 years. Pinus tabulaeformis forests form the most important vegetation cover in the warm temperate regions of China. Similarly, populations of P. tabulaeformis are dominant in existing forest ecosystems. Quercus liaotungensis, Syringa oblata, Populus davidiana, Prunus davidiana, Betula platyphylla and Toxicodendron vernicifluum can be occasionally found in the tree layer and shrub species are abundant. Based on the data collected from 31 plots and 93 soil samples, the state of health of the forest ecosystem is discussed and the appropriate FHA age has been determined. Twelve indices representing vegetation and soil properties in natural, secondary P. tabulaeformis forest ecosystems were generated by sensitivity analysis and an assessment index system for the FHA mode was established. According to the equal distance method, a clustering technique and five grades of an integrated index for evaluating the FHA mode were compartmentalized. The effect of the FHA mode on natural secondary P. tabulaeformis forests was evaluated by an integrated index method with the aid of an analytical hierarchy process (AHP). The results are as follows: values of the integrated index in the FHA mode of 16, 25, 30, 45, 60 and 75 year old stands were 7.25, 6.88, 7.82, 5.51, 4.78 and 2.79 respectively. With an increase over age of the FHA stands, the effect of the FHA mode deteriorated. We conclude that natural forests should not be protected in the FHA way after 45 years. At that stage, mixing suitable tree species, selection cutting and other silvicultural and management measures should be adopted. __________ Translated from Acta Ecologica Sinica, 2007, 27(1): 288–295 [译自: 生态学报]  相似文献   

16.
We investigated conifer plantation management in Japan using high-resolution airborne data based on an individual tree crown (ITC) approach. This study is the first to apply this technique to Japanese forests. We found that forest resources can be measured at the level of a single tree. We also produced a tree-crown map for a test site with Chamaecyparis obtusa, Pinus densiflora, Larix kaempferi, Cryptomeria japonica, other conifers, and broadleaved trees, with a classification accuracy of 78%. Forest-stand polygons with tree-cover types were generated from this map, a tree-density map, and a crown-occupied-area map. Forest information for the stand polygons was extracted automatically and compared with detailed field-survey data. The error between our ITC estimates and the field-survey data ranged from 0.3 to 30.2%, depending on tree crown size, density, and other factors. Errors were highest for high-density stands with mixed compositions and tree crown diameters ≤5.0 m. However, the error for stands with crown diameters ≥6.2 m was 11.6% or less. Therefore, this technique is best suited to pure Japanese conifer plantations without multiple layers or high-density stands.  相似文献   

17.
Tree seedling recruitment was monitored after various types of logging in mixed conifer and deciduous forests of northern British Columbia, Canada. Predicting tree seedling recruitment after disturbance is fundamental to understanding forest dynamics and succession and is vital for forest management purposes. Seedling recruitment success in multi-species northern latitude forests varied as a function of mature tree canopy cover, gap size and position in a gap. Recruitment was abundant within canopy gaps across a wide range of gap sizes (20–5000 m2), but recruit numbers dropped off rapidly under the closed forest canopy and in the open conditions of clearcuts. Inside canopy gaps, recruitment was similar by gap position in small gaps (<300 m2) but, in these northern latitude forests, exhibited a trend of increasing density from the sunny north to shady south end of larger gaps. This was true for all tree species regardless of their shade tolerance ranking. There was no evidence of gap partitioning by any of the tree species during the regeneration phase suggesting that adaptation to the subtleties of gap size during early recruitment are not well developed in these tree species. Favorable locations for emergence and early establishment of germinants were less favorable for growth and survival of established seedlings, i.e. the regeneration niches in these forests were discordant. Tree abundance and species diversity appears to be controlled more by differentiation among growth and survival niches than by the regeneration niches. From the perspective of forest management, abundant natural regeneration of all the dominant tree species of these mixed-species forests can be obtained after partial cutting.  相似文献   

18.
辽东山区长白落叶松天然更新调查   总被引：11，自引：0，他引：11  
调查辽东山区长白落叶松人工林的长白落叶松天然更新数量、幼苗的存活和生长状况以及其幼树出现的生境.结果表明:在结实的长白落叶松人工林内均有1年生落叶松幼苗,其数量与人工林种子库落叶松种子数量有关,其存活和生长(高生长和叶片数量)状况明显受林分开阔度、地面植被盖度和枯落物的影响.1年生长白落叶松幼苗在林内生长缓慢,一般高生长不超过6 cm,地径生长不超过0.1 cm,叶片数量不超过25针.长白落叶松天然更新幼苗4月末开始出现,6月份幼苗数量最多,9月份林内大部分幼苗死亡.这表明长白落叶松幼苗在林下不能正常存活和生长,而影响其存活和生长的主要因素是光照、草本灌木和地面枯落物.另外,长白落叶松林下很难见到幼树,只有在有种源、光照充足、草本灌木和枯落物少,以及土壤较湿润的生境中能见到其幼树.  相似文献   

19.
The relationships between seed size, emergence, stand establishment and seedling height were examined in a trial comparing tree and shrub species as hedgerows for alley cropping in four environments in Haiti. Logarithmic functions relating % emergence, % stand establishment and seedling height to seed weight were positive and significant. These relationships were stronger when species weighing > 10 g 100−1 seed were excluded from the analysis. Higher R2 values were obtained regressing % stand than regressing % emergence on seed weight. Although exceptions were noted, a majority of species having seed weights between 2.5 g and 10 g 100−1 had good stands. Establishment was poor or inconsistent for species with seed < 2.5 g 100−1. The effect of seed size on height was more important for seed weights < 5 or 8.8 g 100−1, depending upon the site, than for larger seed. Species with < 2.5 g 100−1 seed are unsuitable for direct seeding where the chance of drought is high.  相似文献   

20.

Seed properties and seedbed conditions influence forest regeneration following direct seeding. It might therefore be possible to improve result by improving seed quality and seedbed conditions. In a direct seeding experiment designed to determine the effects of seed invigoration (incubation at 30% moisture content for 7 days) and microsite preparation (2-cm-deep pyramidal indentations) on the regeneration, sowing was repeated on eight different occasions, and seedling emergence, survival and height growth were recorded. Seedling emergence, mainly after early sowing, was improved by invigoration when used in combination with microsite preparation. Both survival and growth were impaired by delaying sowing until later in the growing season, but were not significantly affected by invigoration or microsite preparation. It is concluded that an improved and less varying regeneration result after direct sowing may be achieved by using invigorated seeds in combination with microsite preparation.  相似文献   

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