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1.
Abstract

The re-establishment rates of reindeer lichen following conventional soil preparation by disc trenching and gentle soil preparation using HuMinMix apparatus were calculated from data acquired in surveys of 17 Pinus–lichen clear-cut stands in northern Sweden conducted 1–15 years after the treatments. The maximum lichen cover and biomass recorded in areas scarified by disc trenching were 29% and 24% of the lichen cover and biomass measured in adjacent intact lichen mats in the surveyed stands after 15 years. By contrast, 9 years after scarification, the corresponding percentages measured in areas disturbed by the HuMinMix treatment were 91% and 100%, respectively. According to calculated linear regressions, the reindeer lichen cover could be completely re-established just one decade after HuMinMix treatment. Ground lichen diversity in the scarified areas was also inventoried. The significant increase in lichen diversity for HuMinMix, especially of early-successional species, could explain the more rapid re-establishment of late-successional lichens (e.g. Cladonia stellaris spp.), compared with disc trenching. Thus, the study demonstrates that both the degree of initial disturbance and the re-establishment rate of reindeer lichen must be considered when estimating the effects of scarification on lichen loss for reindeer grazing over time.  相似文献   

2.
In northern Sweden, the forests are used simultaneously for both timber production and reindeer husbandry. During the winter months, lichen is the most important fodder for reindeer. Forest management operations are generally considered having a negative impact on reindeer husbandry as harvesting and dense stands remove or obscure the ground lichen cover. In this study, we simulate three different scenarios for forest management, differing in the intensity and types of harvest operations. The resulting 100-year scenarios are analyzed with respect to their estimated suitability for providing reindeer pasture areas. Suitability is determined by vegetation type, stand density and stand height. The results indicate that the current trend of a decrease in lichen area will continue if existing forestry practice prevails. Implementing continuous cover forestry as a management alternative and carrying out precommercial thinning could halt the decrease in reindeer pasture area and even lead to a future increase in pasture area, with losses of approximately 5% in the net present value of forestry.  相似文献   

3.
Abstract

Stump removal procedure increases the extent of exposed mineral soil in the clear felled areas. In this study, our aim was to find out whether the early growth and mycorrhization of Norway spruce (Picea abies (L.) Karst.) seedlings differ between stump removal and mounding sites. Stumps were harvested from five one-hectare study sites and other five sites were mounded after clear felling. Twenty seedlings were planted on mounds at each study site. Although the height of spruce seedlings did not differ between the treatments after three growing periods, their mean growth was ca. 10% higher at the stump removal sites. The community of ectomycorrhizal (ECM) in the seedling roots did not differ between the treatments. The high mycorrhization rate in the nursery still remained after two and three growing seasons in the field and diversity of ECM community slightly increased during the first years. Our results indicate some positive although rather small and transient effects of stump removal on the performance of planted spruce seedlings. Longer-term studies are needed to fully understand the effects of stump removal on the later seedling development and their symbiotic interactions with mycorrhizal fungi.  相似文献   

4.
Conserving forest biodiversity has become a significant global concern. A change in forest composition, compared to pre-industrial values, is an important biodiversity issue. In this regard, there is a significant decrease in Picea glauca densities at the landscape level in boreal mixedwoods of northeastern Canada, compared to pre-industrial abundance. Enrichment planting is considered a promising silvicultural approach for improving P. glauca stocking and re-establishing its natural co-dominance with Abies balsamea. However, optimal enrichment scenarios must be identified, notably regarding site preparation treatments that can enhance planted seedling survival and growth while protecting Abies advance regeneration. We established a field trial in Quebec (Canada), laid out as a complete block split-plot design. Picea glauca and P. mariana seedlings (main plot) were planted according to 3 “microsite” treatments (subplot): (1) planting in the humus layer; (2) planting in the mineral soil, with minimal humus disturbance; and (3) planting in a microsite mechanically-prepared using modified brushsaws. After 3 growing seasons, P. mariana seedlings were taller than P. glauca seedlings, but had smaller ground-level diameters. Seedlings planted in the mineral soil had overcome the initial height difference they had with seedlings planted in the humus and mechanically-prepared microsites, with no interaction with species. Ground-level diameter was similar for seedlings planted in the humus and mechanically-prepared microsites, but smaller for seedlings planted in the mineral soil. Water potentials, available light, and foliar nutrient concentrations were not influenced by treatments or species; survival was >90 %. Our results indicate that P. glauca can be established through enrichment planting in boreal mixedwoods. Localized site preparation is not required to limit initial plantation stress in this ecosystem.  相似文献   

5.
Birch forests cover large areas of the northern Fennoscandia and the mountain birch (Betula pubescens ssp. czerepanovii) often forming the altitudinal and Arctic forest and tree lines. Interpreting the factors leading to landscape changes in mountain birch forest involves disturbance from anthropogenic use and climate as important factors. Using vegetation maps based on aerial photographs and satellite images from 7 years in combination with statistical data and ground estimation data of biomass in the period 1957–2006, we were able to assess the transitions among mountain heaths and different types of forest, the displacement of the altitudinal forest line and hence the change in biomass. The tree biomass as well as the growing stock for birch in Finnmarksvidda doubled in the period 1957–2006. Only lichen biomass has been significantly reduced in the period 1957 to 2000, with a subsequent slight increase of lichen biomass in the period 2000 to 2006. The results presented in this paper show significant and positive relationships between measured tree biomass and the increase in reindeer population in the period 1957–2006 in the study area. The hypothesis concerning the removal of the “lichen barrier effect” by heavy reindeer grazing which leads to increased success for birch seeds to germinate and sprout is therefore considered to be valid. However, also, climate change effects like increased precipitation, moth attacks, freezing and thawing events during winter and long-transported air pollution (e.g. nitrogen) may also have reinforced the changes in biomass. Climatic variables from regional weather stations for the period 1955–2006 did not, however, reveal any consistent change except for that the increased snow depth had a negative impact on the reindeer population while increase in snow depth hindering the reindeer in grazing seemed to have a positive impact on the lichen biomass. The ongoing development of increased forest cover and hence elevated forest line will lead to reduction of open habitats, and hence decreased grazing accessibility for the reindeer. The effects on forestry and carbon sequestration are also discussed, and here one of the suggestions is to let the northern birch forests act as carbon sinks contributing to the reduction of total net emission of CO2 in the Nordic countries. Land use like reindeer husbandry has shown to strongly affect relationships between ecological processes like tree-growth and climate. Moreover, reindeer husbandry is such a widespread human activity in the arctic and boreal region that it might affect the global carbon budget.  相似文献   

6.
The purpose of this study was to examine the effects of type and amount of ground vegetation and substrate on the germination and early survival of Norway spruce (Picea abies (L.) Karst) seedlings in uneven-aged stands on mineral soils in Southern Finland. The study was conducted on two permanent plots representing average site quality for spruce stands in the area. A gradient in selection cut intensity and residual stand density was imposed across each of the two permanent plots. Regeneration and vegetation surveys were repeated annually on a grid of unmanipulated microplots and on a set of soil treatment microplots. The results indicated that the number of spruce germinants per unit area was positively correlated with herb cover but negatively correlated with total plant cover. Germinant density was also related to vegetation-free seedbed conditions, high overstory competition index and presence of larger spruce seedlings. Regeneration microsites dominated by moss or with plants with a relatively high light demand were associated with low germinant numbers. Bare humus layer and decayed stumps seemed to promote germinant emergence. Exposure of mineral soil resulted in germination rates that were 4.3–6.9 times higher than in undisturbed conditions during the first growing season.  相似文献   

7.
Temperatures of sandy podzols of middle taiga pine forests with moss and lichen ground cover were analyzed which had been exposed to ground fires of low to medium intensity. In general, temperatures in lichen and moss plots of the pine forests under study, are close to each similar, but in the first year after a fire a noticeable contrast was observed. The reasons are an increase in the amplitude of daily temperatures on the soil surface and stronger heating of upper mineral layers. Temperatures in the mineral layer with depths up to 30 cm depend on the thickness of the forest fl oor. Analysis of the results show that the duration of postfire effects in pine forests with sandy podzols is determined by a number of factors: the intensity of the fire, the degree of erosion of the ground cover and litter, and the recovery rate of these components.  相似文献   

8.
Five soil treatments in a 4-year-old clearcut in southern Sweden affected biomass increase and net nitrogen uptake by planted Norway spruce (Picea abies (L.) Karst.) seedlings through their on net mineralisation and root growth. The patch soil treatments studied were: (i) soil inversion in an old clearcut; (ii) mineral soil from the clearcut remaining, (iii) mineral soil from the clearcut with fertiliser application during the first season; (iv) mineral soil from a nearby uncut forest replacing the clearcut mineral soil; and (v) an untreated control. Growth increased in seedlings in treatments (i) and (iii), but growth in soil treatments with humus removal was not better than that of seedlings in untreated soil. High N uptake early in the first growing season resulted in increased growth during this season in contrast to late N uptake that resulted in a high N concentration in the seedlings after the first growing season. This in turn led to a high growth rate during the next growing season. Generally, both root growth and net N mineralisation were positively correlated to N uptake in the soil treatments. Therefore, a combination of low net N mineralisation and poor root growth as a result of high soil density appears to explain the low N uptake in seedlings in undisturbed soil. The importance of competition with field vegetation for N and water was not clear. Net mineralisation was larger in soil treatments where the humus layer was retained than where it was removed. Net N mineralisation in soil from old clearcuts was the same as in soil from fresh clearcuts.  相似文献   

9.
The effects of a large holiday resort on the distribution and range use of semi‐domesticated reindeer were studied in the three most important habitat types in Saariselkä, eastern Finnish Lapland. In two of these habitats the relative reindeer densities increased in winter, when outdoor activities declined reaching density equal to the average for the herding association at a distance of about 10 km from the holiday resort. In summer, male reindeer adapted better than females to the proximity of the holiday resort. The feeding site preferences of the reindeer during winter followed, in general, the normal pattern in areas with little human interference. Although the economic value of the range loss appeared small compared to the gain from outdoor enterprises, better planning is needed to avoid the harmful effects of recreation on reindeer management.  相似文献   

10.
Factors affecting seed germination, seedling establishment and growth to reproductive maturity in Eucalyptus pauciflora Sieb. ex Spreng. at the tree line in southeastern Australia were investigated. Timing of germination was determined principally by temperature but could be strongly modified by soil water. Mortality of germinants was heaviest during the first growing season (dependent on soil water) and the first winter (dependent on the depth and duration of snow cover). Neither plant nor soil treatments with insecticide or fungicide had an overriding or consistent effect on germination or seedling establishment. The probability of reaching the sapling stage was greater for planted seedlings than for plants germinated on site, indicating the dependence of survival on accumulated reserves. There was no clear indication of a general decrease in survival with increasing altitude, because of an interplay between elevation and site factors including competing vegetation, snow depth and period of snow cover. Five-year-old saplings planted up to 200 m above the tree line produced reproductive structures and viable seed between the ages of 13 and 18 years. The relevance of the results to an understanding of the location of the present tree line is discussed.  相似文献   

11.
Balisky  Allen C.  Burton  Philip 《New Forests》1997,14(1):63-82
A field trial was conducted investigating the single season growth response of 1+0 313 PSB Engelmann spruce (Picea engelmannii Parry ex Engelm.) and lodgepole pine (Pinus contorta Dougl. ex Loud.) seedlings planted into two different soil thermal regimes at three high-elevation locations spanning 200 km in the Engelmann Spruce-Subalpine Fir (ESSF) biogeoclimatic zone in the Cariboo Mountains of central British Columbia. Temperature treatments represented the extremes of soil temperature commonly found in high-elevation clear-cuts. A warm soil treatment (clear day, mid-afternoon soil temperature in mid-summer of 18 to 25 °C at –10 cm) consisting of a bare mineral soil hummock (average dimensions of 100 cm × 100 cm × 40 cm) was contrasted with a cool soil treatment (clear day, mid-afternoon soil temperature in mid-summer of 10 to 13 °C at –10 cm) comprised of organic forest floor overlying mineral soil. By the end of the growing season, seedlings of both species planted into the warm treatment generally exhibited greater root, stem, foliage, and total seedling biomass than cool treatment seedlings. Measurements of root growth at 30, 60, and 90 days after planting showed that total root number and total root length were consistently greater for warm treatment seedlings than for cool treatment seedlings. Root growth was greater from the bottom rather than from the side of the root plug for all seedlings. These results suggest that the effect of low soil temperatures on outplanted styroblock conifer seedlings is pronounced and may be limiting growth performance in high-elevation plantations in British Columbia. We recommend silvicultural treatments that secure natural regeneration, ensure that warmer microsites are always planted, and utilize seedling stocktypes able to make rapid lateral root growth into warmer surface organic horizons.  相似文献   

12.
Haywood  James D. 《New Forests》1999,18(3):263-276
Several mulches of natural, synthetic, or blends of natural and synthetic fibers were tested around newly planted loblolly pine (Pinus taeda L.) seedlings on a sheared and windrowed site in central Louisiana, USA. The vegetation was primarily winter annuals, some residual grasses and forbs, and sparse woody regrowth. Study 1 was rotary mowed just prior to planting in March 1992, and 35 mulches and an untreated check were established. In Study 2, 15 mulches and an untreated check were established in a 1-year-old rough in March 1993. In both studies, a single loblolly pine seedling formed each plot established in a randomized complete block design, with 10 blocks as replicates. Each block was planted with a separate open-pollinated loblolly pine family.Nearly all mulches had deteriorated to some extent after three growing seasons. Synthetic mulches were generally more durable than the natural or natural/synthetic mulches. Mulching eliminated the established vegetation and germinants, and vegetation did not readily reestablish following the deterioration of a mulch. The soil seed bank apparently was not sufficient to regenerate areas that were once covered with mulch and many of the natural materials deteriorated into a fibrous cover that acted like a natural litter layer. Both of these residual weed control effects -- insufficient soil seed bank and formation of a fibrous cover -- were important in stopping vegetation from reestablishing after a mulch had deteriorated. After three growing seasons, the loblolly pine seedlings generally grew better if mulches were used.  相似文献   

13.
Excessive slash loading could pose a problem for the regeneration of the serotinous lodgpole pine especially in forests at higher elevation where soil temperature is limiting. In the past, these forests have commonly been harvested using full-tree harvesting where trees are processed at roadside; however, recently cut-to-length harvesting has become a more frequent harvesting method. In cut-to-length harvesting the harvested trees are processed in the block, as a result slash accumulation is much higher on these cutblocks. In an experimental field trial, the cone distribution, natural lodgepole pine regeneration, and the growth and establishment of planted lodgepole pine were evaluated in response to slash load, drag scarification, and mounding after cut-to-length harvesting of high elevation lodgepole pine stands in the Rocky Mountains. Twelve sites were established, each contained six plots which were randomly assigned to six treatment combinations of two slash removal (slash and slash removed) and three mechanical soil preparation treatments (no soil preparation, drag scarifying, and mounding). The slash removal reduced slash volume by more than 50% but also reduced the number of lodgepole pine cones available for regeneration by over 33%. However, soil mechanical treatments offset this effect as fewer cones were necessary to achieve high natural pine regeneration densities. Drag scarification of plots resulted in 12 times the number of pine seedlings compared to the non-prepared plots. Although slash removal did not have an effect on the number of naturally regenerated lodgepole pine seedlings, it had a positive effect on their growth performance. Conversely, planted pine seedlings had lower mortality and better growth in soils that had been mechanically prepared and had the slash removed; however, the growth effects became only apparent 4 years after planting. While slash removal and mechanical soil preparation did increase soil temperatures; the slash removal treatment had a more transient effect on soil temperatures than soil preparation. Differences in soil temperature decreased over time which appeared to be mostly driven by a warming of the soils in the plots with no soil preparation, likely a result of the decomposition of the finer slash and feathermosses. Overall, it appears that surface disturbance on these high elevation sites had a far greater effect on lodgepole pine regeneration and growth than the increased accumulation of slash as a result of cut-to-length harvesting.  相似文献   

14.
Economic pressures have driven an ever-widening period during which foresters use machines to plant Scots pine (Pinus sylvestris L.) seedlings. In Fennoscandia, this period has recently stretched to the entire growing season. To evaluate the performance of seedlings planted during this extended period, three experiments were carried out in Central and Northern Finland over 2 years. One-year-old and current-year seedlings were planted in mounds or disc-trenched furrows when soil temperatures were >0°C. When 1-year-old seedlings grown for spring planting and overwintered outdoors were planted after mid-June, more needles browned and growth was reduced, possibly because seedlings were oversized with respect to planting density and the volume of growth media. When current-year seedlings sown in spring were planted from July to November, those planted in late September and October grew less in later years than those planted earlier, but survival was unaffected. No large differences in field performance were found with respect to whether seedlings were planted in mounds or disc-trenched furrows. In conclusion, Scots pine seedlings can be machine planted in mounds or furrows during May and early June (later in the North) and then continued from early August until late September, provided climatic conditions in late spring and early autumn are typical and similar to those experienced in Central Finland.  相似文献   

15.
Over the last few decades, the use of forests both by Sámi reindeer herders and for commercial forestry has been a source of increasing conflict in northern Sweden. Forestry disturbs forest ecosystems and thus damages reindeer pastures, especially those rich in ground lichen, for which reindeer forage during the wintertime by digging through the snow. Despite increased communication between the two groups, the Sámi still feel that their interests are not adequately considered by the commercial forestry sector. This paper presents the results of an interdisciplinary study that comprised semi-directed interviews and participant observations. The objective was to understand the Sámi reindeer herders’ extensive ecological knowledge of winter forest pastures, and the characteristics they observe when managing this resource. The study also analysed specific terminology used by the Sámi herders, to describe, analyse and communicate these properties. These terms, as well as Sámi herder knowledge in general, emphasize the importance of snow cover for reindeer grazing in forest pastures, as well as the effects of forest structure and ground vegetation on variations in snow cover during the winter. Whereas the Western use of the word ‘pasture’ is often associated with a specific plant community, the Sámi herders’ understanding of the word also includes the effect of snow on grazing and for this they use a culturally specific word in their language: guohtun. This term conveys the additional notion of whether it is possible for reindeer to access the pasture under the snow. Sámi herder knowledge and know-how allow them to use the mosaic of forest ecosystems to accommodate variability in snow cover during the wintertime, using different forest areas in response to different grazing conditions. Finally, the authors argue that, in order to promote the balanced, multiple use of boreal forest ecosystems, it is necessary to acknowledge and understand Sámi reindeer herders knowledge and thus to fully integrate their needs and aspirations.  相似文献   

16.

Survival, root egress and height growth of 13-month-old actively growing Norway spruce ( Picea abies (L.) Karst.) container seedlings exposed to preplanting drying treatments (0, 4, and 8 days) and postplanting drought periods (1, 2, 3, and 4 weeks) were studied in the field between 5 July and 18 August 1999. The mortality of seedlings increased and the height growth and root egress decreased throughout the postplanting drought period. Postplanting performance was also affected by preplanting drying treatments. The results indicate that no risk of excessive mortality and growth restrictions occurs when actively growing seedlings are planted in summer, provided that seedlings are well watered before planting and the drought period does not exceed 3 weeks.  相似文献   

17.
Three disturbance treatments were imposed on a palustrine forested wetland (Nyssa aquatica-Taxodium distichum) located in southwestern Alabama in 1986: (i) clearcutting with helicopter log removal (HELI), (ii) HELI followed by rubber-tired skidder traffic simulation (SKID) and (iii) HELI followed by removal of all vegetation during the first two growing seasons via glyphosate herbicide application (GLYPH). After two growing seasons, it was hypothesized that eventual woody plant growth would be best in the HELI-treated areas, because SKID plots had reduced rates of water movement and soil aeration. However, measurements at stand age seven years indicate that SKID actually has greater total above-ground biomass (65979 kg/ha) than HELI (46748 kg/ha) and SKID plots have a higher proportion of the most desirable timber species (Nyssa aquatica). GLYPH areas resemble freshwater marshes, although the areas are being invaded by Salix nigra seedlings. All disturbance treatments have significant groundflora components that have increased sediment accumulation 70–175% relative to an undisturbed reference area. By age seven years, regrowth of vegetation has lowered the water table during the growing season but has had little effect on soil redox potential and pH. Our observations suggest that this wetland system is rapidly recovering from logging disturbance seven years ago.  相似文献   

18.
Aspen (Populus tremula) is capable of reproducing both sexually and asexually from root suckers, the latter being more common. Seedlings are rare even though potential seed production is enormous. The aim of this study was to find out how different moisture conditions affect emergence and survival of aspen seedlings on burned soil. We set up a sowing experiment on mineral soil and on humus. The sowing was done at 240 microsites (θ7 cm) in 15 blocks (1×1 m). Treatments were sowing time, watering and sowing shelter. A total of 1463 seedlings emerged on mineral soil and 26 on humus. Watering, covering, their interaction, block and sowing time had significant effects on the number of seedlings. The largest numbers of seedlings were recorded at microsites that were watered and covered. The survival of seedlings was 20% during the growing season and 45% over winter. We conclude that seedbed conditions and moisture are the most essential factors for sexual reproduction of aspen. Short-term weather conditions have an important effect on seedling emergence and are likely to greatly affect field studies.  相似文献   

19.
A 3-year study measured the effects of ground cover treatments and N fertilization on biomass and nitrogen dynamics in an irridiated hybrid poplar (Populus deltoides Bartr. X P. trichocarpa Torr. and Gray, clone NC-9922) plantation in northern Wisconsin, U.S.A. Annually fertilized (112 kg N ha−1 year−1) and unfertilized plots were maintained weed free (bare soil), allowed to revegetate with native weeds, or seeded to birdsfoot trefoil (Lotus corniculatus L.). Biomass and N in trees and ground-cover vegetation were sampled before and after each growing season.Trees in bare-soil plots responded to fertilization primarily in the third growing season, but total biomass of 3-year-old trees was not increased by annual fertilization. In plots with a ground cover,fertilization increased tree growth but cover crop treatment had no effect. Ground cover biomass peaked during the second growing season, but declined thereafter, primarily due to reductions in below-ground biomass. Estimated recovery of fertilizer N was low in bare soil plots after 3 years, with 2% in the ‘perennial’ portion of the trees and 13% in the leaf litter. In contrast, recovery in the cover crop plots was 44%–51% in years 2–4. During that period, both biomass and N pool dominance shifted from primarily cover crop to primarily trees. The ground cover appeared to reduce tree growth in years 1–3, but total tree biomass after 4 years was greater in fertilized plots with ground cover (22.7 Mg/ha) than in fertilized bare soil plots (16.7 Mg/ha). Biomass production in fertilized trefoil plots in the fourth year (15.1 Mg ha−1 year−1, excluding leaves) exceeds that of local forests by 50%, and may be comparable to corn productivity in the area.  相似文献   

20.
Boreal forest ecosystems are generally highly sensitive to logging and other forestry activities. Thus, commercial forestry has had major effects on the forests and landscape structure in northern Sweden since the middle of the 19th Century, when it rapidly extended across the region. Lichens (which constitute up to 80% of reindeer forage in winter and early spring) have often been amongst the most severely affected ecosystem components. The overall aim of the present study was to analyze how forestry has influenced the potential supply of ground-growing lichens as winter forage for the reindeer in this region over the past ca. 100 years. For this purpose, we analysed changes in forest and stand structure in Scots pine-dominated (Pinus sylvestris L.) reindeer wintering areas in the southern part of the county Norrbotten (covering ca. 58,000 ha) using detailed historical forest inventories and management plans. We found that the amount of the forest types considered potentially good pasture (mainly middle-aged and old pine forests) decreased during the first part of the 20th Century. However, the quality of grazing grounds was improved by forestry during this time mainly because selective logging made the forests more open which benefits lichen growth. During the last part of the 20th century forestry impaired the quality of grazing grounds in several ways, e.g. by clear-cutting and intensified use of various silvicultural measures. We conclude that ca. 30–50% of the winter grazing grounds have been lost in the study area because of intensive forest management during the last century. The spatially precise historical information about the affects of forestry on lichen pasture provided in this study can be used to direct forest management which will facilitate and promote reindeer herding in the future.  相似文献   

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