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1.
The effects of stand density, thinning and pruning on the quality of sawn timber of Scots pine (Pinus sylvestris L.) were studied based on a computer model. Procedures for the tree growth and crown structure produce a tree, whose properties in terms of location, dimensions and quality (living, dead) of branches are known for any moment of the selected rotation. Consequently, the size, quality (living, dead) and location of knots in sawn timber are recognizable allowing the grading of sawn pieces. The model computations showed that the natural dynamics of the crown system in narrowly‐spaced stands could yield sawn timber nearly of the same quality as resulted from pruning in widely‐spaced stands. Thinning increased the branch growth and the branchiness of the wood yielding lower grade for sawn pieces, respectively.  相似文献   

2.
Silvicultural cleaning is commonly carried out in young stands to control competition between conifers and deciduous tree species. However, it has been questioned whether this practice affects moose (Alces alces L.) browsing on Scots pine (Pinus sylvestris L.) with respect to damage at the stand level. In this study, moose winter browsing was examined in relation to food availability in 19 Scots pine stands divided into silviculturally cleaned and non‐cleaned blocks. Cleaning had a negative effect on moose foraging in terms of reduced density, biomass availability, and height of the deciduous tree species. The total biomass consumed by moose was significantly lower on the cleaned blocks than on the non‐cleaned ones. There were no differences between the treatments in the amounts of pine consumed and in moose damage to pines. The density or biomass available of different deciduous tree species did not explain the variation in moose browsing on pine. In conclusion, timing of silvicultural cleaning with reference to moose browsing should be taken into account. If pines are not overtopped by deciduous trees, as in the present study, silvicultural cleaning is not necessary to prevent moose damage.  相似文献   

3.
Abstract

The objective of this study was to compare volume growth and external timber quality properties of lodgepole pine (Pinus contorta Dougl. var. latifolia) stands planted in different spacings (1.41×1.41, 2.00×2.00, 2.83×2.83, 4.00×4.00) in southern Sweden. In northern Sweden lodgepole pine has been grown for a long time and shows superior volume production compared to Scots pine (Pinus Sylvestris L.). The Swedish Forestry Act restricts establishment of lodgepole pine plantations in southern Sweden. However, it is important to increase the knowledge about lodgepole pine and its potential use also in this part of the country. Two experiments in southern Sweden were studied. Each trial was designed as a random block experiment with two blocks. Both investigated stands had an age of 23 years at the time of assessment. The mean diameter and the volume production differed significantly between the spacings. The highest volume production was found in the narrowest spacing, 313% compared to the widest spacing. Significant differences between spacings were also found concerning external quality traits. The frequency of dominant and co-dominant trees without defects was 24% in the most open spacing compared to 46% in the narrowest one. Under current circumstances approximately 2500 seedlings/ha in the initial stand seems to be a reasonable compromise between growth, diameter development and timber quality.  相似文献   

4.
The branch biomass of young Scots pines (Pinus sylvestris L.) was inversely related to the branch number per projected crown area giving greater branch density and smaller branch biomass for narrow crowns than for broad crowns. In particular, in dense stands the small share of branches from the total biomass of narrow‐crowned trees was emphasized. The productivity of narrow crowned trees was, however, smaller than that of trees with broad crowns. The concept of the tree ideotype has been discussed on the basis of theoretical calculations.  相似文献   

5.
Abstract

The quantitative effect of pine-dominated edge stands in southern Finland on the density and growth of pine saplings was addressed in the study. The cumulative effect of a mature edge stand on the development of a pine sapling stand with variable cutblock sizes was also estimated. The data were acquired from 10 study sites with a Scots pine sapling stand with a current dominant height of 1–6 m and an adjoining middle-aged to mature edge stand dominated by Scots pine. Tree level models were constructed to describe and simulate the structure and development of the sapling stands. The results indicated that dividing an average regeneration area (2 ha) into smaller cutblock units (1.0–0.5 ha) appreciably increased the edge stand effect. Total stand volume growth was 6% lower on 1 ha cutblocks, and 31% lower on 0.5 ha blocks, than on 2 ha blocks.  相似文献   

6.
Variations in fine root biomass of trees and understory in 16 stands throughout Finland were examined and relationships to site and stand characteristics determined. Norway spruce fine root biomass varied between 184 and 370 g m(-2), and that of Scots pine ranged between 149 and 386 g m(-2). In northern Finland, understory roots and rhizomes (< 2 mm diameter) accounted for up to 50% of the stand total fine root biomass. Therefore, the fine root biomass of trees plus understory was larger in northern Finland in stands of both tree species, resulting in a negative relationship between fine root biomass and the temperature sum and a positive relationship between fine root biomass and the carbon:nitrogen ratio of the soil organic layer. The foliage:fine root ratio varied between 2.1 and 6.4 for Norway spruce and between 0.8 and 2.2 for Scots pine. The ratio decreased for both Norway spruce and Scots pine from south to north, as well as from fertile to more infertile site types. The foliage:fine root ratio of Norway spruce was related to basal area and stem surface area. The strong positive correlations of these three parameters with fine root nitrogen concentration implies that more fine roots are needed to maintain a certain amount of foliage when nutrient availability is low. No significant relationships were found between stand parameters and fine root biomass at the stand level, but the relationships considerably improved when both fine root biomass and stand parameters were calculated for the mean tree in the stand. When the northern and southern sites were analyzed separately, fine root biomass per tree of both species was significantly correlated with basal area and stem surface area per tree. Basal area, stem surface area and stand density can be estimated accurately and easily. Thus, our results may have value in predicting fine root biomass at the tree and stand level in boreal Norway spruce and Scots pine forests.  相似文献   

7.
The effect of tree species mixture on stand volume yield and on tree-species-specific diameter and height growth rates were analysed in managed mixed stands of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Ehrn.).Data were obtained from 14 repeatedly measured stands located in Southern Finland on mineral soil sites with varying admixture of Scots pine and silver birch. Statistical analysis was carried out for studying the effect of species mixture on the development of stand characteristics. For the analysis, the plots were categorised into three groups (plot types) according to the species dominance. In order to analyse species-specific growth rates, individual-tree mixed linear growth models for tree diameter and height growth were developed for both tree species.The results clearly show that the yield of the managed mid-rotation, mixed stands was greater for stands dominated by Scots pine than for stands dominated by birch, and the stand volume increment decreased with an increasing proportion of silver birch. Analysis of diameter and height growth by tree species revealed that the main reason for this pattern is the negative impact of birch competition on the growth of pine trees. The increase in diameter of pine was clearly hampered if the proportion of birch was high. An abundance of birch also slightly decreased the growth in height of Scots pine, although the effect was less than on diameter growth. Species mixture did not affect the diameter growth of birch but did have a significant effect on height development. Height growth of birch was considerably greater in pine-dominated stands than in birch-dominated stands. In pine-dominated mixed stands, the height growth of birch was quite close to that of dominant pine trees, and birches can endure in competition with pines for light.The results apply for even-aged and single-storey managed stands, where stocking density and structure are controlled with pre-commercial and commercial thinnings. The results are not applicable to unmanaged mixed stands undergoing self-thinning. This study provides new information on mixed stands from a silvicultural perspective, which can be applied in decisions involving the management of mixed stands.  相似文献   

8.
油松—白桦混交林种间关系研究   总被引：4，自引：0，他引：4  
聂道平  沈国舫 《林业科学》1997,33(5):394-402
在调查河北隆化县油松－白烨混交林的林木生物量（地上部生物量及根系生物量）、林下草灌木、林地枯落物的总量以及不同混交距离油松地上地下部分生长发育状态的基础上，研究了混交林的种间关系。结果表明，该混交林作为华北地区广泛分布的一种林型，林下草灌木繁茂，林地枯落物分解迅速，有利于土壤理化性质和林分营养状况的改善，种间关系处于基本协调状态。由于林龄及密度原因，混交林生物量略低于油松纯林，在混交林中白桦对油松的生长有一定的抑制作用，抑制作用因混交距离而变化。本文提出了合适的混交距离和混交方式。  相似文献   

9.
Variability in short root morphology of the three main tree species of Europe's boreal forest (Norway spruce (Picea abies L. Karst.), Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth)) was investigated in four stands along a latitudinal gradient from northern Finland to southern Estonia. Silver birch and Scots pine were present in three stands and Norway spruce was present in all stands. For three fertile Norway spruce stands, fine root biomass and number of root tips per stand area or unit basal area were assessed from north to south. Principal component analysis indicated that short root morphology was significantly affected by tree species and site, which together explained 34.7% of the total variability. The range of variation in mean specific root area (SRA) was 51-74, 60-70 and 84-124 m(2) kg(-1) for Norway spruce, Scots pine and silver birch, respectively, and the corresponding ranges for specific root length were 37-47, 40-48 and 87-97 m g(-1). The range of variation in root tissue density of Norway spruce, Scots pine and silver birch was 113-182, 127-158 and 81-156 kg m(-3), respectively. Sensitivity of short root morphology to site conditions decreased in the order: Norway spruce > silver birch > Scots pine. Short root SRA increased with site fertility in all species. In Norway spruce, fine root biomass and number of root tips per m(2) decreased from north to south. The differences in morphological parameters among sites were significant but smaller than the site differences in fine root biomass and number of root tips.  相似文献   

10.
We determined fine root biomass and production of 15-, 35- and 100-year-old Scots pine (Pinus sylvestris L.) stands during three growing seasons. Fine roots were sampled by the soil core method. Mean (+/- SE) annual fine root biomass of Scots pine in the 15-, 35- and 100-year-old stands was 220 +/- 25, 357 +/- 21 and 259 +/- 26 g m(-2), respectively. Fine root biomass of the understory vegetation was 159 +/- 54 g m(-2), 244 +/- 30 and 408 +/- 81 g m(-2), and fine root necromass was 500 +/- 112, 1,047 +/- 452 and 1,895 +/- 607 g m(-2) in the sapling, pole stage and mature stands, respectively. Both understory and Scots pine fine root production increased with stand age. Mean annual Scots pine fine root production was 165 +/- 131, 775 +/- 339 and 860 +/- 348 g m(-2) year(-1) in the sapling, pole stage and mature stand, respectively. The respective mean annual production of all fine roots (Scots pine and understory) was 181 +/- 129, 1,039 +/- 497 and 1,360 +/- 869 g m(-2) year(-1). The Scots pine and understory fine root biomass, necromass and production varied in relation to stand age, although the variation was not statistically significant.  相似文献   

11.
In this study, the effect of pine mistletoe (Viscum album subsp. austriacum) on basal area increment of Crimean pine and Scots pine was investigated. Dendrochronological data were collected from 223 (71 uninfected and 152 infected) Crimean pines and 195 (77 uninfected and 118 infected) Scots pines located in Kastamonu province of Turkey in 2014. Infected sample trees were classified as light, moderate or severe infection levels. Growth trends and basal area increment loses were compared between uninfected and infected trees for the periods of the last 10, 20 and 30 years. In addition, infection status of forest stands was investigated using temporary sample plots; 27 plots in Crimean pine stands and 26 plots in Scots pine. Results demonstrated that basal area increments were negatively affected by pine mistletoe for both species. Mean basal area increment losses of infected trees for the last decade were determined as 24% for Scots pine and 26% for Crimean pine. Basal area increment losses varied by infection levels (light, moderate and severe) as follows: 25%, 20% and 28% for Scots pines and 20%, 32% and 9% for Crimean pines. Scots pine stands were more severely infected by pine mistletoe than Crimean pine stands. There were negative correlations between number of infected trees and stand density for both species, while positive correlation was detected between the number of infected trees and mean diameter for Scots pine. The results of this study indicate that the pine mistletoe infection has negative effect on radial growth of Scots pine and Crimean pine trees. The results can be an important contribution to the forest management and protection activities in mistletoe-infected stands.  相似文献   

12.
The objective of this study was to compare the survival and volume of conifer stands at 26 years of age with their status at planting. Survival, growth and damage were studied in eight clear felled stands regenerated in 1972. Five of the areas were planted with Norway spruce (Picea abies (L.) Karst.) and three with Scots pine (Pinus sylvestris L.). The plantings were examined in 1972 and 1974. In 1974, the number of living undamaged planted seedlings was low (10–15%). However, the number of undamaged seedlings was supplemented by naturally regenerated conifer and birch seedlings. The total number of undamaged seedling in 1974 was equivalent to 20–30% of the number of seedlings planted. In 1998, the main species in three stands had changed from Norway spruce to Scots pine, and in one stand from Norway spruce to birches. Actual volume in 1998 for the stands was compared to stand volume generate according to five scenarios based on recommended and actual seedling number in 1972 and 1974. The actual volume was 64% of that expected if the recommended number of trees had been planted. Naturally regenerated Scots pine and Norway spruce increased stand density in 1998. The actual volume was 37% higher than the average volume in the surrounding county. On average, 36% of the trees were damaged. More than 50% of the total damage was caused by moose (Alces alces L.). For Scots pine, moose or other browsing animals damaged 30% of the trees. The results of this study indicate that the 1998 volume was higher than expected, considering the low number of undamaged seedlings in 1974. This was mainly due to the large amount of naturally regenerated plants. In addition, the results indicate that the volume could have been higher if the initial conditions had been better. Despite the low number of undamaged seedlings in 1974, seven of the eight studied stands produced a higher volume than the average stand for the region. In practise, high numbers of seedlings should be planted on scarified areas. In most cases there will be a supply of naturally regenerated seedlings.  相似文献   

13.

New silvicultural regimes with high within-stand competition require new functions for estimation of standing stock and growth of biomass components, since the allometry of trees is changed by light competition. This paper presents functions for estimation of the aboveground biomass dry weights for stem wood, stem bark, branches and leaves of young (diameter at breast height <10 cm) Scots pine (Pinus sylvestris L.), Norway spruce [Picea abies (L.) Karst.] and birch (Betula pendula Roth. and Betula pubescens Ehrh.) trees growing in dense mixed stands. The functions were derived from a sample consisting of 84 Scots pine, 43 Norway spruce and 66 birch trees from six stands in northern Sweden with high stand densities (>10000 st ha-1). The logarithmically transformed power function displayed a good ability to stabilize the variance of dry weights and showed a good fit to the material (0.37< R 2 <0.99). A comparison with the most commonly used biomass functions in Sweden today showed that they overestimated the weight of stem wood and branches, while the weight of foliage was underestimated. The nature of these discrepancies suggested that the precision of biomass estimations might also be improved for young trees at wider spacing.  相似文献   

14.
G. abietina causes severe dieback in pole-stage stands of Scots pine in Britain. The susceptibility of ll provenances of Scots pine and of three other conifer species was tested by inoculations. Significant differences in susceptibility were found amongst Scots pine provenances, with those originating from a native Scottish pinewood at Loch Maree showing the highest levels of disease. Corsican pine was shown to be the most susceptible species tested although sporulation was greatest on Scots pine. Norway spruce was also shown to be susceptible when inoculated late in the growing season. Infection in lodgepole pine was negligible. High levels of β-phellandrene were tentatively linked with resistance among trees in a single stand of Scots pine.  相似文献   

15.
It was hypothesized that maximum living crown removal can be determined from experimental data using linear statistical models. Pruning experiments were carried out in four Scots pine (Pinus sylvestris L.) stands. Zero to nine living branch whorls were removed in three stands in southern Finland, and 0–4.5 m of the living crown was removed in a stand in northern Finland. Only dead branch whorls were removed from the control trees. A total of 517 trees were analyzed. Diameter increment was measured five years after pruning. The growth reduction was statistically significant when 40% or more of the living crown was removed by pruning. According to the linear models, green pruning decreased diameter growth by 0 to 33% at defoliation levels below 40%. This loss in diameter growth is, from an economical point of view, compensated by the knotfree timber resulting from pruning 2–3 years earlier.  相似文献   

16.
Scots pine (Pinus sylvestris L.) and European beech (Fagus sylvatica L.) dominate many of the European forest stands. Also, mixtures of European beech and Scots pine more or less occur over all European countries, but have been scarcely investigated. The area occupied by each species is of high relevance, especially for growth evaluation and comparison of different species in mixed and monospecific stands. Thus, we studied different methods to describe species proportions and their definition as proportion by area. 25 triplets consisting of mixed and monospecific stands were established across Europe ranging from Lithuania to Spain in northern to southern direction and from Bulgaria to Belgium in eastern to western direction. On stand level, the conclusive method for estimating the species proportion as a fraction of the stand area relates the observed density (tree number or basal area) to its potential. This stand-level estimation makes use of the potential from comparable neighboring monospecific stands or from maximum density lines derived from other data, e.g. forest inventories or permanent observations plots. At tree level, the fraction of the stand area occupied by a species can be derived from the proportions of their crown projection area or of their leaf area. The estimates of the potentials obtained from neighboring monospecific stands, especially in older stands, were poorer than those from the maximum density line depending on the Martonne aridity index. Therefore, the stand-level method in combination with the Martonne aridity index for potential densities can be highly recommended. The species’ proportions estimated with this method are best approximated by the proportions of the species’ leaf areas. In forest practice, the most commonly applied method is an ocular estimation of the proportions by crown projection area. Even though the proportions of pine were calculated here by measuring crown projection areas in the field, we found this method to underestimate the proportion by 25% compared to the stand-level approach.  相似文献   

17.
Stand Density Control Diagram (SDCD) is a stand-level mathematical model, which describes the relationships between yield, density and mortality throughout all stages of stand development. The SDCD is primarily used to derive density control schedules by management objectives. The main objectives of the present study are to define a modified model of SDCD for application to Scots pine (Pinus sylvestris L.) and Austrian black pine (Pinus nigra Arn.) plantations in Bulgaria, to examine the fitness of the model with representative experimental data sets from plantations of both species and to present a way of direct application of the SDCDs for practical purposes. The constructed SDCDs characterize the spatial-temporal dynamics of the pine plantations in a broad range of densities, forest sites and growth stages from 4 to 26 (28) m of dominant height class. The full density lines were fixed with self-thinning exponents α = 1.69 and α = 1.75 for Scots pine and Austrian black pine, respectively, and the trajectories of natural thinning for 23 initial densities (444-40000/ha) were determined. A direct way for application of the SDCDs to the plantation management was designed to estimate the optimal initial densities for the maximum attainable final yield and large-size wood production by self-thinning stands.  相似文献   

18.
The breeding of the pine shoot beetle (Tomicus piniperda) after thinning of young Scots pine stands in late autumn, and subsequent shoot damage, were studied in southern, central and northern Finland during three successive years (1977–79). No distinct increase in the attack density or beetle population was observed on the felled trees during the study period. Shoot damage on the standing trees remained low, and was not considered to have any marked negative effect on tree growth or further development of the pine stand. No noticeable difference was found between the different parts of the country. The economic importance of beetle damage resulting from late autumn thinnings was estimated to be low.  相似文献   

19.
Pensa M  Jalkanen R 《Tree physiology》2005,25(10):1265-1271
Latitudinal variation in needle longevity of conifers in response to climatic variability is a well-known phenomenon, but its significance has only rarely been studied. Scots pine (Pinus sylvestris L.) stands were investigated in four locations in Lapland (northern Finland and northern Sweden, 67-68 degrees N) and in four locations in Estonia (59 degrees N) to test the hypothesis that plasticity in needle longevity results in similar needle biomass per foliated shoot length across latitudes. The needle-trace method (NTM) revealed the dynamics of needle fascicles attached to stem shoots. Long-term mean needle age was 2 years greater in Lapland than in Estonia. In both regions, mean needle age was strongly and negatively related to yearly needle-fascicle production rate relative to the number of needles on the same shoot axis. Although significantly fewer needle-fascicles were produced annually in trees in Lapland than in Estonia, the overall number of fascicles attached to the stem shoots, needle-fascicle density and individual needle dry mass did not differ between regions. Consequently, needle biomass per foliated shoot length was similar in trees in both regions. Thus, our results support the theory that plasticity in needle longevity helps Scots pine to compensate for reduced needle production so that the same foliar biomass per shoot is retained under a wide range of growth conditions.  相似文献   

20.
Background: Bioenergy is re-shaping opportunities and imperatives of forest management. This study demonstrates,through a case study in Scots pine(Pinus sylvestris L.), how forest bioenergy policies affect stand management strategies.Methods: Optimization studies were examined for 15 Scots pine stands of different initial stand densities, site types, and temperature sum regions in Finland. Stand development was model ed using the Pipe Qual stand simulator coupled with the simulation-optimization tool Opti For Bioenergy to assess three forest bioenergy policies on energy wood harvest from early thinnings.Results: The optimal solutions maximizing bare land value indicate that conventional forest management regimes remain optimal for sparse stands. Energy harvests occurred only when profitable, led to lower financial returns. A forest bioenergy policy which included compulsory energy wood harvesting was optimal for denser stands. At a higher interest rate(4 %), increasing energy wood price postponed energy wood harvesting. In addition, our results show that early thinning somewhat reduced wood quality for stands in fertile sites. For less fertile sites, the changes were insignificant.Conclusions: A constraint of profitable energy wood harvest is not rational. It is optimal to carry out the first thinning with a flexible forest bioenergy policy depending on stand density.  相似文献   

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