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1.
Artificial freezing tests were performed with two sets of 12 full‐sib families of Picea abies. Each set was selected from a complete diallel cross performed within a natural population. Significant differences in autumn frost hardiness were observed between the two populations, which originate from the same altitude and longitude approximately 60 km apart. Substantial variation in frost hardiness was observed within both populations. Significant pheno‐typic correlations at the individual level were found between freezing injury and the two traits terminal bud‐set and height growth year one. However, no significant relationships were present between freezing injury and bud‐set at the family level, indicating that bud‐set cannot be used to predict autumn frost hardiness for families.  相似文献   

2.
Autumn frost hardiness of offspring from clones (originating from 66.0–68.8° N) in six southern (62° N) and one northern (66.5° N) Pinus sylvestris seed orchards in Finland, was evaluated. One‐year‐old seedlings derived from open pollinated grafts of young, intermediate, and fairly mature age, and natural stand seedlings, were artificially freeze tested. The northerly located orchard produced the least injured offspring. There were no significant systematic differences between age classes. Neither were there any indications of decreased freezing injury with later age classes. The family variance was significant in four orchards. The estimated injury levels of orchard offspring were, in general, closer to those of natural stands in the neighbourhood of the orchard than natural stands at the place of clonal origin. It is suggested that not only differences in mother clone hardiness but also clonal differences in pollen contamination rate caused the observed family variation.  相似文献   

3.
Reciprocal families of Scots pine, Pinus sylvestris (L.), from a Swedish seed orchard were artificially freeze tested to ‐10°C at the end of the first growth period. The degree of freezing damage was used as an indication of the cold acclimatization achieved at the time of freeze testing. Both one‐year cold acclimatization and one‐year height were mainly additively inherited. Specific combining ability, reciprocal and maternal effects were small. On the family level, freezing damage was non‐significantly correlated with field survival and field height after ten years. One‐year height was positively correlated with 10‐year field height and negatively correlated with field survival on the family level. Within families, plants from early germinated seeds cold acclimated earlier and were higher at the end of the first growth period than plants from lately germinated seeds. The results indicate that conclusions made from first‐year cold acclimatization studies can be influenced by variation in the rate of seed germination and seedling/germinant development.  相似文献   

4.
Seeds from open pollination of individual female stroboli on different days in June 1990 in a coastal clone archive of Pinus sylvestris (L.) in northern Sweden (latitude 63°39’ N) were used for a short‐term progeny experiment to study differences over time in the genetic composition of the background pollen. Analysis of progeny growth and cold acclimatization in artificial whole‐plant freezing tests suggested no genetic differences in the fertilizing pollen between pollination days. A clinal variation with the latitude of pollen origin was observed for reference progenies from fertilization with pollen of known geographical origin. Comparisons between open‐pollinated progenies and reference progenies indicate that the pollen cloud originated from approximately 1° latitude (based on height growth) or 1.5° latitude (based on freezing injury) north of the pollination site.  相似文献   

5.
The reliability of field, greenhouse and cut‐shoot screening procedures for the assessment of the susceptibility of Scots pine (Pinus sylvestris) genotypes to Melampsora pinitorqua, the causal agent of twisting rust, was evaluated. Fourteen genotypes evaluated on half‐sib progenies after natural infection of 2‐year‐old plants showed comparable rank for rust susceptibility with those evaluated after controlled inoculation of 1‐year‐old seedlings in a greenhouse. The dynamic of pycnia production was assessed at 2‐day intervals in cut‐shoot assays under controlled conditions, giving reliable rankings of Scots pine genotypes between years and being significantly correlated with rust susceptibility assessed after natural infection of 2‐year‐old progenies. These results underlined the importance of some critical factors such as inoculum load and host phenology in the reliability of susceptibility evaluations assessed under different experimental conditions. Artificial inoculation of 1‐year‐old seedlings in greenhouse experiments could provide a useful early test for the management of Scots pine breeding programmes and study of inheritance of twisting rust susceptibility. However, in specific investigations the cut‐shoot assay would constitute a reliable laboratory test for studying host–pathogen interactions and the variability in pathogenicity of Melampsora pinitorqua populations.  相似文献   

6.
No differences were found in the frost resistance of the different‐aged (2–7 weeks) germlings of Scots pine. The critical temperature was ‐6°C to —3°C. A temperature of ‐4°C killed half of the germlings and produced damage in one fifth of the surviving seedlings; the needles turned brown and growth ceased. The frost resistance of the previous year's shoots of two‐year‐old seedlings, measured by the impedance method, decreased during the shoot elongation period from — 10°C to — 4°C. Damage to the current shoots during shoot elongation appeared as deformation and discolouration of the needles and abnormal development or death of the apical bud. The results indicate that frost is not the cause of growth disturbances observed in Finnish nurseries.  相似文献   

7.
The time constant of the dehardening of one‐year‐old Scots pine (Pinus sylvestris L.) seedlings was studied in three experiments started at different times of the year. The seedlings were transferred from outside to the different controlled conditions in January (Exp. 1), March (Exp. 2) and April (Exp. 3). Changes in the frost resistance of the seedlings were followed on the basis of the temperature response of the specific impedance difference in the shoot. The time constant for dehardening at a temperature of 7.5°C was found to be about 12 days in Exp. 1, about 8 days in Exp. 2, and about 5 days in Exp. 3. the terminal bud burst when the frost resistance of the shoot had risen to about ‐10°C.  相似文献   

8.
After‐effects of maternal soil treatment, controlling for photoperiod and weather conditions, were tested in a nursery experiment with Pinus sylvestris L. seedlings. Three full‐sib crosses were replicated in 4 different soil treatment plots of a grafted experimental seed orchard (63"54'N). Weight of filled seeds, percentage of empty seeds, first year hypocotyl and epicotyl length, and autumn frost hardiness (from artificial freeze testing) were evaluated. Systematic effects from maternal soil treatment were small compared to family differences for all characters studied. However, epicotyl length of the progenies showed a positive relationship with N needle content of the mother grafts. The results also indicated an important interaction between maternal soil treatment and autumn frost hardiness of the families. It is suggested that the altered mother tree treatment could lead to a change in progeny performance, corresponding to a 1.5 degree difference in latitudinal origin of natural stand seedlings. The results demonstrate the advantages of providing the mother tree with uniform conditions in early progeny testing.  相似文献   

9.
短轮伐期日本落叶松家系生长性状遗传参数的变化   总被引：12，自引：0，他引：12  
以湖北省建始县长岭岗林场 15年生日本落叶松自由授粉家系子代测定林为研究对象 ,研究纸浆材轮伐期内不同年龄阶段生长性状的家系间遗传变异、早 -晚相关关系及主要遗传参数随时间变化的趋势 ,确定早期选择年龄及早期选择因子。结果表明 :各年龄胸径、树高不仅在家系水平上存在着极显著的变异 ,家系内个体间也存在着丰富的变异 ,具有很大的遗传改良潜力 ,家系和家系内个体选择是日本落叶松改良的主体 ;性状的方差分量、遗传力、变异系数在树木生长过程中一直处于动态变化中 ,并表现出一定的规律性 ;胸径、树高的早 -晚相关十分紧密 ,且这种相关关系主要受遗传因素制约 ;对胸径和树高进行家系早期选择的最适年龄为 6年生 ,进行家系内个体选择年龄为 8年生 ,胸径作为早期选择指标较树高更能准确地表达成熟时的信息。  相似文献   

10.
利用湿地松改良种子园的建园无性系生产的8个全同胞家系和20个自由授粉家系苗木共营建测定林3块。3－4年生的测定林数据分析结果表明，参试的自由授粉家系作全同胞家系的平均生长量显著地大于台山湿地松初级种子园的，20个自由授粉家系的树高、胸径、材积平均增益分别为5．54％，7．77％和23．41％，初步实现了建园目标；同一无性系的不同采种年份的自由授粉子代生长存在着差异，有随着种子园母树年龄的增大而表现渐佳的趋势；改良园中还存在着少量的误选无性系，需要继续作测定和淘汰。从无性系亲子的生长量相关分析中发现，生长较差的自由授粉家系，较大部分产自生长量缩小的母本无性系。  相似文献   

11.
银杏叶中化学成分的遗传变异   总被引：3，自引：0，他引：3  
对有代表性的 7个产区 ,每产区 4个家系及分布区南北两端 2个产区各 1家系内 10个单株 2a生幼苗叶片中 3种黄酮类化合物和白果内酯及银杏内酯的含量进行了测定 ,结果表明 :产区间、产区内家系间和家系内单株间均有广泛的遗传差异。黄酮类的含量与各内酯含量呈负相关 ,但达不到统计学显著水平。 2a生幼苗叶中的内酯含量与 16a生树的无明显差异  相似文献   

12.
Seedlings representing offspring from 46 Swedish natural stands of Pinus sylvestris (L.) from latitude 63° to 68° and altitude 75 to 675 m were artifically frost hardened and tested for autumn frost hardiness using artifical freeze testing in a programmable freezing chamber. A clinal variation in frost hardiness was observed over latitude. Altitude had no effect on the frost hardiness. The results are discussed in relation to the reproduction‐ and migration biology of P. sylvestris.  相似文献   

13.

The frost hardiness of non-juvenile Norway spruce [Picea abies (L.) Karst.] populations growing in northern Sweden (63°54' N) was monitored during 1996-1997. The investigated progenies originated from 12 natural populations and six seed orchards located between 58° N and 68° N in Sweden. Frost hardiness of needles was assessed by measuring chlorophyll fluorescence and electrolyte leakage after freezing. The loss of frost hardiness in 1-yr-old needles during spring occurred slightly earlier in populations originating north of 63°30' N than in those originating further south. Dehardening was slightly delayed in selected populations compared with natural populations of similar origin. The level of frost hardiness during autumn was higher in populations originating north of 63°30' N than in those originating south of this latitude, but there were no clear differences in frost hardiness between selected and natural populations of similar origin. The results are discussed in relation to climatic factors and inherent growth rhythms.  相似文献   

14.
The magnitude and practical importance of family × site interactions for growth and form traits considered in radiata pine (Pinus radiata D.Don) breeding were analysed by four different methods: type B genetic correlation, regression coefficient, mean rank deviation and ecovalence. The material analysed consisted in growth, form and frost resistance assessments of 58 open pollinated half-sib families at 3 to 4 years old across three sites in Galicia (NW Spain). Analysis of variance revealed that family × site interactions were quantitatively important for all traits (diameter, volume, branch angle, straightness and frost resistance). The losses in potential genetic gain for a breeding population were important for all traits under individual selection but only for diameter and frost resistance under family selection. Omission of the ten most interactive families from the analysis considerably reduced the losses for all traits, although losses in potential genetic gain remained important for frost resistance under family selection. The results indicate that elimination of these families from breeding programmes would be an effective strategy for selecting for stability in performance across sites for all traits except frost resistance. In order to overcome this problem, selecting varieties for frost resistance specifically adapted to various regional contexts would be an option. However further studies under controlled conditions are required before making final decisions for radiata pine breeding programmes.  相似文献   

15.
In a 14-week study, 1-year-old Aleppo pine seedlings were grownin two growth chambers. Seedlings were artificially hardenedby decreasing photoperiod and temperature. In each chamber halfof the seedlings were fertilized with nitrogen (8.4 mg seedling–1).In order to determine the relative importance of the hardeningenvironment versus fertilization, each chamber was programmedto decrease night temperatures down to a low of 8 or 4°C.Chlorophyll fluorescence and frost hardiness was measured fivetimes during the experiment. A sample of seedlings from eachtreatment was exposed to an artificial frost at –5°Cand the freezing effects were assessed by measurements of chlorophyllfluorescence and visual evaluation of needle damage. Seedlingsincreased their frost hardiness during the experiment in allthe treatments but the ratio of variable to maximal chlorophyllfluorescence (Fv/Fm) measured before freezing did not vary duringthe experiment. This indicates that Aleppo pine maintains itsphotosynthetic ability during hardening in contrast to otherconiferous species from colder climates. The effect of nitrogenfertilization on frost hardiness was small in comparison withchamber effect. Nitrogen fertilization slightly delayed theacquisition of hardening in the coldest chamber. Seedlings inthe warmest chamber did not become fully resistant to –5°C,but in the coldest chamber, where night temperature reached4°C, all the seedlings were resistant to the frost. Severedamage caused by frost could be related to a rapid rise of minimalfluorescence (F0) but the best index of damage was the dropof Fv/Fm after freezing.  相似文献   

16.
The susceptibility of Pinus oocarpa to freezing temperatures limits the commercial deployment of the highly productive Pinus patula × P. oocarpa hybrid in South Africa. Identifying P. oocarpa germplasm with increased frost tolerance is important. Twenty-three P. oocarpa provenances, originating from Mexico, Honduras, Guatemala and Nicaragua, were therefore assessed for their tolerance to freezing conditions by analysing field survival after frost events, subjecting needles to freezing temperatures and assessing damage using the electrolyte leakage test, and exposing young plants to freezing temperatures in a semi-controlled environment and scoring tissue damage based on a visual assessment. The performance of many of the provenances represented in the field and artificial freezing studies were similar and there was a strong correlation between provenance ranking in the whole plant freezing and electrolyte leakage test. We therefore support the use of these techniques as a means to assess cold tolerance in P. oocarpa at the provenance level. Provenances from north-western Mexico demonstrated more frost tolerance than those from southern Mexico. Provenances representing Honduras and Guatemala appear to be highly susceptible to frost.  相似文献   

17.
Cuttings were made from six‐year‐old seedlings of Picea abies L. (Karst.) that had been studied in the Stockholm phytotron. The seedlings belonged to four full‐sib families, one being of French origin, one of North‐Swedish origin and two being hybrids between parents of North‐Swedish and French origin. Nine clones from each family were included. After the rooting phase, the cuttings were exposed to the same cultivation regimes as the seedling ortets. Characteristics of growth rhythm, percentage free growth and leader length were recorded during the 2nd to the 5th growth period (GP). Plagiotropic growth habit was recorded in the 3rd to the 5th growth period, and, characteristics of the rooting phase were studied. A juvenile behaviour very similar to that characterizing growth periods 2–3 of the seedlings was observed for growth rhythm characteristics and for free growth capacity during GP2 of the cuttings. The absolute value of the differences between each observation of the seedling ortets in GP6 and the means of the cuttings in GP2 produced a new variable that quantified the degree of rejuvenation. This new variable was meaningfully calculated only for growth rhythm and growth pattern variables. A gradual reversion of the growth rhythm occurred for the cuttings, which corresponded to the change with increasing age for the seedlings. The capacity for free growth, on the contrary, disappeared already in GP3 for the cuttings, except for the French family. A large variation in the rejuvenation ability between families was observed. Furthermore, the rejuvenation ability was negatively correlated with the ability to grow plagiotropically.  相似文献   

18.
This study examined 108 15‐year‐old Douglas‐fir (Pseudotsuga menziesii var. menziesii) trees to investigate whether trees exhibiting less severe Swiss needle cast (SNC) symptoms were more resistant (had less fungal colonization) or more tolerant (maintained healthy foliage under similar infection levels). Trees were sampled from six open pollinated families that were categorized into three disease severity groups (two families for each group; mild, moderate and severe disease symptoms). The amount of retained foliage and level of discoloration were visually assessed on trees in the field. Fungal colonization (as determined by proportion of stomata occluded with pseudothecia and by amount of Phaeocryptopus gaeumannii DNA in sampled needles) was measured on 1‐ and 2‐year‐old needles in the laboratory. Trees in the different disease severity groups were similar with respect to amount of fungus in their needles, yet the trees in the mild symptom group retained higher proportions of needles and maintained greener foliage. The relationship between amount of P. gaeumannii in needles and SNC symptom severity was statistically significant (p < 0.05) for amount of fungal DNA in 1‐year‐old needles and average needle retention (NR) over the last four growing seasons. Average NR decreased with increased amount of pathogen DNA in the mild disease symptom families. This relationship was reversed in the severe disease symptom group and there was no relationship in the moderate disease symptom group. Because the amount of P. gaeumannii DNA in foliage did not differ significantly among the groups, differences in symptom severity were attributed to tolerance, not resistance. Visual scoring of individual trees for average NR over the past four growing seasons could be used to effectively assess for SNC tolerance in Douglas‐fir.  相似文献   

19.
Plants utilize two general strategies to cope with pathogen attack. They either limit or resist the pathogen (termed ‘resistance’) or they cope with the disease by surviving and growing (termed ‘tolerance’). Both strategies tend to increase plant fitness; however, there are possible costs, trade‐offs and interactions associated with each strategy. This study focused on five half‐sib interior Douglas‐fir families [Pseudotsuga menziesii var. glauca (Beissn.) Franco] that were putatively classified as either resistant or susceptible to Armillaria ostoyae in a previous greenhouse study of seedling families challenged with A. ostoyae. We compared their survival rates in the greenhouse study with results of juvenile trees from the same five families that were artificially inoculated in field conditions. Both resistance and tolerance appeared to be operating in the field test trees, and a possible trade‐off between resistance and tolerance was detected. Significant differences were detected among the five families for stem radial growth following infection at the tree root collar. Compared with the putatively susceptible families, resistant families had smaller lesions and lower proportional root collar girdling. Tolerant families generally had larger lesions but demonstrated better growth when diseased than resistant families. One tolerant family that was a good survivor in the greenhouse survival study presented vertically shaped lesions that were large in area but had greatly reduced proportional root collar girdling. While a second family showed tolerant traits in the field study, its poor survival in the greenhouse study agreed with the large horizontally spreading lesions associated with high root collar girdling in the field study trees. Survival rankings of the five families in the greenhouse study mostly agreed with results from the field study based on the proportion of collar girdling among families. These host responses are discussed with respect to stability, quantity and quality of stands and products.  相似文献   

20.
In full‐sib families of Picea abies (L.) Karst. (Norway spruce) the dynamics of juvenile growth and the inheritance of the most important growth characters were analysed in six growth periods in climate chambers followed by two additional growing seasons in a nursery. The full‐sib families were of French and North Swedish origin including both within and between provenance crosses. All growth rhythm characters changed over time. For instance, number of days to budbreak increased, whereas days to budset decreased, as growth periods advanced. Concurrent with the changes in growth rhythm, free growth decreased with increasing age. Half of the total variation for different growth rhythm characters within a growth period could in most cases be explained by provenance effects. Early selection is, therefore, meaningful to reduce damage by frost and mortality. Furthermore, the actual distances between family means decreased substantially with increasing age, and the actual length of the shoot elongation period decreased from 31 days during the second growth period to 16 days during the sixth. We suggest therefore that selection together with some treatment in the nursery that alters the growth rhythm of the seedlings are the best ways to increase frost resistance. Tree height was influenced by a set of complex gene actions that changed over time. Provenance effects were important during the first two growth periods. Subsequently, non‐accumulative within provenance family variance took over. If this pattern holds true in general, juvenile selection for height growth in Norway spruce cannot be made as early as that for growth rhythm.  相似文献   

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