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1.
The growth and structural development of Scots pine (Pinus sylvestris L.) trees growing at different spacing was simulated using a model based on the dry matter production per needle biomass unit and its allocation to needles, branches and stem. Special emphasis was given to the effect of stand density on the growth of the crown system and its implications on branchiness and timber quality. The simulations showed that the needle biomass culminates considerably earlier than the branch biomass with a time lag inversely related to the stand density. The lengths of living and dead crown were also inversely related to stand density. The resulting differences in branchiness were especially obvious in the early development of the tree stands. In the long run these differences tend to disappear, indicating equal external branchiness independently of the initial spacing for mature stands of Scots pine. The internal branchiness, however, was particularly sensitive to the initial spacing.  相似文献   

2.
A crown profile model was developed for radiata pine (Pinus radiata D. Don) in Galicia (northwestern Spain). Data from 443 trees located in 56 permanent plots, established by the Unidade de Xestión Forestal Sostible (UXFS) of the University of Santiago de Compostela in plantations of this species in the region, were used. The crowns of the trees were measured by a visual method based on similar triangles. Both simple geometric shapes and mathematical equations were used to describe the crown profiles. As crown profile models usually require variables that are expensive to measure, equations to estimate the maximum crown radius and the height to the maximum crown radius were also developed, using other easily measured tree and stand variables. Several models were fitted using a system of equations approach and accounting for an autocorrelated, heteroscedastic error structure. The selected crown profile model consists of a system of two allometric equations for the crown below (primarily shade needles) and above (primarily sun needles) maximum crown radius. The model explained 88% of the variability in crown radius with a mean error of 0.24 m.  相似文献   

3.
Precommercial thinning is regarded as one of the most important measures for influencing timber quality in stands of Scots pine (Pinus sylvestris L.). The present study considers the influence of both thinning intensity and tree height at the time of thinning on diameter of the thickest branch and crown ratio. A total of 115 plots (either naturally regenerated, planted or sown, and either thinned down to 528–8000 stems?ha?1 or untreated) included in 20 sites in southern Sweden were analysed. The average tree height after thinning varied from 1.2 to 8.3?m. An increasing number of remaining stems resulted in a reduction in branch diameter, although the reduction appeared to be only minor if the number of stems after thinning was more than 3000 stems?ha?1. It was found that late thinning reduced the diameter of the thickest branch. The crown ratio decreased with stand height, number of stems after thinning and average height at thinning. The results were consistent for all trees and for the 500 thickest trees per hectare.  相似文献   

4.
Abstract

This study compares the results of the prediction of crown height characteristics using airborne laser scanner (ALS) data and intensive field measurements in boreal forests. The data consisted of 31 sample plots located in Kalkkinen, southern Finland. Crown height models were constructed at both the tree and plot level. Scots pine, Norway spruce and birches were used. The models included independent variables of tree levels, such as tree height, crown area and independent plot-level variables, i.e. canopy height and density quantiles and proportion of vegetation hits. Field measurement-based models used tree height and diameter at breast height as the independent tree-level variables, whereas basal area, mean diameter and height were used as the plot-level variables. The results indicated that the ALS-based crown height models were more accurate than the field measurement-based models when plot-level information was used as independent variables. However, the field measurement-based tree-level models for Scots pine and Norway spruce were more accurate than the ALS-based models. Even so, the accuracy of the different models was very similar and the study data set was quite small. The results of this study can be used for different tree growth studies and for the assessment of tree stock quality in boreal forests.  相似文献   

5.
The research described in this paper was performed in the Niepolomice Forest (Southern Poland) in 2001 as part of the Forest Environmental Monitoring and Management System (FOREMMS; 5FP IST) project. The material for the present study consisted of the measurement results of the biomass of Scots pine shoots with needles and needles alone carried out on 113 felled sample trees. The purpose of this study was to construct empirical equations for estimating the foliage biomass of Scots pine from easy to measure parameters. To achieve this aim, the dependence of the foliage biomass of Scots pine on stem diameter, height, age, crown length, basal area increment of the trees was analyzed. Using the biometric characteristics such as: tree diameter at breast height (dbh), basal area increment, age, height, and crown length empirical equations for estimating the foliage biomass of Scots pine reasonably precisely have been established. The created empirical equation gives accurate foliage biomass estimates. The explained variability varies between 65 and 85%, it depends on the number of variables applied in the equation. The equations presented in this paper were created with a view to their possible use in ecological studies where biomass quantity may be used, for example, in modeling carbon circulation in the forest ecosystem. From the point of view of forestry practice, these equations may help to assess biomass production in Scots pine stands.  相似文献   

6.

Reducing competition by point cleaning may be an attractive alternative to conventional precommercial thinning in forestry. In this study, survival and development of main stems of Scots pine (Pinus sylvestris L.) following point cleaning were examined in a mixed stand of mainly Scots pine and birch (Betula pendula Roth and Betula pubescens Ehrh.) in northern Sweden. Treatments studied were: no precommercial thinning, conventional precommercial thinning, point cleaning of all secondary stems, and point cleaning of only the secondary stems that were higher than the main stem, within a radius of 0.8 m from the main stem, respectively. A randomised complete block design with three replicates was used. Measurements taken 13 growing seasons after these treatments showed that conventional precommercial thinning was the best treatment for retaining Scots pines as main stems. However, conventional precommercial thinning resulted in the poorest timber quality of the remaining Scots pines in terms of stem taper, thickest branch diameter and relative live crown height. In conclusion, point cleaning has to be followed by a complementary cleaning operation if the objective is to retain Scots pines as main stems in such mixed stands.  相似文献   

7.
The role of stand and site characteristics in a 50‐year‐old, sowed Scots pine (Pinus sylvestris L.) stand on the decline of Scots pine caused by Gremmeniella abietina was studied. Treewise damage of Scots pine was modelled using stand and site variables and stepwise regression analysis. The significant variables included in the model were stand elevation from the main cold air centre and the pH of the humus layer, while the other less significant variables were the number of stems per hectare of Scots pine and Norway spruce, content of Cu and Ni in the humus layer, and the thickness of the humus layer and the A2 horizon. The studied variables excluded from the model were the number of broad‐leaved trees per hectare, Al and Pb contents in the humus layer, and Al, Cu, Pb and Ni contents in the A2 horizon. The results strongly suggest that the vicinity of water as well as local depressions act as vulnerable sites for Scots pine and the said vulnerability is enhanced by increased pH of the humus layer.  相似文献   

8.
G. abietina causes severe dieback in pole-stage stands of Scots pine in Britain. The susceptibility of ll provenances of Scots pine and of three other conifer species was tested by inoculations. Significant differences in susceptibility were found amongst Scots pine provenances, with those originating from a native Scottish pinewood at Loch Maree showing the highest levels of disease. Corsican pine was shown to be the most susceptible species tested although sporulation was greatest on Scots pine. Norway spruce was also shown to be susceptible when inoculated late in the growing season. Infection in lodgepole pine was negligible. High levels of β-phellandrene were tentatively linked with resistance among trees in a single stand of Scots pine.  相似文献   

9.
Altogether 82 plots (261 estimations) of Picea abies (L.) Karst, and 193 plots (360 estimations) of Pinus sylvestris (L.) stands were estimated by a vertical tube. The “crown free projection”, CFP, of stands thinned in three methods with different thinning grades was measured: unthinned, heavily and very heavily thinned, heavily thinned delayed first thinning, extra heavily thinned and thinned from the top. Basal area (m2ha?1) density (stems ha?1) and diameter sum (m ha?1) were plotted against CFP. Basal area was the best practical measure of stand in this study. Generally Scots pine stands have higher CFP and the curves are steeper than in Norway spruce stands. Depending on the grade of thinning, heavily and very heavily thinned spruce stands, delayed first thinning included, have CFP values of 10–15% and stands thinned from the top, 20–40%, compared with 30–80% and 30–60% respectively in pine stands. Extra heavily thinned stands have the highest CFP, 20–80% in spruce and 50–90% in pine stands. The CFP levels after thinning are too high in pine stands for avoidance of sucker and sprout production of aspen and birch. In dense Norway spruce stands thinned from the top or heavily and very heavily thinned, the CFP values are low enough (≤30%) to diminish the production of suckers.  相似文献   

10.
基于大比例尺航片的针叶树种冠幅的提取   总被引：2，自引：0，他引：2  
基于凉水国家级自然保护区2009年拍摄的1：2000航空像片和同期森林资源二类调查的固定样地数据,采用子像元分类方法分别提取出红松、落叶松和云冷杉的专题影像图。在此基础上,将栅格专题影像图转换为矢量图形,采用目视解译的方法提取上层针叶林的树冠信息。通过将针叶树冠形似为圆形提取出各树种的冠幅,用固定样地实测数据进行对比分析和精度评价,并建立航片上提取冠幅与实测冠幅之间的一元线性回归模型。结果表明：红松、落叶松和云冷杉冠幅的提取精度分别达到83．50％、84．35％和82．26％,其预测精度分别达到83．60％、81．46％和83．57％。  相似文献   

11.
It was hypothesized that maximum living crown removal can be determined from experimental data using linear statistical models. Pruning experiments were carried out in four Scots pine (Pinus sylvestris L.) stands. Zero to nine living branch whorls were removed in three stands in southern Finland, and 0–4.5 m of the living crown was removed in a stand in northern Finland. Only dead branch whorls were removed from the control trees. A total of 517 trees were analyzed. Diameter increment was measured five years after pruning. The growth reduction was statistically significant when 40% or more of the living crown was removed by pruning. According to the linear models, green pruning decreased diameter growth by 0 to 33% at defoliation levels below 40%. This loss in diameter growth is, from an economical point of view, compensated by the knotfree timber resulting from pruning 2–3 years earlier.  相似文献   

12.
The patterns of current‐year shoot, needle and terminal bud elongation in seedlings of three Scots pine (Pinus sylvestris L.) and three lodgepole pine (Pinus contorta Dougl. var. latifolia Engelm.) provenances were compared during the third and fourth growing seasons after planting. Lodgepole pine produced longer shoots and buds than did Scots pine, mainly because lodgepole pine formed more stem units and elongated at a faster rate. Stem unit length and the duration of shoot and bud elongation differed relatively little between species and provenances. Lammas or polycyclic growth occurred in some lodgepole pine provenances, but not in any Scots pine provenance, and was associated with enhanced shoot elongation. Needle elongation commenced earlier, proceeded at a faster rate, and was greater in lodgepole pine than in Scots pine, but ceased about the same time in all species and provenances. The heat sum required to attain 50% of final length was lower for shoots and needles in lodgepole pine than in Scots pine, and for shoots in northern provenances than in southern ones. Mitotic activity in the apical meristem of the terminal bud, which occurred less than one week after the seedlings were free from snow, started and ceased about the same time in each species, but was higher in lodgepole pine than in Scots pine early in the shoot elongation period.  相似文献   

13.
The aim of this study was to investigate the differences in infections caused by Agrobacterium tumefaciens in a conifer, Scots pine (Pinus sylvestris), and in a non-host deciduous species, silver birch (Betula pendula). All the Agrobacterium tumefaciens strains tested caused crown-gall formation in both tree species, but the infection rates varied remarkably. In Scots pine, the development of galls was rare, and slower than in silver birch. Inoculation into the base of the stem were the most successful in gall induction. Silver-birch galls were large, often surrounding the whole stem, in contrast to Scots pine galls, which were characterized by their small size and neck-like connection with the host plant. In silver birch, no other morphological changes could be seen. In Scots pine, abnormal phenotypes with proliferating short shoots above the galls were observed during the second and third growing season. The results indicate that, of the two non-host tree species, the deciduous one, silver birch, is more susceptible to an A. tumefaciens infection than the conifer, Scots pine. The matrix for A. tumefaciens infection in silver birch differs from that in Scots pine, since the terpene compounds of Scots pine seem either to kill the agrobacteria or to suppress their growth. The differences between the species could be partly caused by their difference in sensitivity to phytohormones. These features reflect evolutionary incompatibility between A. tumefaciens and a gymnosperm.  相似文献   

14.
Summary The aim of this study was to screen Scots pine embryogenic tissue culture material to Gremmeniella abietina. According to the results Scots pine (Pinus sylvestris) embryogenic tissue lines (27 lines) could be ranked based on significant differences in G. abietina effects estimated by changes in glucosamine concentrations, growth, and dry matter contents of inoculated tissue pieces. The ranking order of the extreme tissue lines stayed basically similar, regardless of the variable measured. These findings imply a possibility to use the ranked embryogenic tissue lines in more detailed investigations of interactions between G. abietina and its host. This is the first report of using embryogenic tissue cultures in studying host–parasite interaction between Scots pine and G. abietina.  相似文献   

15.
The effects of stand density, thinning and pruning on the quality of sawn timber of Scots pine (Pinus sylvestris L.) were studied based on a computer model. Procedures for the tree growth and crown structure produce a tree, whose properties in terms of location, dimensions and quality (living, dead) of branches are known for any moment of the selected rotation. Consequently, the size, quality (living, dead) and location of knots in sawn timber are recognizable allowing the grading of sawn pieces. The model computations showed that the natural dynamics of the crown system in narrowly‐spaced stands could yield sawn timber nearly of the same quality as resulted from pruning in widely‐spaced stands. Thinning increased the branch growth and the branchiness of the wood yielding lower grade for sawn pieces, respectively.  相似文献   

16.
In this study, the effect of pine mistletoe (Viscum album subsp. austriacum) on basal area increment of Crimean pine and Scots pine was investigated. Dendrochronological data were collected from 223 (71 uninfected and 152 infected) Crimean pines and 195 (77 uninfected and 118 infected) Scots pines located in Kastamonu province of Turkey in 2014. Infected sample trees were classified as light, moderate or severe infection levels. Growth trends and basal area increment loses were compared between uninfected and infected trees for the periods of the last 10, 20 and 30 years. In addition, infection status of forest stands was investigated using temporary sample plots; 27 plots in Crimean pine stands and 26 plots in Scots pine. Results demonstrated that basal area increments were negatively affected by pine mistletoe for both species. Mean basal area increment losses of infected trees for the last decade were determined as 24% for Scots pine and 26% for Crimean pine. Basal area increment losses varied by infection levels (light, moderate and severe) as follows: 25%, 20% and 28% for Scots pines and 20%, 32% and 9% for Crimean pines. Scots pine stands were more severely infected by pine mistletoe than Crimean pine stands. There were negative correlations between number of infected trees and stand density for both species, while positive correlation was detected between the number of infected trees and mean diameter for Scots pine. The results of this study indicate that the pine mistletoe infection has negative effect on radial growth of Scots pine and Crimean pine trees. The results can be an important contribution to the forest management and protection activities in mistletoe-infected stands.  相似文献   

17.
Scots pine (Pinus sylvestris) and lodgepole pine (Pinus contorta) needle litters were compared in terms of nutrient composition and its change during decomposition. Initial nutrient composition differed between the species, with lodgepole pine needle litter having significantly higher concentrations of P, Mg and Mn. However, no difference was found for concentrations of N, Ca or K. Increases in concentrations of N, P and K during decomposition were significant in both litter types. For Ca the pattern of concentration changes followed a quadratic function as decomposition proceeded. Concentrations of Mg and Mn decreased in lodgepole pine needle litter. In Scots pine litter there was also an initial decrease, but it was followed by an increase in most incubations. For both Mg and Mn, changes in concentrations during decomposition differed significantly between species. In the late decomposition stages, concentrations of Mg and Mn became similar in both litter types. Nutrient concentrations generated by the models were compared with those of the humus (F and H) layer in the stands. The model was quite accurate in predicting concentrations of N and P for both species and the concentration of Mg for lodgepole pine. By contrast, it was not accurate in predicting concentrations of Ca and Mn. Nutrient release was estimated for the two species using both measured litterfall data and long‐term estimates, and regression models were used to predict concentration changes. Rates of release of P, Mg and Mn in the lodgepole pine stands were found to be about twice as high compared with those in Scots pine. Calcium was also released to a greater extent although the difference was not significant.  相似文献   

18.
Abstract

Pine stumps are not being treated against Heterobasidion spp. in Sweden. To determine whether they should be, the frequency of stump infections and the species of Heterobasidion involved were investigated in nine newly thinned pure Scots pine (Pinus sylvestris L.) stands in southern Sweden. The incidence of Heterobasidion was measured in roots of standing Scots pine in another 15 stands. Infections by both H. annosum (Fr.) Bref. s.s. and H. parviporum Niemelä & Korhonen were numerous in stumps six months after a summer thinning. The pathogen, mostly H. annosum, was found in 44 of 60 sampled root systems, from 14 of the 15 stands. Twenty of the infected pines were assessed as healthy on the basis of crown symptoms while 24 trees had defoliated crowns. Infected root systems were most frequent among trees with thin crowns in stands on former agricultural land, where previous thinnings had been carried out during the growing season when airborne spores are plentiful. The study suggests that stump treatment may be a profitable way to reduce disease development in Scots pine monocultures on sandy soils as well as in mixtures with Norway spruce [Picea abies (L.) Karst.] on any soil.  相似文献   

19.
We tested hypotheses on the effect of growing space and biomechanics on tree growth, using data from two field experiments where Scots pine seedlings had been planted in a fixed but systematically irregular pattern. After 16 or 18 years, respectively, stem diameter was measured at 1.3 m (d1.3) and 0.3 m above ground (d0.3). Total tree height, length and diameter of the crown, and height to crown gravity point were also measured. Analyses were split in two parts: influence of growing space on tree and crown dimensions, and biomechanical influence of the crown on stem diameter. The results showed that crown size was closely related to tree height, and that there was a small but statistically significant effect of growing space on both crown size and shape. The biomechanical model explained about 85% of the variation in stem diameter, but growing space had no influence on this relationship, indicating that the wind load absorbed by a given crown size was not affected by growing space. The results were discussed in relation to future growth modelling and new ways of estimating site productivity.  相似文献   

20.
Increased mortality rates in Scots pine (Pinus sylvestris) forests have recently been observed in the inner alpine Swiss Rhone valley. Drought, in combination with stand competition, mistletoe infections as well as nematode and insect infestations, appears to be the main factor for the decline. In focus of this study was the occurrence and role of fungal pathogens in the decline dynamics. Branches, stems and roots of 208 trees in five different crown transparency classes were collected and inspected for blue stain and fungal infections. Neither Armillaria species nor Heterobasidon annosum s. str. were detected, but blue stain was commonly observed. Visible blue stain increased with increasing crown transparency. Among the recently dead trees, 80% showed visible blue stain in the branches, 90% in the roots and 100% in the stems. In the crown transparency classes 2 and 3 (25–60% crown transparency), five of the 103 trees showed visible blue stain in the roots, one of 130 trees in the stem but none in the branches. Blue‐stain fungi were isolated from all parts of the trees and from all crown transparency classes. Overall incidence of blue stain was highest in the roots and lowest in the branches. In class 2, roots of 60% of the trees were visibly blue‐stained or developed blue stain in culture, but stems of only 24% and branches of 14% of the trees. In the roots Leptographium species, mostly L. serpens, dominated. From stems and branches, mainly Ophiostoma species were isolated. The positive relationship between the incidence of blue stain and crown transparency, in combination with the high infection levels of roots of fairly vigorous Scots pines, indicates the pathogenic potential of the blue‐stain fungi. Hence, these fungi together with their insect vectors may well act as an important contributing factor involved in pine decline.  相似文献   

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