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1.
We quantified biomass accumulation and nitrogen (N) retranslocation, allocation, and utilization of Changbai larch (Larix olgensis) seedlings subjected to four fertilization treatments (24, 59, 81, 117 kg·ha-1 N) with an unfertilized control during summer and autumn 2009. Ammonium phosphate (18-46-0) was the fertilizer used in all treatments. On both sampling dates, the needles had greater biomass and N content than new (2009) stems and old (2008) stems, and coarse, medium and fine roots (diameters of >5, 2-5 mm, and 0-2 mm, respectively). Higher N concentration was observed in old stems and coarse roots than that in new stems and medium roots. In mid-summer, fine roots had higher N concentration than coarse roots. The treatment with 24 kg·ha-1 N had the greatest biomass and N content in needles and old stems, and highest net N retranslocation (NRA) and amount of N derived from soil. On September 21, no N translocation was observed, while the treatment with 24 kg·ha-1 N had the highest N utilization efficiency and fertilizer efficiency. Vector analysis revealed that all four fertilization treatments induced N excess relative to the control. The treatments with 59, 81, 117 kg·ha-1 N induce N excess compared with treatments at 24 kg·ha-1 N. We conclude that the traditional local fertilizer application rates exceeded N requirements and N uptake ability for Changbai larch seedlings. The application rate of 24 kg·ha-1 N is recommended.  相似文献   

2.
We measured respiration of 20-year-old Pinus radiata D. Don trees growing in control (C), irrigated (I), and irrigated + fertilized (IL) stands in the Biology of Forest Growth experimental plantation near Canberra, Australia. Respiration was measured on fully expanded foliage, live branches, boles, and fine and coarse roots to determine the relationship between CO(2) efflux, tissue temperature, and biomass or nitrogen (N) content of individual tissues. Efflux of CO(2) from foliage (dark respiration at night) and fine roots was linearly related to biomass and N content, but N was a better predictor of CO(2) efflux than biomass. Respiration (assumed to be maintenance) per unit N at 15 degrees C and a CO(2) concentration of 400 micro mol mol(-1) was 1.71 micro mol s(-1) mol(-1) N for foliage and 11.2 micro mol s(-1) mol(-1) N for fine roots. Efflux of CO(2) from stems, coarse roots and branches was linearly related to sapwood volume (stems) or total volume (branches + coarse roots) and growth, with rates for maintenance respiration at 15 degrees C ranging from 18 to 104 micro mol m(-3) s(-1). Among woody components, branches in the upper canopy and small diameter coarse roots had the highest respiration rates. Stem maintenance respiration per unit sapwood volume did not differ among treatments. Annual C flux was estimated by summing (1) dry matter production and respiration of aboveground components, (2) annual soil CO(2) efflux minus aboveground litterfall, and (3) the annual increment in coarse root biomass. Annual C flux was 24.4, 25.3 and 34.4 Mg ha(-1) year(-1) for the C, I and IL treatments, respectively. Total belowground C allocation, estimated as the sum of (2) and (3) above, was equal to the sum of root respiration and estimated root production in the IL treatment, whereas in the nutrient-limited C and I treatments, total belowground C allocation was greater than the sum of root respiration and estimated root production, suggesting higher fine root turnover or increased allocation to mycorrhizae and root exudation. Carbon use efficiency, the ratio of net primary production to assimilation, was similar among treatments for aboveground tissues (0.43-0.50). Therefore, the proportion of assimilation used for construction and maintenance respiration on an annual basis was also similar among treatments.  相似文献   

3.
Abstract

Knowledge of the canopy lichen flora of managed forests is poor, but needs more focus since, for example, slash (tops, branches and twigs) harvest for biofuel may pose a threat to epiphytic lichen diversity. This study compared lichen species richness, density and composition between stems, tops, branches and twigs of mature Norway spruce (Picea abies) and aspen (Populus tremula) in managed boreonemoral forests in south–central Sweden. The stems were also compared with the slash fractions pooled together. All comparisons were made separately for each tree species. In total, 30 lichen species were found on Norway spruce and 46 on aspen. No significant differences in species richness or species density between fractions were found for Norway spruce, whereas aspen tops were significantly less species rich and species dense than the other fractions. Moreover, aspen slash was significantly more species dense than the stem. The lichen species composition of the stems clearly differed from that of the tops, branches and twigs in both tree species. Thus, lichen communities other than those removed with stems by conventional forestry are removed from the stands owing to slash harvest. However, these species are rather common and widespread in Sweden. The impact of slash harvest on the epiphytic lichen flora may therefore be of minor importance in forests established after clear-cutting or on former arable land.  相似文献   

4.
Lack of information concerning root growth of trees limits our knowledge of plant development and fertilizer response. The objective of this work was to study root growth dynamics of an E. urophylla forest after harvesting and the supply of nutrients from the roots and the soil to the new sprouts originating from the stumps. About 7-year-old eucalypt trees were felled and the sprouts and roots were sampled at 0, 60, 120, 180, 240, 330 days after harvesting. The roots were separated into fine roots (<1 mm), medium roots (1–3 mm), coarse roots (>3 mm), and taproot. Nutrient supply to sprouts from the old roots and the soil was calculated based on the change in nutrient content of the roots with time and accumulation of nutrients in the sprouts. Fine, medium and coarse root biomass increased with time after harvesting. However, the increase was more pronounced with fine roots. Between harvesting and day 60 of the new growth, all nutrients allocated to the sprouts, excluding potassium, were supplied by the soil. K was the nutrient most dependent on root reserves for the initial growth of sprouts. The contribution of the old roots to N, P, Ca, and Mg accumulation in the sprouts increased between day 60 and 120. At 330 days after harvesting, about 9.2, 23.9, and 12.6% of the N, K, and Mg, respectively, that had accumulated in the sprouts were supplied by the roots, while all P and Ca were supplied by the soil.  相似文献   

5.
The decomposition of Pinus radiata (D. Don) stems, coarse woody roots and stumps was studied in Tarawera forest, Bay of Plenty region, North Island, New Zealand. The study examined the residues from two thinning events with 6 and 11 years of decay. Changes in the mass of stems, and density of roots and stumps were used to estimate the decay rate constants using a single exponential model. The decay rate of stems was not significantly related to DBH and averaged 0.1374 year−1 (22 years for 95% mass loss). The decay rate of coarse woody roots was not significantly different to stem decay and averaged 0.1571 year−1 (19 years for 95% mass loss). A large range in stump decay rates was measured and a significantly lower decay constant was observed for stumps (0.1101 year−1, 27 years for 95% mass loss), possibly due to the stumps being kept alive after felling through root grafting and a resistance to decay due to the presence of resin. The concentration of C remaining in stems and stumps increased with mass loss from 52% to 55% C after 11 years of decay. The C concentration in coarse woody roots initially increased but then declined near to the original level of 50% after 11 years of decay. Nitrogen concentrations increased substantially in all components with decay.  相似文献   

6.
Bareroot Changbai larch (Larix olgensis Henry.) seedlings were reared with inorganic fertilizer (nitrogen (N):phosphorus (P) = 1:1, W/W) applied at a rate of 100 (F100) or 200 kg N ha−1 (F200) with (+) or without (−) chicken manure as a soil amendment (O) in north-eastern China. An unfertilized control treatment was included. Inorganic and organic fertilizer treatments tended to increase soil ammonium, nitrate, available P, total P, organic carbon content and electrical conductivity, and biomass and N concentration in seedlings. Organic amendment improved first order lateral root number, tap root length, fine root morphology (length, surface area, volume) in seedlings, while the F100 treatment increased N accumulation in needles and stems compared to the F200 treatment, on average. Most fertilizer treatments tended to increase P content in combined stems and roots, but F200 − O and F100 + O treatments diluted P in needles. Organic amendment combined with inorganic fertilizer at a rate of 100 kg N and P ha−1 is recommended to improve seedling growth and N reserves in woody tissues.  相似文献   

7.
The attacks of bark‐ and wood‐boring Coleoptera on broken conifer stems after severe snow‐breakage in early 1988 were studied in autumn 1988 and 1989. The study included twelve stands in the county of Värmland in central Sweden differing in age and edaphic conditions. The tops on the ground as well as the remaining rooted stem stumps of 94 Picea abies and 61 Pinus sylvestris were inspected. The presence of living branches on the stumps strongly influenced the incidence of insect attack. All stumps without branches were attacked during the two‐year period, whereas only a few spruce stumps with more than ten branches and no pine stumps with more than five branches were attacked. On spruce stumps, the most frequently encountered Coleoptera were all scolytids, i.e. the species Pityogenes chalcographus, Hylurgops palliatus, and the genera Polygraphus, Dryocoetes, and Trypodendron. On pine stumps, Tomicus piniperda was the most common species. Most of the spruce tops and virtually all pine tops were attacked during the two‐year period, and most of these attacks occurred during the first summer. P. chalcographus, Dryocoetes, and weevils of the genus Pissodes were the most frequent Coleoptera on spruce tops. On pine tops, the dominant insects belonged to the genera Pissodes and Pityogenes.  相似文献   

8.
研究表明，在松材线虫发生区，疫病的扩散和流行与松树的采伐剩余物有着密切的关系。松树的新鲜采伐剩余物，如梢，枝，干，桩，每平方分米表面积松褐天牛幼虫蛀入孔数分别为1．46个，1．41个，0．18个，0．29个。次年，从这些剩余物上每平方分米分别能出现0．55个，0．84个，0．15个，0．27个松褐天牛羽化孔，经镜检，受松褐天牛寄生的采伐剩余物均带有松材线虫。由此可见，松树采伐剩余物是松材线虫病扩散流行的一个重要的中间环节，改善松林卫生状况，规范松林采伐技术，科学处理剩余物是抑制松材线虫病扩散和流行的重要环节和措施。  相似文献   

9.
Effects of harvest residues on nutrient leaching and soil chemical properties were studied in a lysimeter experiment. Treatments were: (A) forest floor litter and harvest residues, other than branches, incorporated into the soil, (B) as A, but with branches cut in 20 cm long bits and placed on the soil surface, (C) as B, but with bits incorporated into the soil, (D) as B, but with branches chopped into chips, (E) as C, but with branches chopped into chips, (F) forest floor litter and harvest residues on the soil surface, with branches cut in 20 cm long bits, (G) as F, but with branches chopped into chips, and (H) absence of harvest residues (control). Treatments were applied in zero-tension lysimeters containing 25 kg of soil. Leachates were collected for a 6-year period. At the end of the experiment, lysimeters were dismantled and soil was divided in four layers. Residues, other than branches, increased N leaching, as compared with the control. Branches on the soil surface reduced N leaching when cut in chips. Branches incorporated into the soil reduced leaching independently of their size. Organic residues on the soil surface showed similar effect to those incorporated into the soil. However, harvest residues on the soil surface increased leachate volume, and reduced Ca and P losses. Such a placement of residues led to high contents of Ca and P in the 0–5 cm top soil layer. Contents of organic C, total N and base cations were not affected by the treatments.  相似文献   

10.
The soil acidity of four coniferous sites, two in S Sweden and two in N Sweden, were studied 7–9 years after clear‐felling. Each site had three different slash treatments; (a) all slash harvested, (b) only needles remaining, and (c) all slash remaining. Our results verify that harvesting of slash after clear‐cutting means an acidification of the organic horizon in acid forest soils. On plots subjected to full slash harvest the pH‐value of the forest floor was 0.2–0.4 units lower than that of slash‐covered ones, and at three of the four study sites the exchangeable acidity was significantly higher on the slash‐cleared plots. Plots with only needles remaining were generally intermediate in acidity. The change in exchangeable acidity corresponded to approximately 10% of the theoretically estimated proton load caused by slash harvesting. The rate of proton flux associated with whole‐tree harvesting varied between 7–65 meq m‐3 yr‐1, which roughly corresponds to the amounts deposited in precipitation. Harvesting of stems accounted for about one third and needles for one third of the proton load.  相似文献   

11.
Abstract

Biomass, total nitrogen (N) and total carbon (C) stocks were determined in trees, roots, field vegetation and soil in plots given two different site preparation treatments, deep soil cultivation (DSC) approximately 50?cm deep and patch scarification (PS), at three locations in Sweden 10?years after treatment. One location was planted with Pinus contorta, one with Picea abies and one with a mixture of P. abies and Pinus sylvestris. No differences were found in total ecosystem (trees, roots, field vegetation and soil) C and N stocks between the DSC and PS plots. In the DSC plots the tree biomass, tree N and C contents and total biomass were higher than in the PS plots, but the opposite was found for stocks in field vegetation. Biomass and C stocks in the total vegetation (trees, roots and field vegetation) were higher in the DSC plots. However, vegetation N stocks did not differ between the soil treatments, probably because the combined amount of leaf tissue in the trees and field vegetation did not differ between them. The proportions of biomass allocated to roots, stems and needles did not differ between the two treatments. However, the rooting was deeper in DSC plots, possibly because nutrient availability was higher, and subsoil density lower, following DSC than following PS.  相似文献   

12.
We estimated above- and belowground biomass and net primary production (NPP) of a 73-year-old Scots pine (Pinus sylvestris L.) forest stand in the Belgian Campine region. Total biomass for the stand was 176 Mg ha(-1), of which 74.4% was found in stems. The root system contained 12.6% of total biomass, most of it in coarse roots (> 5 mm). Fine roots (< 5 mm) comprised only about 1.7% of total biomass, and more than 50% of fine root biomass was retrieved in the litter layer and the upper 15 cm of the mineral soil. The ratio of belowground biomass to aboveground biomass was 0.14, which is lower than that of other Scots pine forests and other coniferous forests. Between 1995 and 2001, mean annual NPP was 11.2 Mg ha(-1) year(-1), of which 68.7% was allocated to aboveground compartments. Stems, needles and cones made relatively high contributions to total NPP compared with branches. However, branch NPP was possibly underestimated because litterfall of big branches was neglected. The proportion of total NPP in belowground components was 31.3%. Coarse root NPP (2% of total) was low compared with its biomass. Fine root NPP was 3.3 Mg ha(-1) year(-1), representing about 29.5% of total NPP; however, the estimate of fine root NPP is much more uncertain than NPP of aboveground compartments. The ratio NPP/GPP (gross primary production) was 0.32, which was low compared with other coniferous forests.  相似文献   

13.
We studied the biomass and its allocation in natural secondary forests, as well as the amounts, accumulation and distribution of nutrient elements (N, P, K, Ca and Mg) in sample plots established in the Loess Plateau in Shanxi Province, northern China. The results show that biomass in natural secondary forests amounted to 36.09 t/hm2, of which the tree layer accounted for 46%, the shrub layer for 29%, the herb layer for 13% and the litter layer for 12%. The total storage of the five nutrient elements is 1089.82 kg/hm2. Nutrient storage in the tree layer is the largest, at 41%. The sequence of storage of the elements varied among different layers and is given as follows: shrub layer 31.27%, herb layer 12.55% and litter layer 15.36%. The accumulation of nutrient elements in the tree layer, ordered from high to low, is: branches > roots > stems >bark > leaves. The total storage of the five nutrient elements in the soil is 634.97 t/hm2, where the accumulation of the nutrients accounts for 95.32% (N), 99.64% (P), 99.91% (K), 99.84% (Ca) and 99.95% (Mg) of the total amounts. The accumulation coefficients of different organs in the tree layer are, from high to low: leaves > branches > roots > bark > stems. The accumulation coefficients in the different layers are listed as follows: shrub layer > tree layer > herb layer and for the elements as: N > P > Ca > K > Mg. __________ Translated from Journal of Beijing Forestry University, 2008, 30(3): 57–62 [译自: 北京林业大学学报]  相似文献   

14.
三峡库区马尾松根和叶片的生态化学计量特征   总被引：3，自引：0，他引：3       下载免费PDF全文
[目的]对马尾松根系和叶片养分(C、N、P、K、Ca、Mg)进行研究,探讨生长季和非生长季根系和叶片的化学计量学特征和养分分配特征,以及根和叶片养分的相关性。[方法]本研究分别在2014年7月和11月,利用土柱法采集根样和高枝剪采集枝条第二节上的健康叶样,然后再进行室内测试分析。[结果]表明:(1)非生长季与生长季相比,根C养分含量、C:N和N:P比值均下降,N、P、K、Ca、Mg养分含量含量均增加。(2)根C养分含量随直径增大而增加,N、P、K、Ca、Mg养分含量随直径增大而减小。(3)径级、取样时间以及两者交互作用对根系C、N、P、K、Ca、Mg养分含量和C、N、P养分化学计量比存在极显著影响(P0.01),同时取样时间对粗根C、N养分含量以及C:N比无显著影响。(4)细根(2 mm)C和N元素含量存在极显著的负相关关系,N和P元素含量存在极显著的正相关关系,同时C:N比值和N:P比值变异分别由各自两元素含量共同决定;粗根(2 3 mm)C、N计量比值和N:P比值变异分别主要由N元素含量和P元素含量决定;(5)从地上叶到地下根,C、N、P、K养分含量呈减少趋势,Ca、Mg养分含量呈增加趋势。同时,根和叶片养分含量相关性较弱,除C、K养分之外。[结论]马尾松根系养分含量与叶片养分的关系较弱,且分别对各养分的相对需求量不同;与生长季相比,非生长季马尾松根系N、P、K、Ca、Mg含量显著增加;C、N元素和N、P元素的耦合关系只出现在细根中。  相似文献   

15.
Coarse woody debris (CWD) has become recognised as an important component of the carbon (C) pool in forest ecosystems. In Ireland, managed Sitka spruce (Picea sitchensis (Bong) Carr.) forests account for 52.3% of the total forest estate. To determine the stock and decay dynamics of above and belowground CWD, field surveys using fixed area sample plots, were conducted in six even-aged Sitka spruce stands, representing the young, intermediate and mature stages of a typical commercial rotation. The volume, mass, density loss and C:N ratio of all CWD types (logs, stumps, and coarse roots) were determined using a five-decay class (DC) system. The decay rates and half life of CWD was also determined. To estimate CWD coarse root mass; roots associated with stumps classified in different decay classes were excavated. The coarse roots were categorised into small (2-10 mm), medium (10-50 mm) and large (>50 mm) diameter classes.CWD C-mass ranged from 6.98 to 18.62 Mg ha−1 and was highest in an intermediate forest (D35), while the aboveground volume varied from 6.31 to 42.27 m3 ha−1. Coarse roots accounted for 21% to 85% of the total CWD C-pool in the surveyed stands. The total CWD C-mass was poorly correlated with the number of thinning events (R2 = 0.29), when data from D35 was excluded. The density loss was significant in logs (45%), stumps (58%), and small- (38%), medium- (50%) and large roots (38%) as decay progress from DC 0 to 4. There was a 46%, 41%, 51%, 72% and 57% decline in C:N ratio of logs, stumps, small-, medium- and large roots, respectively, as decay progressed from DC 0 to 4. The density decay rates were 0.059, 0.048 and 0.036 kg m−3 year−1 for logs, stumps and coarse roots, respectively. The size classification of roots did not significantly affect their decay rate. The half life (50% decomposition) of CWD was estimated has 12-, 14- and 19 years for logs, stumps and roots of Sitka spruce. Regression curves showed a strong correlation between the density and C:N ratio (R2 = 0.69, 0.74 and 0.93 for logs, stumps and coarse roots, respectively). The long term storage of C and its slow rate of decomposition make CWD a vital structural and functional component of the CWD C-pool and a major controller of forest ecosystem C-retention.  相似文献   

16.
Models for cycles for organic matter and nutrients element (N, P, K, Ca and Mg) are presented for the agroforestry systems of cacao (Theobroma cacao) withCordia alliodora orErythrina poeppigiana in Turrialba, Costa Rica. For the models, system reserves (soil, humus, vegetation divided into leaves, branches, stems, fine roots, fruits) and transference between compartments (production and decomposition of litter residues) inputs (fertilizer, rainfall) and outputs (harvests) of the system are considered. The implications of the models are discussed in detail. Aspects of net primary production in the systems studied are considered. N fixation is calculated on the basis of balances. Analysis of soil water showed high variations that coincided with rainfall patterns and pruning of theE. poeppigiana. For part I see Vol. 4, No. 3, 1986 For part II see this issue Agroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesellschaft für Technische Zusammenarbeit), Turrialba, Costa Rica  相似文献   

17.
Models for cycles for organic matter and nutrients element (N, P, K, Ca and Mg) are presented for the agroforestry systems of cacao (Theobroma cacao) withCordia alliodora orErythrina poeppigiana in Turrialba, Costa Rica.For the models, system reserves (soil, humus, vegetation divided into leaves, branches, stems, fine roots, fruits) and transference between compartments (production and decomposition of litter residues) inputs (fertilizer, rainfall) and outputs (harvests) of the system are considered.The implications of the models are discussed in detail.Aspects of net primary production in the systems studied are considered.N fixation is calculated on the basis of balances. Analysis of soil water showed high variations that coincided with rainfall patterns and pruning of theE. poeppigiana.For part I see Vol. 4, No. 3, 1986 For part II see this issueAgroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesellschaft für Technische Zusammenarbeit), Turrialba, Costa Rica  相似文献   

18.
Fall nutrient loading of deciduous forest nursery seedlings is of special interest because of foliage abscission and varied translocation patterns. For non-deciduous seedlings in the nursery, fall fertilization typically can reverse nutrient dilution and possibly increase nutrient reserves; however, this technique has received little attention with deciduous conifer trees that translocate nutrients before abscising foliage. We evaluated how fall nitrogen (N) fertilization affected N storage and translocation in the deciduous conifer Olga Bay larch (Larix olgensis Henry) seedlings during the hardening period. Seedlings were supplied with 25 mg N seedling?1 for 15 weeks before hardening and fall fertilization treatments began with a three week application period of K15NO3 at 0, 5, 10 and 15 mg N seedling?1. During the hardening period, fall N fertilization had little effect on seedling morphology. The N concentration and content of needles decreased dramatically as needles abscised, while that of stems and roots increased. Six weeks after fall N fertilization ceased, all seedlings translocated similar net N from their needles. For the control seedlings, this accounted for 84 % of the N stored in stems and roots. For fall fertilized seedlings, however, the proportion of N stored in stems and roots translocated from needles accounted for only 41–61 % of the total because of absorption of fall fertilizer that was translocated directly to stems and roots. Six weeks after fall fertilization, the distribution pattern of N concentration and content in seedlings was found in this order: stems > fine roots > coarse roots > needles. Our results suggest that providing deciduous conifer seedlings N during hardening, in this case Olga Bay larch, is a way to promote nutrient loading during nursery production.  相似文献   

19.
《林业研究》2021,32(5)
Previous research has shown that competition between plants can have differential effects on leaf stoichiometry and non-structural carbohydrate(NSC) in different environments.However,little attention has been given to understanding these effects on non-photosynthetic organs,particularly of deciduous tree species.Here we assess the impact of competition on below and aboveground biomass,stoichiometry,nutrient composition and NSC in pure and mixed forests of two Larch species,Larix kaempferi and L.olgensis under nitrogen(N) addition.Nitrogen enrichment did not result in stronger intraspecific competition for both species and L.olgensis benefited from the presence of L.kaempferi under different N levels.Stems kept relatively stable C/N compared to roots and branches in response to competition,while N addition imposed stronger impacts on N/P of different organs rather than competition.In contrast to stable C concentrations,starch and soluble sugar concentrations were more easily impacted by competition and the addition of nitrogen.Competition forced L.kaempferi and L.olgensis to allocate more carbon into storage by increasing their starch concentration and starch/soluble sugar of stems under competition.However,no significant differences in stoichiometry and NSC concentration between intra-and interspecific competition were found.NSC and nutrient pools of L.kaempferi stems,branches and coarse roots consistently declined due to competition regardless of N addition.Coarse and fine roots of L.kaempferi accumulated more N when in competition with L.olgensis than with a conspecific neighbor under N addition.Our results show that NSC was more sensitive to competition relative to stoichiometric traits(N and P) of non-photosynthetic organs.  相似文献   

20.
The response of N-fertilisation, irrigation, acid irrigation, and liming on concentrations of elements in needles, fluxes of elements in litter, volume increment, and crown density on a full stocked, healthy, and vigorously growing mature spruce stand was investigated. The plots of the Höglwald site in Southern Bavaria exhibited a high volume increment with regularly more than 20 m3 ha−1 per year, despite the high age of the stand (77 years at the beginning of the experiment in 1984). Neither a distinct growth reduction of the stand due to acid irrigation, or N oversaturation, nor an enhancement of growth due to N-fertilisation, irrigation, or liming of the stand could be detected. For the years 1984 and 1985 a marked decrease in crown density was detected for all plots. This was followed by a stagnation for 2 years. Afterwards the crown density improved until the end of the investigation for all plots. Neither acid irrigation, nor liming altered the amount of litter fall. Ca fluxes in litter, and concentrations in needles were enhanced on most of the limed plots five to six years after liming. Acid irrigation reduced Ca content in litter, but other elements were not or only slightly influenced. The nutritional status of all plots as shown by the concentrations of elements in needles indicates that for most of the years a sufficient to high supply of N, P, Ca, and Mg for all investigated plots, whereas K concentration in needles exhibited wide year to year variations. Most of the elemental concentrations in needles and fluxes in litter were not influenced by any of the treatments. Also, N-fertilisation did not enhance the N concentrations in needles significantly.  相似文献   

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