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1.
We applied a physiological individual‐based model for the foraging and growth of cod (Gadus morhua) and haddock (Melanogrammus aeglefinus) larvae, using observed temperature and prey fields data from the Irish Sea, collected during the 2006 spawning season. We used the model to estimate larval growth and survival and explore the different productivities of the cod and haddock stocks encountered in the Irish Sea. The larvae of both species showed similar responses to changes in environmental conditions (temperature, wind, prey availability, daylight hours) and better survival was predicted in the western Irish Sea, covering the spawning ground for haddock and about half of that for cod. Larval growth was predicted to be mostly prey‐limited, but exploration of stock recruitment data suggests that other factors are important to ensure successful recruitment. We suggest that the presence of a cyclonic gyre in the western Irish Sea, influencing the retention and/or dispersal of larvae from their spawning grounds, and the increasing abundance of clupeids adding predatory pressure on the eggs and larvae; both may play a key role. These two processes deserve more attention if we want to understand the mechanisms behind the recruitment of cod and haddock in the Irish Sea. For the ecosystem‐based management approach, there is a need to achieve a greater understanding of the interactions between species on the scale a fish stock is managed, and to work toward integrated fisheries management in particular when considering the effects of advection from spawning grounds and prey–predator reversal on the recovery of depleted stocks.  相似文献   

2.
Atlantic cod (Gadus morhua) is one of the commercially most important fish species in the North Atlantic and plays a central role in several ecosystems. Fishing pressure has been heavy over a prolonged period and the recent decades have shown dramatic decline in abundance of many stocks. The Arcto‐Norwegian (or North‐east Arctic) cod stock in the Barents Sea is now the largest stock of Atlantic cod. Recruitment to this stock has varied extensively during the last 60 yr. There is evidence for fluctuations in climate, particularly sea temperature, being a main cause for this variability, higher temperatures being favourable for survival throughout the critical early life stages. Our studies of time series present compelling evidence for a strengthening of the climate–cod recruitment link during the last decades. We suggest this is an effect of the age and length composition of the spawning stock having changed distinctly. The age of the average spawner has decreased by more than 3 yr from between 10 and 11 in the late 1940s to 7–8 in the 1990s, average length from just above 90 cm to around 80 cm. The number of age classes contributing to the spawning stock has also decreased, while the number of length groups present increased slightly. Significant decrease in age of spawners has frequently been described for other heavily fished stocks worldwide. We therefore find it likely that the proposed mechanism of increased influence of climate on recruitment through changes in the spawning stock age and size composition is of a general nature and might be found in other systems.  相似文献   

3.
There is considerable interest in understanding the effect of the North Atlantic Oscillation (NAO) on recruitment to North Atlantic cod stocks. An earlier analysis of data for the North‐east Arctic cod stock showed that, while there is an empirical relationship between recruitment and the NAO over the period 1973–96, this relationship was absent over the period 1946–72. It has recently been suggested that the effect of the NAO on recruitment depends on the size of the spawning stock. Here, the possibility that this explains the earlier result for North‐east Arctic cod is tested and rejected.  相似文献   

4.
A comparative analysis of the fish condition (Fulton’s K) of 10 cod stocks in the north Atlantic in relation to the temperature of their habitat, growth rates and their reproductive potential is presented. It is shown that the cod stocks in the north Atlantic display different levels of mean condition, which is partly due to the different temperature regimes of their habitats. Cod living in colder waters, e.g. Greenland, Labrador and Grand Bank stocks, were found to be in poorer condition than cod living in warmer waters, e.g. North Sea and Irish Sea stocks.

Poor condition causes reduced productivity in terms of slow growth and low recruitment potential. Stocks in better condition display significantly higher weights at age 4 than stocks in poor condition. The a coefficients (function’s slopes) obtained from standardised Ricker’s recruitment–spawning stock biomass (SSB) relationships were defined as indicators for the recruitment potential of stocks. These a coefficients were found to be positively correlated with the mean condition factor of the 10 stocks analysed. This indicates that stocks consisting of individuals in poor condition appear to be very susceptible to reduced recruitment at low SSB, while the stocks that consist of fish in good condition seem to behave more robustly with a higher probability of good recruitment at low SSB. The positive effect of the cod condition on their reproductive potential generally implies that the stocks in good condition in the temperate regions of the northeast and west Atlantic can sustain higher exploitation rates than stocks in poor condition in the colder regimes of the northwest Atlantic (Greenland, Labrador and Grand Bank). This is confirmed by the positive relationship established between the estimated biological management reference points Fmed and the mean cod condition factors, as well as by the recent status of these stocks.  相似文献   


5.
《Fisheries Research》2007,87(2-3):262-267
Within commercial fisheries, particularly mixed fisheries, both target and non-target species are often discarded. Discarding represents a potentially significant loss to the productivity of fish stocks; it can have damaging ecological consequences, and is a potential cause of the failure of recovery plans. The Nephrops fishery in the North Sea is classified as a mixed fishery. Nephrops trawls are constructed with smaller meshes than trawls used to target whitefish; consequently, the bycatch of juvenile fish can be substantial. Several new Nephrops trawl designs have been tested in the North Sea. The data from these trials are used to investigate the potential impact of their implementation on cod, haddock and whiting stocks in the North Sea (including the Kattegat and Skagerrak).The model examines five trawl designs, and also the scenarios of a cessation of discarding in all North Sea fisheries and in just the Nephrops fishery. The model is deterministic, and evaluates the relative differences between scenarios assuming all other variables remain constant. If discarding of cod, haddock and whiting in the North Sea fisheries were eliminated, stocks would increase by 41%, 14% and 29%, respectively, within 10 years. Eliminating discarding in the Nephrops fishery alone would increase stocks by 2%, 1% and 13%, respectively, reflecting the relative proportion of catches of these species in the Nephrops fishery. For cod and haddock, the introduction of the Nephrops trawl with a grid with a square-mesh codend was the only scenario in which a notable increase in stock number was observed. This trawl design facilitates the escape of fish of all ages/sizes from the trawl, effectively making the Nephrops fishery a single-species fishery. For whiting, stock numbers and landings increased under all scenarios, but forecasted landings were lower than if current discard patterns continued in all except the no-discards scenario. The dependency of the results on the validity of the assumptions and on the accuracy of the input data is discussed.  相似文献   

6.
Sea temperature has earlier been shown to have a large influence on the recruitment of Arcto-Norwegian cod, Gadus morhua. We here hypothesize that this linkage is partly due to the direct effect of temperature on larval and juvenile growth. Secondly, temperature acts as a proxy for both biotic and abiotic factors influencing recruitment. Indices of abundance of early juvenile cod (2–3 months old), 0-group cod (4–5 months old) and 3-year-old cod are analysed in more detail against the environmental temperature, wind stress components, wind-induced turbulent energy and the spawning stock biomass. To deal with autocorrelation, non-stationar-time and nun-normality, which complicate a statistical time series analysis, randomization and Box-Jenkins methods are applied. In addition to the important effect of high sea temperature during the early life stage in forming strong year classes, the results show that the spawning stock biomass is nearly as important. Also, alongshore southerly wind stress anomalies during the period of pelagic drift (from April through summer) and offshore wind stress anomalies during egg and early larval stages (in April) act favourably on recruitment. The beneficial effect of southerly wind anomalies could he linked partly to high temperature, but the flux of zooplankton-rich water from the Norwegian Sea into the feeding areas of the Barents Sea may also be increased. The favourable influence of offshore winds in April is less predominant and causal links are also less clear; possible explanations for this might be increased offshore spreading of eggs and early larvae, resulting in reduced risk of predation, and increased compensation inflow of intermediate Norwegian Sea water which, in this restricted period of time, has a high concentration of spawning copepods suitable as prey for the developing cod larvae.  相似文献   

7.
Oceanographic and predation processes are important modulators of fish larvae survival and mortality. This study addresses the hypothesis that immature Norwegian spring‐spawning herring (Clupea harengus), when abundant in the Barents Sea, determine the capelin reproduction success through consumption of Barents Sea capelin (Mallotus villosus) larvae. Combining a hydrodynamic model and particle‐tracking individual‐based model, a realistic spatio‐temporal overlap between capelin larvae and predatory immature herring was modelled for the summer seasons of 2001–2003. Capelin larvae originating from western spawning grounds became widely dispersed during the summer season, whereas those originating from eastern spawning grounds experienced a rapid drift into the southeastern Barents Sea. Herring caused a 3% mortality of the capelin larvae population in 2001 and a 16% mortality in 2003, but the effect of predation from herring on capelin larvae was negligible in 2002. Despite a strong capelin larvae cohort and a virtual absence of predatory herring, the recruitment from the capelin 2002 year class was relatively poor from a long‐term perspective. We show that the choice of capelin spawning grounds has a major impact on the subsequent capelin larvae drift patterns, constituting an important modulator of the capelin larvae survival. Variation in drift patterns during the summer season is likely to expose the capelin larvae to a wide range of hazards, including predation from young cod, sandeel and other predators. Such alternative predators might thus have contributed to the poor capelin recruitment during 2001–2003, leading to the collapse of the capelin stock in the subsequent years.  相似文献   

8.
The survival of fish eggs and larvae, and therefore recruitment success, can be critically affected by transport in ocean currents. Combining a model of early‐life stage dispersal with statistical stock–recruitment models, we investigated the role of larval transport for recruitment variability across spatial scales for the population complex of North Sea cod (Gadus morhua). By using a coupled physical–biological model, we estimated the egg and larval transport over a 44‐year period. The oceanographic component of the model, capable of capturing the interannual variability of temperature and ocean current patterns, was coupled to the biological component, an individual‐based model (IBM) that simulated the cod eggs and larvae development and mortality. This study proposes a novel method to account for larval transport and success in stock–recruitment models: weighting the spawning stock biomass by retention rate and, in the case of multiple populations, their connectivity. Our method provides an estimate of the stock biomass contributing to recruitment and the effect of larval transport on recruitment variability. Our results indicate an effect, albeit small, in some populations at the local level. Including transport anomaly as an environmental covariate in traditional stock–recruitment models in turn captures recruitment variability at larger scales. Our study aims to quantify the role of larval transport for recruitment across spatial scales, and disentangle the roles of temperature and larval transport on effective connectivity between populations, thus informing about the potential impacts of climate change on the cod population structure in the North Sea.  相似文献   

9.
Similar to many other commercial marine fish species, Atlantic cod (Gadus morhua L.) migrate towards specific sites at spawning. These temporal aggregations are generally the most targeted by the commercial fishery. The Kattegat cod has undergone a substantial reduction over the past 25 years and both stock size and spawning stock biomass have remained at very low levels since the end of the 1990s. There is, therefore, an urgent need to map and document spawning grounds still in use. In the present study, spawning sites of Atlantic cod were identified in the Kattegat using a combination of commercial and fishery independent data from 1996 to 2004. Moreover, putative spawning and non-spawning areas were also sampled before and during the reproductive season between 2002 and 2006, and the proportion of mature females together with the individual physiological status were used to validate and strengthen the spatial analyses.

The spatial analyses identified several spawning areas in the region and two areas in the southeastern part of the Kattegat which appeared to be the most important. The results showed the presence of cod spawning aggregations, although reduced in size, in areas that have been utilized for more than 25 years according to historical information. Some local spawning grounds may have also disappeared. The proportion of mature females was higher in putative spawning than in non-spawning areas (p < 0.001) and females from spawning areas had higher gonadosomatic (p < 0.05) and hepatosomatic (p < 0.001) indices than those from non-spawning areas.

Knowledge of stock spatial and temporal distribution is essential in designing recovery strategies for depleted fish populations. The unambiguous stability of the locations of spawning aggregations over time, as shown in this study, represents a useful aid in order to efficiently implement a recovery plan for the collapsing cod stock in this area.  相似文献   


10.
Two approaches were used to qualify observed variability in Greenland cod (Gadus morhua) recruitment. In the first analysis, we used the linear trend of the Greenland cod recruitment time series and climatic variables, such as air temperatures from the Denmark Strait and wind conditions off East Greenland and Southwest Greenland, to explain the interannual variation in cod recruitment off Greenland. The model resulting from this ‘trend/environmental approach’, explained 79% of the interannual variation in cod recruitment off Greenland. In the second, analytical approach, the ‘regime approach’, multiple linear regression models were used, with the input data being the time series of cod recruitment and spawning stock biomass (SSB) from Iceland and Greenland, sea surface and air temperatures around Greenland, and zonal wind components between Iceland and Greenland. Model results indicated that, during the decades between 1950 and 1990, there were three different cause–effect regimes which significantly influenced the variability of cod recruitment. The three regimes included: (a) the 1950s and 1960s, a regime with favorable sea surface temperatures and a self‐sustaining cod stock off Greenland with high SSB that produced a series of above‐average, strong year classes; (b) the 1970s and 1980s, a regime of declining SSB and recruitment, with recruitment dependent on advection from Iceland; and (c) the 1990s, when the advective potential for recruitment from the Icelandic cod stock was the only available source for replenishment of the Greenland cod stocks, because cod recruitment in Greenland waters was negligible. The three models explained 76–77% of the observed interannual variation in cod recruitment off Greenland. Both approaches suggested that advective factors were the dominant influences for cod recruitment in the ‘Iceland–Greenland System’.  相似文献   

11.
Depletion of sturgeon stocks world-wide has increased interest in aquaculture-based restoration programmes. The Caspian Sea Sturgeon Ranching Programme (SRP) of the former Soviet Union represents a unique opportunity to evaluate expense, benefits and potential ecological and genetic effects of such restoration programmes. The SRP was initiated in the 1950s to compensate for lost spawning habitat in the Volga River and elsewhere. After its completion in 1962, the Volgograd Dam reduced spawning grounds in the Volga River system, the principal spawning tributary of the Caspian Sea, by ∼80%. For two of the three commercial sturgeon species (Russian sturgeon, Acipenser güldenstädti , and stellate sturgeon, A. stellatus ), yields improved after the imposition of the 1962 moratorium on sturgeon harvests in the Caspian Sea. Volga River fisheries were managed for spawning escapement. Although imprecisely known, the contribution of the millions of stocked Russian and stellate juveniles during 1962–91 was most likely important to sustaining fisheries, although less so (contributing to <30% of the adult stock) than natural recruitment. Apparently, reduced spawning grounds, supplemented with artificial spawning reefs were sufficient to support reproduction and large fishery yields of Russian and stellate sturgeons. For beluga sturgeon, Huso huso , harvests in the Volga River were nearly all dependent upon hatchery stocking. Beluga sturgeon spawning grounds were mostly eliminated with the construction of the Volgograd Dam. Without the hatchery programme, beluga sturgeon in the Volga River and Caspian Sea would in all likelihood have been extirpated. Currently, sturgeons are severely depleted in the Volga River and Caspian Sea due to poaching and lack of co-operation between countries exploiting the species. Aquaculture-based restoration in Russia is now viewed a chief means of rebuilding stocks of Caspian Sea sturgeons.  相似文献   

12.
The introduction of 200 n.m. exclusive economic zones (EEZs) in the late 1970s required increased collaboration among neighbouring coastal states to manage transboundary and straddling fish stocks. The established agreements ranged from bilateral to multilateral, including high‐seas components, as appropriate. However, the 1982 United Nations Convention on the Law of the Sea does not specify how quotas of stocks crossing EEZs should be allocated, nor was it written for topical scenarios, such as climate change with poleward distribution shifts that differ across species. The productive Northeast Atlantic is a hot spot for such shifts, implying that scientific knowledge about zonal distribution is crucial in quota negotiations. This diverges from earlier, although still valid, agreements that were predominately based on political decisions or historical distribution of catches. The bilateral allocations for Barents Sea and North Sea cod remain robust after 40 years, but the management situation for widely distributed stocks, as Northeast Atlantic mackerel and Norwegian spring‐spawning herring, appears challenging, with no recent overall agreements. Contrarily, quotas of Northern hake are, so far, unilaterally set by the EU despite the stock's expansion beyond EU waters into the northern North Sea. Negotiations following the introduction of EEZs were undertaken at the end of the last cooler Atlantic Multidecadal Oscillation (AMO) period, that is, with stock distributions generally in a southerly mode. Hence, today's lack of management consensus for several widely distributed fish stocks typically relates to more northerly distributions attributed to the global anthropogenic signal accelerating the spatial effect of the current warmer AMO.  相似文献   

13.
The reproductive success of marine ectotherms is especially vulnerable in warming oceans due to alterations in adult physiology, as well as embryonic and larval survival prospects. These vital responses may, however, differ considerably across the species' geographical distribution. Here we investigated the life history, focusing on reproductive ecology, of three spatially distant populations (stocks) of Atlantic cod (Gadus morhua, Gadidae) (50–80° N), in the Irish/Celtic Seas-English Channel Complex, North and Barents Seas, under past and projected climate. First, experimental tracking of spawning behaviour evidenced that the ovulation cycle is highly distressed at ≥9.6 (±0.25)°C (Tup). This knife-edge threshold resulted in erratic spawning frequencies, whereas vitellogenin sequestration remained unaffected, indicating endocrine rather than aerobic scope constraints. Cod in the Celtic Sea-English Channel are, therefore, expected to show critical stock depensation over the next decades as spawning grounds warm above Tup, with Irish Sea cod subsequently at risk. Second, in the relatively cooler North Sea, the northward retraction of Calanus finmarchicus (Calanidae) and Para-Pseudocalanus spp. (Clausocalanidae) (1958–2017) limit cod larvae feeding opportunities, particularly in the southernmost subarea. However, the contrasting increase in Calanus helgolandicus (Calanidae) does not counteract this negative effect, likely because cod larvae hatch ahead of its abundance peaks. Overfishing again comes as a twin effect. Third, in the still relatively cold Barents Sea, the sustainably harvested cod benefit from improved food conditions in the recent ice-free polar region but at the energetic cost of lengthier and faster spawning migrations. Consequently, under climate change local stocks are stressed by different mechanistic factors of varying management severity.  相似文献   

14.
Reference points based on fishing mortality (F) and spawning stock biomass (SSB) are a requirement of many fisheries management frameworks. SSB is assumed to be a proxy for stock reproductive potential (SRP). Limit reference points based on SSB are used to indicate the level of biomass below which productivity is affected. SSB fails to account for changes in fecundity, egg viability and sex ratio, and it has been argued that total egg production (TEP) provides a better reflection of SRP. We explore how accounting for TEP impacts limit reference points and evidence for a relationship between stock and recruit. Time series of SSB and TEP are compared for three North Sea stocks: cod (Gadus morhua), herring (Clupea harengus) and plaice (Pleuronectes platessa). Dynamics based on TEP are different from those based on SSB for cod and plaice, but the stock–recruit relationships were not ‘improved’ using TEP. Shifts in productivity (spawner per recruit) occur in all three time series and SSB underestimated uncertainty. Yet again, it was shown that assumptions of stationarity about fish population productivity are incorrect. We argue that the use of TEP does improve the realism in our understanding of stock dynamics, and demographically, more complex management strategy evaluation is required to develop management procedures that are robust to uncertainty and integrate F and the demographic health of a stock. Empirical feedback control systems based on fisheries independent indices including surveys of eggs, larvae, recruits, juveniles or spawning adults should be evaluated and compared to traditional approaches.  相似文献   

15.
To evaluate genetic differences in growth and survival potential as a function of light regime, cod larvae from two stocks (CC: coastal cod and AC: Arcto-Norwegian cod, spawning at latitudes of 60°N and 69°N respectively) were co-reared in mesocosms. The experiment was carried out with two artificially illuminated light regimes, corresponding to the natural photoperiod at peak spawning of the respective stocks (CC: 7–8 h distinct dark period, AC: ‘mid-night sun’ as continuous light with 2–3 h reduced light intensity). One additional regime with ambient natural light analogous to 60°N was used. At termination on day 36 post-hatch, larvae of the co-reared stocks were distinguished from each other by a genetic marker in the CC stock. Stock-specific differences in growth rate and survival were indicated, with CC larvae growing and surviving better than AC larvae. This is in agreement with earlier comparisons of larval growth in these stocks. The 69°N light regime did not enhance growth for any of the two stocks. However, natural light greatly increased growth rate of both stocks, indicat ing important mechanisms in regulation of larval feeding relative to light quality or intensity. The results may be important for choice of stock and illumination in intensive aquaculture of this species.  相似文献   

16.
Stock level of Japanese sardine (Sardinops melanostictus) was high from 1980s to early 1990s and low from late 1990s to 2000s. The warm and cold water masses in the vicinity of the Kuroshio axis from winter to early spring used to be critical for the recruitment in the high‐stock period, because most of the larvae were distributed there. However, the environmental fluctuation might not affect the recruitment in the low‐stock period. Some studies reported that spawning location and spawning season, and hence the larval habitat, differ depending on the stock level. Three points were investigated in this study: (a) how spawning location and spawning season shifted from the late 1990s, (b) confirmation of the distribution area of larvae in the recent low‐stock period and (c) whether the water temperature in the vicinity of the Kuroshio axis was still related to the recruitment in the low‐stock period. The spawning location and spawning season clearly changed after 1995. Consequently, particle tracking experiments suggested that the larvae appeared in the vicinity of the Kuroshio axis from winter to early spring decreased. Nevertheless, only the ambient temperature of larvae that appeared in the vicinity of the Kuroshio axis from winter had a significant negative correlation with an index of the recruitment in the low‐stock period. It is suggested that the warm and cold masses in the vicinity of the Kuroshio axis are critical for the recruitment regardless of the stock level.  相似文献   

17.
Assumptions about the future productivity of a stock are necessary to calculate sustainable catches in fisheries management. Fisheries scientists often assume the number of young fish entering a population (recruitment) is related to the biomass of spawning adults and that recruitment dynamics do not change over time. Thus, managers often use a target biomass based on spawning biomass as the basis for calculating sustainable catches. However, we show recruitment and spawning biomass are not positively related over the observed range of stock sizes for 61% of 224 stocks in the RAM Legacy Stock Assessment Database. Furthermore, 85% of stocks for which spawning biomass may not drive recruitment dynamics over the observed ranges exhibit shifts in average recruitment, which is often used in proxies for target biomasses. Our results suggest that the environment more strongly influences recruitment than spawning biomass over the observed stock sizes for many stocks. Management often endeavours to maintain stock sizes within the observed ranges, so methods for setting management targets that include changes within an ecosystem may better define the status of some stocks, particularly as climate changes.  相似文献   

18.
An individual-based modelling approach was developed to investigate the spatial and temporal patterns in the recruitment processes of North Sea haddock, Melanogrammus aeglefinus . The approach was based on the realization that the survivors to recruitment of an annual cohort are most probably not drawn at random from the initial population of eggs, but represent the fastest-growing individuals. Individual growth rates reflect the unique exposure of each larva to the environment along its drift trajectory. In this context, the environment refers to a wide range of factors affecting growth such as food, turbulence and temperature. A combination of a model of egg production by the adult stock, a particle-tracking scheme, and a model of larval growth and mortality rate was used to simulate the dispersal trajectories, and the survival of haddock larvae spawned at different times and locations on the continental shelf. The particle tracking was driven by flowfields from a climatological implementation of the Hamburg Shelf–Ocean Model (HAMSOM) for the North Sea and NE Atlantic. The system was able to resolve spatial and temporal patterns in the recruitment process and indicated that the surviving population of larvae was drawn from a restricted part of the spawning distribution. The results have the potential to guide the development of future conservation measures in fisheries management.  相似文献   

19.
Spawning of fishes takes place across a wide area of the North Sea. However, more intense spawning is seen in restricted areas, indicating that such areas present favorable conditions. To update information on fish spawning in the North Sea and analyze potential linkages to hydrographic characteristics, an internationally coordinated survey was conducted in the winter/spring of 2004. Oblique hauls for fish eggs and larvae and vertical profiles of temperature and salinity were carried out at 393 stations across the entire North Sea. The hydrography was strongly influenced by the interfacing of water masses of different salinity, and frontal zones were seen along all coastal areas and off the Dogger and Fisher Banks. Total abundances of eggs and larvae, including fish species such as cod, haddock, plaice, long rough dab and sandeel, peaked in the vicinity of the frontal areas. Hence our findings indicate that the main spawning locations of fish are linked to recurrent hydrographic features such as salinity fronts. Such a linkage may provide survival advantages, as the fronts present favorable feeding conditions, and the related physical processes may confine egg and larval dispersal and transport them towards suitable nursery habitats.  相似文献   

20.
A substantial improvement in the bycatch selectivity of Norway lobster Nephrops norvegicus trawls is required, particularly with respect to cod Gadus morhua, whose stocks are at low levels in several areas. Conventional escape windows are not adequate to properly release cod and other bycatch species caught in the trawls. To address this issue, we developed a novel sorting box concept consisting of a four-panel section with a window on the top in order to improve the escape of cod and other bycatch species through an escape window while retaining the target catch of Norway lobster. The concept was tested on a commercial trawler in Kattegat and Skagerrak. Two different window mesh sizes and two different sorting box heights were tested using a traditional codend cover and a dual codend cover. We observed greatly reduced bycatches of both cod and other fish species compared to a standard codend. The reduction in bycatch decreased with decreasing mesh size and increasing height of the sorting box. Escape of Norway lobster through the escape window was limited. A modified version of the sorting box concept was implemented in the Kattegat fishery from 2009 onwards.  相似文献   

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