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1.
Many ecosystems are now dominated by introduced species, and because dominant species drive ecosystem properties, these changes lead to increased uncertainty in estimates of carbon storage and cycling. We examined aboveground biomass in forests dominated by the introduced tree Rhamnus cathartica (common buckthorn) relative to forests dominated by native species, and measured aboveground biomass increment over a three-year period (2005-2008). Three of the four lowest biomass levels occurred in R. cathartica-dominated forests, and biomass in these forest types was stored primarily in trees 10-20 cm DBH. By contrast, forests dominated by native trees (including those with R. cathartica understories) had the six highest biomass levels, and biomass was stored primarily in trees >50 cm DBH. On average, forests dominated by R. cathartica stored half as much aboveground biomass (14.6 ± 3.3 kg/m2) as forests dominated by native tree species (28.9 ± 8.3 kg/m2). R. cathartica-dominated forests also had half the aboveground biomass increment of native-dominated forests (0.28 vs. 0.60 kg/m2/year). Although known anecdotally as a fast-growing species, R. cathartica growth rates declined with increasing size. Between 2005 and 2008, R. cathartica individuals <10 cm DBH grew faster than native species; however, R. cathartica individuals >10 cm DBH grew consistently slower than native species. Overall, our findings indicate that intrinsic size limitations on R. cathartica will lead to lower biomass stocks in forests where it acts as a canopy dominant relative to forests dominated by native tree species.  相似文献   

2.
Ficus insipida Willd. (Moraceae) is a fast growing tree species of early successional stages in the Amazonian nutrient-rich white-water floodplains (várzea). The species is one of the most economically important low-density wood species in the community-based forest management project in the Mamirauá Sustainable Development Reserve (MSDR) in Central Amazonia, where timber species are managed using a polycyclic selection system with a minimum logging diameter (MLD) of 50 cm and a cutting cycle of 25 years. In this study we analyze the floristic composition, stand structure and forest regeneration of a natural 20 year-old stand at an early successional stage and we model tree growth of diameter, height and volume of F. inspida based on tree-ring analysis to define management criteria. The volume growth model indicates that the preferred period for logging should be at a tree age of 17 years when the current annual volume increment peaks. This age corresponds to a diameter of 55 cm, which would be an appropriate MLD.  相似文献   

3.
Long-term tree and seedling growth and survivorship data from permanent sample plots established in a neotropical dry forest in Jamaica from 1998 to 2008 were used to (1) model growth (periodic annual increment) and survivorship dynamics, (2) cluster structural and functional types, and (3) estimate the age of selected tropical dry forest tree species. A statistical comparison of parameter estimates derived from a generalized linear model (GLM) of each species to a reference species was used to group individuals based on size (DBH and height), and demographic dynamics (periodic annual increment and survivorship). We identified two groups of species based on structural types (canopy and sub-canopy species), three groups of species based on periodic annual increment (fast, intermediate, and slow growing) and four groups of species based on the probability of survivorship of seedlings and trees (very low probability of seedling survivorship but high tree survivorship (two groups); high survivorship throughout the DBH classes; very low survivorship, regardless of stem size). The composition of the groups was mixed, and included individuals of both structural types, and with different periodic annual increment and survival probabilities. The dichotomy of guilds found in tropical rainforests (pioneer and climax species) was not found in this forest. Individual and group GLMs incorporating empirical relationships between periodic annual increment and survivorship, across a spectrum of ontogenies and DBH’s, were also generated. The periodic annual increment models were then used to estimate the time taken by a newly germinated seedling to reach the largest recorded DBH. The fastest growing species was the hemi-cryptophyte Clusia flava which was estimated to take 74 years to reach its maximum recorded size (12.1 cm DBH), whereas the slowest growing species, Ziziphus sarcomphalus, was estimated to take 399 years to reach its maximum size (24.4 cm DBH). These dry forest trees were estimated to reach their maximum size (which was one-half or one-third of the largest DBH recorded for tropical rainforests) in a time similar to tropical rainforest trees. Some of the tree species are ubiquitous to other neotropical dry forests; therefore, our equations for periodic annual increment and survivorship can be applied elsewhere in the region.  相似文献   

4.
Against a background of increasing human populations in developing countries, and global climate change, conservation of tropical forests remains one of the most important ecological challenges of our time. One of the biggest difficulties for ecologically sustainable management of tropical forests is obtaining reliable growth data for trees, which is a prerequisite for determining harvesting volumes and cutting cycles. GOL is the first concept for sustainable management of tropical timber resources in Amazonian floodplain forests (várzea) based on species-specific management criteria, such as minimum logging diameters (MLDs) and cutting cycles. From timber species with varying wood densities of different successional stages, volume stocks have been estimated in 1-ha plots and 12 growth models have been developed based on tree rings, which are annually formed as a consequence of the regular, long-term flooding. The MLDs of timber species vary between 47 and 70 cm and the estimated cutting cycles differ the 10-fold, from 3 to 32 years. These enormous differences in the growth rates between tropical timber species are not considered in current management practices, which apply only one diameter cutting limit and one cutting cycle to harvest many tree species. This practice risks the overexploitation of slow-growing timber species, while the fast-growing timber species with low wood densities cannot be efficiently used. Based on the timber stocks and lifetime growth rates, the GOL concept has been created as an aid to improve forest management in the Central Amazonian várzea. The model is unique for tropical silviculture.  相似文献   

5.
Large tree species have a disproportional influence on the structure and functioning of tropical forests, but the forces affecting their long-term persistence in human-dominated landscapes remain poorly understood. Here we test the hypothesis that aging forest edges and small fragments (3.4–295.7 ha) are greatly impoverished in terms of species richness and abundance of large trees in comparison to core areas of forest interior. The study was conducted in a hyper-fragmented landscape of the Atlantic forest, northeast Brazil. Large tree species were quantified by recording all trees (DBH ≥ 10 cm) within fifty-eight 0.1-ha plots distributed in three forest habitats: small forest fragments (n = 28), forest edges (n = 10), and primary forest interior areas within an exceptional large forest remnant (n = 20). Large tree species and their stems ≥10 cm DBH were reduced by half in forest edges and fragments. Moreover, these edge-affected habitats almost lacked large-stemmed trees altogether (0.24 ± 0.27% of all stems sampled), and very tall trees were completely absent from forest edges. In contrast, large trees contributed to over 1.5% of the whole stand in forest interior plots (2.9 ± 2.8%). Habitats also differed in terms of tree architecture: relative to their DBH trees were on average 30% shorter in small fragments and forest edges. Finally, an indicator species analysis yielded an ecological group of 12 large tree species that were significantly associated with forest interior plots, but were completely missing from edge-affected habitats. Our results suggest a persistent and substantial impoverishment of the large-tree stand, including the structural collapse of forest emergent layer, in aging, hyper-fragmented landscapes.  相似文献   

6.
Stand-level tree diameter growth patterns were explored for evergreen moist forests in the southern Cape, South Africa. Results of standard multiple regression analyses, involving 934 permanent sample plots with data spanning a 10-year interval, revealed that stand-level increment of canopy species in the canopy layer (>30 cm dbh) was significantly determined by inherent species-specific growth capacities (species composition of the stand), water availability, forest matrix crowding and tree condition impairment (age-related manifestations of reduced vitality indicated by signs of crown die-back, damage and stem rot). In contrast, stand-level increment of trees of canopy species in the subcanopy layer (10-20 cm dbh) was prominently shaped by light availability, as mainly determined by the degree of canopy-level disturbance (mortality rate of trees >30 cm dbh), crowding (canopy-level overhead and forest matrix crowding) and proximity to conspecific adults (within 6-8 m). In addition to species-inherent and resource factors, considerable variation in stand-level growth resulted from site-climate interactions. For 507 of the permanent sample plots, increment data was available for two consecutive 10-year intervals; permitting the analysis of spatiotemporal interactions of growth patterns (repeated measures ANOVA). In the Knysna forests higher canopy-level increment rates were associated with the moister southerly facing slope sites in comparison with the drier northerly facing and ridge sites during the first increment period. During the second increment period, increment rates on the drier, but better illuminated sites had increased disproportionately. In contrast, in the Tsitsikamma forests, higher increment rates during the second increment period were encountered on moister flat bottomland sites (with extended periods of subsoil wetness) than on the comparatively drier southerly facing slope sites (increment period × site-based water availability × forests interaction). In both forests relatively higher growth performance of subcanopy-level trees during the second increment period was associated with stands experiencing conditions of enhanced light availability. Atmospheric temperatures were higher during the second increment period (mean periodic Tmax: + 0.64 °C). The detected spatiotemporal interactions were interpreted as site × climate interactions where site-related conditions of favourable light or water availability resulted in enhanced temperature-linked growth responses during the second increment period. A metabolic performance trade-off model provided a framework for the interpretation of these complex site-climate interactions by placing the patterns of forest growth into an ecophysiological explanatory context.  相似文献   

7.
This study examined riparian forest and instream large wood characteristics in a 2.7 km reach of the West Branch of the Sheepscot River in Maine in order to increase our basic knowledge of these components in a system that is known to have undergone multiple land conversion. The West Branch is approximately 40 km long, drains a 132 km2 watershed and is vitally important to the remnant population of Atlantic salmon (Salmo salar) and other native species. The riparian forest is comprised of relatively small trees with a mean DBH of 21 cm (SD ± 10.92) with 56% of the trees having a DBH <20 cm. Balsam fir (Abies balsamea) and red maple (Acer rubrum) are the most common species (54%), and 75% of all trees are short-lived, small diameter species. These data suggest the riparian forest in the West Branch Sheepscot River is dominated by young forest stands, a legacy of land use. During a survey conducted in 2005, 210 pieces of large woody debris (LWD) were identified in the study reach; an average of 78 pieces km−1. The total volume of pieces was 8.5 m3 or 3.2 m3 km−1 (LWD in this study is defined as pieces ≥10 cm in diameter and >2 m in length). The mean diameter of LWD was 17 cm with 75% of all pieces having a diameter <20 cm. Most pieces were oriented parallel or nearly parallel to the channel and did not appear to influence channel morphology. In contrast, larger pieces were more often in perpendicular or nearly perpendicular orientations, and were more likely to have a pool-forming function. Overall, the reach has low levels of stable large wood, which do not have a major influence on stream habitats.  相似文献   

8.
Selective logging is an important socio-economic activity in the Congo Basin but one with associated environmental costs, some of which are avoidable through the use of reduced-impact logging (RIL) practices. With increased global concerns about biodiversity losses and emissions of carbon from forest in the region, more information is needed about the effects of logging on forest structure, composition, and carbon balance. We assessed the consequences of low-intensity RIL on above-ground biomass and tree species richness in a 50 ha area in northwestern Gabon. We assessed logging impacts principally in 10 randomly located 1-ha plots in which all trees ?10 cm dbh were measured, identified to species, marked, and tagged prior to harvesting. After logging, damage to these trees was recorded as being due to felling or skidding (i.e., log yarding) and skid trails were mapped in the entire 50-ha study area. Allometric equations based on tree diameter and wood density were used to transform tree diameter into biomass.Logging was light with only 0.82 trees (8.11 m3) per hectare extracted. For each tree felled, an average of 11 trees ?10 cm dbh suffered crown, bole, or root damage. Skid trails covered 2.8% of the soil surface and skidding logs to the roadside caused damage to an average of 15.6 trees ?10 cm dbh per hectare. No effect of logging was observed on tree species richness and pre-logging above-ground forest biomass (420.4 Mg ha−1) declined by only 8.1% (34.2 Mg ha−1). We conclude from these data that with harvest planning, worker training in RIL techniques, and low logging intensities, substantial carbon stocks and tree species richness were retained in this selectively logged forest in Gabon.  相似文献   

9.
Dead wood is an important component of forest ecosystems and volumes vary depending on forest age, management intensity and productivity. This is the first large-scale study to quantify dead wood in Irish forests and to compare them to forests in other locations. We measured the volume and size distribution of logs, the density and size distribution of snags and the volume of dead wood contained in stumps in Oak (Quercus spp.) and Ash (Fraxinus excelsior) forests and in Sitka spruce (Picea sitchensis) plantations throughout Ireland. We also assigned each log, snag and stump to one of three decay classes (intact, part-rotted and well-rotted). We found no significant difference in log volume between any of the forest types. The majority (>90%) of logs were less than 20 cm in diameter, and large logs (>40 cm diameter) were scarce. We found a relatively high density of snags in all forest types but, as in the case of logs, over 90% of snags were <20 cm DBH and large snags (>40 cm DBH) were rare. The volume of dead wood contained in stumps was significantly higher in plantations than in Oak or Ash forests as a result of thinning and harvesting. Most logs and snags were moderately decayed but, in plantations, most stumps were intact. Log volume and the size of logs and snags were considerably lower than in old-growth forests in other regions. These patterns may reflect historical use of Irish forests for coppice and timber production. Management for biodiversity should aim to accelerate dead wood accumulation to increase the frequency of large-diameter logs and snags. Although management seeking to replicate the dead wood volumes of old-growth forests is ideal, it may be unrealistic in the short term.  相似文献   

10.
To improve the silvicultural targets for ecologically sustainable forestry, we quantified functionally important structural features for the first time in a representative set of old-growth forests in hemiboreal Europe. Altogether, 23 old-growth stands of four site-type groups were compared with mature commercial stands nearby in the Estonian state forests that hold the Forest Stewardship Council (FSC) certificate of sustainable forestry. These two treatments did not differ significantly in terms of tree-species diversity, volumes of woody debris of <20 cm diameter (including fine woody debris) and its decay-stage composition. However, mature stands had many more early-successional trees and lacked late-successional deciduous species; they also had a higher overall density and volume of live trees, due to abundant individuals of 10–39 cm diameter at breast height. Old-growth stands had at least twice as many live trees ≥40 cm, standing dead trees ≥30 cm and lying wood ≥20 cm in diameter, any freshly fallen debris, and regeneration. For lying wood ≥20 cm in diameter, the treatment effect depended on site type: both treatments of Vaccinium-type dry boreal forests were remarkably deadwood-poor (indicating historical management of the old-growth stands), while mature eutrophic stands of Aegopodium-type were most impoverished relative to old-growth levels. We conclude that many functional characteristics of old growth were present in the FSC-certified, mostly naturally regenerated, commercial stands. The main problem is the lack of very large trees, particularly of late-successional deciduous species, which should be addressed by their well-planned retention in cut areas and reconsideration of salvage logging strategies. A dense regeneration in old-growth stands also indicated the potential of selection cuttings. The study highlighted the need for region- and site-type specific numerical targets for sustainable forest management, which in the hemiboreal region should address the characteristic occurrence of late-successional deciduous trees on fertile soils and higher natural deadwood volumes than in typical boreal forests. For certification, the issues of structural impoverishment revealed both the inadequacy of some silvicultural practices and some indicators set by the national FSC-standard in Estonia.  相似文献   

11.
Allometric equations were developed and applied to forest inventory data to estimate biomass and carbon stocks for temperate species and forests of Durango and Chihuahua and for tropical dry forests of Sinaloa, Mexico. A total of 872 trees were harvested and dissected into their component parts: leaves and branches, boles, and coarse roots. Coarse roots of 40 temperate trees ranging in diameter at breast height (DBH) from 6.0 to 52.9 cm were excavated in their entirety (i.e., >0.5 cm diameter). The species sampled (number of trees) in tropical dry forests (39) were Lysiloma divaricata (Jacq) Macbr. (10), Haematoxylon brasiletto Karst. (10), Cochlospermum vitifolium (Wild.) (5), Ceiba acuminata (S. Watson) Rose (5), Bursera penicillata (B. inopinnata) (5), and Jatropha angustifolia Mull. Arg. (4) and in temperate forests (833) were Quercus spp. (118) (Q. rugosa Neé, 15, Quercus sideroxylla Humb. & Bonpl, 51, Quercus spp., 52), Pinus herrerae Martinez 1940 (19), Pinus oocarpa Schiede ex Schlectendal 1838 (31), Pinus engelmannii Carriere 1854 (7), Psudotsuga menziesii (Mirb.) Franco (19), Pinus leiophylla Schiede ex Schlectendal et Chamisso 1831 (27), Pinus teocote Schiede ex Schlectendal et Chamisso (55), Pinus ayacahuite Ehrenb. ex Schltdl. (58), Pinus cooperi Blanco (48), Pinus durangensis Martinez 1942 (385), and Pinus arizonica Engelmann 1879 (66). Allometric equations having only DBH as an independent variable were developed for each component of each species. Since Pinus herrerae, Pinus engelmannii, Pinus oocarpa and Pseudotsuga menziensii had a small number of trees, an individual allometric equation was developed for these species. We used non-linear regression to fit parameters of the typical allometric power equation. The resulting 31 equations (10 species or groups of species, three biomass components; bole, branch and leaves, and total aerial; and the generalized equation for coarse roots) fit the data well and enable the user to predict biomass by component for each of the 10 different groups of species or each of six temperate species. A single allometric equation that incorporates the basic specific gravity for aboveground biomass of all temperate tree species also fit the data well, and this equation provides both the detail and the accuracy supplied by species-specific, plant-part-specific equations. Biomass equations coupled with forest inventory data for temperate (637 circular, 1/10 ha plots) and tropical dry forests (166 20 m × 20 m-quadrats) of northwestern Mexico predict a mean (confidence intervals) of 130 Mg ha−1 (4.2 Mg ha−1) and 73 Mg ha−1 (7.1 Mg ha−1) for total tree and total aboveground biomass, respectively. Large sample sizes and the economic and ecological importance of the species studied make this data set uniquely useful for biomass estimations and for understanding the inherent heterogeneity of tree structure in dynamic tropical and temperate environments of northwestern Mexico.  相似文献   

12.
Oriental Beech is the most important commercial tree species in northern Iran. In recent years wood production companies interested in felling large beech trees for profit have challenged advocates of close-to-nature silviculture who favor conservation. Our study objective was to assess the economic value of over-mature beech trees by relating tree diameter (DBH) to amount of decay. Based on the location of onset of decay, we categorized three types of decay as stump, stem, and crown decay. Trees of greater diameter (age) typically showed greater decay in the stem. Percent of decayed volume, diameter of decayed tissue, and length of decay in tree stems varied between 0.5%?64.3%, 15 cm?75 cm, and 2.0?19.5 m, respectively. With increasing trunk diameter, the proportion of truck decay increased. Red heart and dark red heart constituted 25% and 14.3% of sampled trees, respectively. However, we found no correlation between intensity of stem decay and morphological characteristics of trees. Seedlings were not abundant around the bases of over-mature trees, suggesting that the trees did not contribute to regeneration of the stand. Beech trees of diameter >1 m do not provide valuable round wood for industries and cause to raise wood production costs. We recommend that these trees >1 m DBH should be retained in forest stands because of their low commercial value but high ecological and conservational values such as maintaining biodiversity in forest ecosys-tems.  相似文献   

13.
The aim of this study was to develop and test a new basal area growth model in mixed species continuous cover forests in northern Iran.Weanalyzed 421 core samples from 6 main species in the forest area to develop our growth model.In each plot,we measured variables such as total tree height(m),diameter at breast height(DBH)(cm)and basal area of larger trees as cumulative basal areas of trees(GCUM)ofDBH[5 cm.The empirical data were analyzed using regression analysis.There was a statistically significant nonlinear function between the annual basal area increment,as the dependent variable,and the basal area of the individual trees and competition as explanatory variables.Reference area from the largest trees,was circular plot with area of 0.1 ha.GCUM was estimated for trees of DBH>5 cm.Furthermore,we investigated the dependencies of diameter growth of different species on stand density at different levels of competition,and diameter development of individual trees through time.The results indicate that competition caused by larger neighborhood trees has a negative effect on growth.In addition,the maximum diameter increment is affected by competition level.Therefore,the maximum diameter increment of species occurs when the trees are about 35–40 cm in dense-forest(40 to 0 m^2 per ha)and when the trees are about 60 to 70 cm in very dense forest(60 to 0 m^2 per ha)which is more likely to Caspian natural forests with high level density due to uneven-aged composition of stands.  相似文献   

14.
Developing sustainable extractive industries in otherwise intact tropical forest regions requires a sound understanding of the production potential of key resource populations. The oleoresin extracted from Copaifera trees is an economically important non-timber forest product harvested throughout the lowland Amazon basin. We studied oleoresin extraction from four species of Copaifera trees with known harvest histories within two contiguous extractive reserves in western Brazilian Amazonia. We conducted a large-scale experimental harvest of 179 previously unharvested Copaifera trees, in both seasonally flooded (várzea) and adjacent unflooded (terra firme) forests. The likelihood of trees yielding any oleoresin was principally determined by their species identity: C. multijuga was the only species to regularly yield oleoresin (70% of trees). Yield volumes varied both amongst species and forest types: C. multijuga (restricted to terra firme forest) had the highest mean yield of 505 ml, whilst C. guyanensis produced higher volumes of oleoresin in várzea (139 ml) than terra firme (15 ml) forest. Intraspecific differences were driven mainly by tree size. To assess extraction sustainability, we reharvested a sample of C. multijuga trees and compared the oleoresin production of 24 conspecific trees that had been initially harvested one year previously with that of 17 trees initially harvested three years previously. Reharvested trees produced just 35% of the oleoresin volume compared to that when originally drilled, but this response was not affected by the time interval between consecutive harvests. We demonstrate that, within a population of Copaifera, both morphological and environmental factors restrict total productivity; consideration of these factors should inform sustainable management practises. We additionally raise methodological considerations that may improve the comparability of studies.  相似文献   

15.
Allometric equations can be used to estimate the biomass and carbon stock of forests. However, so far the equations for Dipterocarp forests have not been developed in sufficient detail. In this research, allometric equations are presented based on the genera of commercial species and mixed species. Separate equations are developed for the Dipterocarpus, Hopea, Palaquium and Shorea genera, and an equation of a mix of these genera represents commercial species. The mixed species is constructed from commercial and non-commercial species. The data were collected in lowland mixed Dipterocarp forests in East Kalimantan, Indonesia. The number of trees sampled in this research was 122, with diameters (1.30 m or above buttresses) ranging from 6 to 200 cm. Destructive sampling was used to collect the samples where diameter at breast height (DBH), commercial bole height (CBH), and wood density were used as predictors for dry weight of total above-ground biomass (TAGB). Model comparison and selection were based on Akaike Information Criterion (AIC), slope coefficient of the regression, average deviation, confidence interval (CI) of the mean, paired t-test. Based on these statistical indicators, the most suitable model is ln(TAGB) = c + αln(DBH). This model uses only a single predictor of DBH and produces a range of prediction values closer to the upper and lower limits of the observed mean. Model 1 is reliable for forest managers to estimate above-ground biomass, so the research findings can be extrapolated for managing forests related to carbon balance. Additional explanatory variables such as CBH do not really increase the indicators’ goodness of fit for the equation. An alternative model to incorporate wood density must be considered for estimating the above-ground biomass for mixed species. Comparing the presented equations to previously published data shows that these local species-specific and generic equations differ substantially from previously published equations and that site specific equations must be considered to get a better estimation of biomass. Based on the average deviation and the range of CI, the generalized equations are not sufficient to estimate the biomass for a certain type of forests, such as lowland Dipterocarp forests. The research findings are new for Dipterocarp forests, so they complement the previous research as well as the methodology of the Good Practice Guidance for Land Use and Land Use Change and Forestry (GPG-LULUCF).  相似文献   

16.
This paper estimates the difference in stand biomass due to shorter and lighter trees in southwest (SW) and southern Amazonia (SA) compared to trees in dense forests in central Amazonia (CA). Forest biomass values used to estimate carbon emissions from deforestation throughout, Brazilian Amazonia will be affected by any differences between CA forests and those in the “arc of deforestation” where clearing activity is concentrated along the southern edge of the Amazon forest. At 12 sites (in the Brazilian states of Amazonas, Acre, Mato Grosso and Pará) 763 trees were felled and measurements were made of total height and of stem diameter. In CA dense forest, trees are taller at any given diameter than those in SW bamboo-dominated open, SW bamboo-free dense forest and SA open forests. Compared to CA, the three forest types in the arc of deforestation occur on more fertile soils, experience a longer dry season and/or are disturbed by climbing bamboos that cause frequent crown damage. Observed relationships between diameter and height were consistent with the argument that allometric scaling exponents vary in forests on different substrates or with different levels of natural disturbance. Using biomass equations based only on diameter, the reductions in stand biomass due to shorter tree height alone were 11.0, 6.2 and 3.6%, respectively, in the three forest types in the arc of deforestation. A prior study had shown these forest types to have less dense wood than CA dense forest. When tree height and wood density effects were considered jointly, total downward corrections to estimates of stand biomass were 39, 22 and 16%, respectively. Downward corrections to biomass in these forests were 76 Mg ha−1 (∼21.5 Mg ha−1 from the height effect alone), 65 Mg ha−1 (18.5 Mg ha−1 from height), and 45 Mg. ha−1 (10.3 Mg ha−1 from height). Hence, biomass stock and carbon emissions are overestimated when allometric relationships from dense forest are applied to SW or SA forest types. Biomass and emissions estimates in Brazil's National Communication under the United Nations Framework Convention on Climate Change require downward corrections for both wood density and tree height.  相似文献   

17.
Secondary evergreen broadleaved forests are precious remnants for biodiversity conservation and templates for sustainable management of natural forests in subtropical China. Floristic composition, size structure, and spatial pattern of dominant tree species have been investigated for a subtropical secondary evergreen broadleaved forest in the Huitong Yingzuijie National Forest Reserve, Hunan, China. The location of all trees greater than 4 cm in diameter at breast height (DBH) were mapped within a 0.96-ha plot in which species, DBH, and total tree height were recorded. Ripley’s K(t) function was used to analyze spatial patterns and associations. The secondary forest consisted of 74 tree species and 1,596 stems per hectare. A reverse-J shaped DBH classes distribution was observed for all stems and trees of later seral species whereas trees of earlier successional species were distributed irregularly. Significant aggregated spatial patterns were observed for all trees within the forest and for conspecific trees of each dominant species. This result, and a repulsive spatial pattern for interspecific trees of Choerospondias axillaries and Cyclobalanopsis glauca against other dominant tree species, support segregation hypothesis. Contributions of seed dispersal, topographic heterogeneity, and competition to spatial patterns of conspecific trees vary depending on tree species. Attractive spatial pattern among interspecific trees of Liquidambar fortunei, Liquidambar formosana, and Pinus massoniana reflects stochastic colonization of pioneer tree species and a facilitation relationship. Although deciduous species are long-lived and persist over long successional processes, they will eventually be replaced by late seral evergreen species within the secondary forest if no disturbance events occur.  相似文献   

18.
In the climate change discussion, the possibility of carbon sequestration of forests plays an important role. Therefore, research on the effects of environmental changes on net primary productivity is interesting. In this study we investigated the influence of changing temperature, precipitation and deposition of sulphur and nitrogen compounds on forest growth. The database consisted of 654 plots of the European intensive monitoring program (Level II plots) with 5-year growth data for the period 1994–1999. Among these 654 plots only 382 plots in 18 European countries met the requirements necessary to be used in our analysis. Our analysis was done for common beech (Fagus sylvatica), oak (Quercus petraea and Q. robur), Scots pine (Pinus sylvestris) and Norway spruce (Picea abies). We developed an individual tree growth model with measured basal area increment of each individual tree as responding growth factor and tree size (diameter at breast height), tree competition (basal area of larger trees and stand density index), site factors (soil C/N ratio, temperature), and environmental factors (temperature change compared to long-term average, nitrogen and sulphur deposition) as influencing parameters. Using a mixed model approach, all models for the tree species show a high goodness of fit with Pseudo-R2 between 0.33 and 0.44. Diameter at breast height and basal area of larger trees were highly influential variables in all models. Increasing temperature shows a positive effect on growth for all species except Norway spruce. Nitrogen deposition shows a positive impact on growth for all four species. This influence was significant with p < 0.05 for all species except common beech. For beech the effect was nearly significant (p = 0.077). An increase of 1 kg N ha−1 yr−1 corresponds to an increase in basal area increment between 1.20% and 1.49% depending on species. Considering an average total carbon uptake for European forests near 1730 kg per hectare and year, this implies an estimated sequestration of approximately 21–26 kg carbon per kg nitrogen deposition.  相似文献   

19.
Producing high value veneer wood requires that the tree bole be branch-free. This can be accomplished by natural or artificial pruning. Since wild cherry does not self prune well, pruning artificially is the only practical option. The study analysed the effect of conventional whorl-wise pruning and selective pruning, on height growth, diameter growth and secondary shoot development of wild cherry. Four pruning treatments were applied on cherry trees in summer 2007, one group of cherries was left unpruned to serve as a control: treatment C1 (upper 5 whorls left), C2 (upper 3 whorls left), S1 (removal of branches larger than 3 cm or with an angle to the stem < 40°), S2 (removal of branches larger than 2 cm or with an angle to the stem < 40°), N (unpruned). Data showed that height growth was not affected by pruning. In contrast, diameter growth at breast height of the C2 pruned cherry was reduced by approximately 5% (SE = 2.7%) in the year of pruning (trees were pruned in July). This pruning treatment produced significant (p = 0.028) nine percent less diameter growth than the control in the second year following pruning. The diameter increment of the C1 pruned trees with five whorls left after pruning and the selective pruned cherries were only about 4% (SE = 4.0%) smaller than the control after two years. This loss was statistically not significant. Analyses showed that on selective pruned trees the survival rate of secondary shoots was significantly reduced compared to those on whorl-wise pruned trees. Significant differences in the size of the secondary shoots were only found between the C1 and S1 (p < 0.05) pruned trees. We did not find differences in the total number of secondary shoots per tree among pruning treatments. Solely from a tree growth perspective, the moderate whorl-wise pruning treatment C1 and the selective prunings were equally effective in minimizing the reduction of diameter growth and are recommended in practice. However it was found that the survival of secondary shoots was reduced on selective pruned trees although the amount of pruning work needed in selective pruning was slightly greater than conventional moderate pruning.  相似文献   

20.
Woodpeckers, able to excavate holes in trees, can provide resources critical for non-excavator hole users. Supply of woodpecker-made holes in forests depends on excavation rates by the birds and holes’ persistence times. I use 30 years of data from a primeval forest (strictly protected reserve, Bia?owie?a National Park, E Poland) to determine how long woodpecker-made holes persist, and whether their persistence varies across forest types, tree species and conditions, and woodpecker species. I followed the fate of 719 breeding holes, excavated by eight woodpecker species, for up to 27 years, from 1979 to 2010. Almost 80% of hole losses were caused by collapse of either the tree or the section supporting the hole. Holes were retained for (median) 6-7 years in riverine and oak-hornbeam forest but 10 years in coniferous forest. These differences can be explained by almost completely non-overlapping sets of tree species used in these different habitats. Lifespan of holes varied by tree species, ranging from four (Picea abies) to >22 years (Pinus sylvestris, almost 100% dead). The long lifespan of holes in the dead Pinus was exceptional, as otherwise, persistence was much lower for holes excavated in dead trees or limbs (5 years) than for those in living substrates (9 years). Tree species with higher frequency of holes in dead wood showed lower persistence times of holes. Lifespans of holes excavated by individual woodpecker species varied widely and was strongly dependent on frequency with which the species excavated in dead wood. Holes of Dendrocopos minor and Dendrocopos leucotos (only in dead wood) persisted for four years, while holes of Dendrocopos major (able to excavate in living sapwood of some trees) lasted for nine, and those of Dryocopus martius for 18 years. Retention of dead P. sylvestris, decaying Quercus robur in stands and addition/retention of aspens (Populus tremula and Populus tremuloides) in them would provide conditions to increase the availability of relatively persistent woodpecker holes in forests of the Northern hemisphere.  相似文献   

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