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1.
Forest disturbance and transformations into agricultural land alter tropical landscapes at drastic rates. Here, we investigate bryophyte assemblages on trunk bases in natural forest, selectively logged forest and cacao agroforests that are shaded by remnants of natural forest in Central Sulawesi. Overall, bryophyte richness per site did not differ between forest types. However, mosses and liverworts reacted differently in that moss richness was lowest in cacao agroforests, whereas liverwort communities were equally rich in all forest types. In terms of cover, mosses remained unaffected while liverwort cover decreased significantly in disturbed forest. Species composition of bryophytes clearly changed in cacao agroforests as compared to natural forests and selectively logged forests. In particular some drought-sensitive species were rare or absent in cacao agroforests and were replaced by drought-tolerant ones, thus underlining the importance of microclimatic changes. Moreover, differences in bryophyte species composition between large and small trees were only pronounced in cacao agroforests, presumably due to concomitant changes in stemflow of precipitation water. In conclusion, the bryophyte assemblages of selectively logged forests and cacao agroforests were as rich as in natural forest, but species turn-over was particularly high towards cacao agroforests probably due to microclimatic changes. Maintenance of shade cover is crucial to the conservation of the drought-sensitive forest species.  相似文献   

2.
A modelling system (MS) for evaluation of current increment (net and gross growth) and mortality in Russian forests is presented. The informational basis of the MS are the data of the Russian forest state account (growing stock by dominant species, age, site indices, stocking) and some additional indicators (types of age structure, average species composition, type and intensity of disturbance regimes by ecoregions and/or landscapes. The MS includes model blocks generated with available information including yield tables (general, regional, full-stocked, with different stocking and types of age structure), growth tables, models, and experimental data for managed and unmanaged, disturbed and undisturbed forests. Two-dimensional (by age and site index) and three-dimensional (age, site index and stocking) models by species and ecoregions, based on regularities of coefficients of Mitscherlich growing functions, were used as the nuclei of the MS. The preliminary results, received for several Russian forest forming species gave a good fit to experimental data.  相似文献   

3.
Fragmentation of old-growth forests and greatly reduced amounts of coarse dead wood in managed forests threat the persistence of many saproxylic species in boreal Fennoscandia. Individual old-growth forest remnants may lose species over time as they pay off their extinction debt. We tested this by comparing the observed site occupancy of individual wood-inhabiting fungal species in isolated old-growth stands (i.e. woodland key habitats; WKHs) with statistical predictions of their occupancy assuming potential extinction debt had already been paid off. The occupancy of species was analysed in two sets of WKHs differing in time since isolation (i.e. recent and old isolates).Few species occurred more frequently than expected in WKHs. However, patterns across species and across all WKHs masked important differences among species in their risk of facing future extinction. The site occupancy decreased significantly between recent and old isolates for a group of annual, red-listed specialist fungal species, suggesting that an extinction debt in WKHs may exist among specific species confined to coarse dead wood and old-growth forest habitat. Generalist species that also occur in the surrounding matrix showed no negative trends, or actually increased in site occupancy, making future extinctions less likely. Thus, continuing loss of threatened species are likely if not preservation of WKHs are combined with other conservation efforts in managed forest landscapes. Natural forest landscapes may serve as important references when aiming to identify species in risk of future extinction but more detailed knowledge about the biology of the most vulnerable species is also required.  相似文献   

4.
5.
In European temperate forests, changes in the compositional similarity among local plant communities (beta diversity) have rarely been studied due to the lack of adequate baseline data. Several studies, however, report the spread of common, generalist species or a decline in specialized, rare species. Both processes may lead to increased similarity among communities, i.e., biotic homogenization.To quantify changes in beta diversity, we resampled the herb layer composition of ancient broadleaf forests at 175 semi-permanent plots distributed across the Weser-Elbe region in NW Germany 20 years after first sampling. We hypothesized that beta diversity would have decreased on average as a result of a spread of habitat generalists and a decline in habitat specialists.After two decades, the forest communities did not yet exhibit severe biotic impoverishment, although there was a broad trend towards homogenization. The actual magnitude of change depended on which beta diversity measure was applied. The downward trend was primarily the result of the spread of native species that are able to tolerate broad pH and moisture ranges. A distinction between forest specialists (closely tied to forest habitats) and generalists (also found in open habitats) did not help explain changes in beta diversity. The study shows that on the regional scale and in habitats not yet threatened by the invasion of alien species, shifts in native species can promote biotic homogenization.  相似文献   

6.
Riparian forests in the Sahel zone of Africa are of considerable importance for human resource use and for their ecological and conservation value. Typical riparian forest tree species are dependent on river flows and a shallow aquifer, and the community and population structure of riparian forests is related to spatial and temporal patterns of flooding at a site. Therefore, changes in hydrology caused by dams cause changes in floodplain vegetation. This paper explores the spatio-temporal complexity of these changes through an analysis of regeneration in a Nigerian floodplain affected by dams built in the early 1970s. The study shows that under certain conditions, and given sufficient time, riparian forests may be changing to forest types more characteristic of unflooded, upland areas. The results of the study have implications for the way in which the sustainability of developments downstream of large dams are analysed, suggesting that assessment of impacts must take place within temporal parameters relevant to ecological processes and at spatial scales that encompass spatial variation in floodplain ecosystems.  相似文献   

7.
The perceived health of forest ecosystems over large temporal and spatial scales can be strongly influenced by the frames of reference chosen to evaluate both forest condition and the functional integrity of sustaining forest processes. North American forests are diverse in range, species composition, past disturbance history, and current management practices. Therefore the implications of changes in environmental stress from atmospheric pollution and/or global climate change on health of these forests will vary widely across the landscape. Forest health surveys that focus on the average forest condition may do a credible job of representing the near-term trends in economic value while failing to detect fundamental changes in the processes by which these values are sustained over the longer term. Indications of increased levels of environmental stress on forest growth and nutrient cycles are currently apparent in several forest types in North America. Measurements of forest ecophysiological responses to air pollutants in integrated case studies with four forest types (southern pine, western pine, high elevation red spruce, and northeastern hardwoods) indicate that ambient levels of ozone and/or acidic deposition can alter basic processes of water, carbon, and nutrient allocation by forest trees. These changes then provide a mechanistic basis for pollutant stress to enhance a wider range of natural stresses that also affect and are affected by these resources. Future climatic changes may ameliorate (+ CO2) or axacerbate (+ temperature, + UV-B) these effects. Current projections of forest responses to global climate change do not consider important physiological changes induced by air pollutants that may amplify climatic stresses. These include reduced rooting mass, depth, and function, increased respiration, and reduced water use efficiency. Monitoring and understanding the relative roles of natural and anthropogenic stress in influencing future forest health will require programs that are structured to evaluate responses at appropriate frequencies across gradients in both forest resources and the stresses that influence them. Such programs must also be accompanied by supplemental process -oriented and pattern -oriented investigations that more thoroughly test cause and effect relationships among stresses and responses of both forests and the biogeochemical cycles that sustain them.  相似文献   

8.
The relative importance of time since disturbance and habitat variables in creating diversity in old-growth forests will influence conservation strategies. However, the independent roles of these factors are not well understood, as they are rarely examined independently. This study examines the respective roles of habitat variables and time (stand age) in determining bryophyte diversity in Picea mariana (Mill. (BSP)) forests. Bryophytes are frequently used as indicators of old-growth forest, but their true dependence on forest continuity is unknown. Bryophytes were classified into taxonomic-habitat guilds: true mosses (forest), forest liverworts, bog liverworts and sphagna (bog). Diversity increased with age and peaked at approximately 275 years since fire, driven by liverworts. Multiplicative habitat modeling indicated that time and habitat played different roles for the different taxonomic-habitat guilds. True mosses and forest liverworts were primarily influenced by habitat variables, while sphagna and bog liverworts were influenced by time and habitat variables. The models for sphagna were particularly strong, indicating that many important factors were included, while forest liverwort models were particularly weak. This unexplained variability may represent site specific random factors, such as secondary disturbances that create habitat in a fully occupied space. Overall, high richness was created by small species dependent on habitat variables and chance factors for establishment. Therefore, time since disturbance was not the primary factor limiting richness in these forests. These results suggest that in boreal North America, where forest fragmentation is limited and recent, conservation strategies that emphasize habitat variables rather than forest continuity may be effective for some bryophytes.  相似文献   

9.
北京市低山区水土保持林评价与分级   总被引:1,自引:0,他引:1  
以北京低山区同龄油松(Pinus tabulaeformis)林、侧柏(Platycladus orientalis)林、刺槐(Robinia pseudoacacia)林和栎类(Ouercus liaotungensis)林为研究对象,设置32个30m×30m标准样地,结合当地实际情况,运用数理统计方法及专家咨询法筛选出三级共12个评价指标,构建水土保持林评价指标体系,建立评价综合模型,计算林分综合评价指数,并对各林分进行了评价。将水土保持林分为四级:一级水土保持林,综合评价值S≥8;二级水土保持林,综合评价值5≤S<8;三级水土保持林,综合评价值2≤S<5;四级水土保持林,综合评价值S<2。探讨了不同级别的水土保持生长状况及立地特征,并提出了合理的改造措施。  相似文献   

10.
Range limits and broad-scale geographic variations in the productivity of boreal and northern deciduous tree species in Europe are simulated with a bioclimatic model (STASH). STASH is based on a small number of distinct mechanisms by which climate is thought to affect the survival, regeneration and growth of trees. Survival is limited by summer warmth requirements and winter cold tolerance; regeneration, additionally by winter chilling requirements; and growth rate by net assimilation, which in turn is related to photosynthetically active radiation, growing-season length, temperature (relative to species-specific optima) and soil moisture supply (relative to evaporative demand). These mechanisms are quantified either as thresholds (for survival and regeneration) or multipliers (for growth), based on bioclimatic variables computed from monthly climate normals interpolated three-dimensionally to a 10-minute grid. Growing-season and growing degree day calculations take into account the effects of chilling. The drought calculations also take into account the effects of soil-moisture storage by means of a physically based evapotranspiration calculation coupled to a one-layer soil hydrology model. STASH is used to examine changes in potential range limits under a 2xCO2 climate-change scenario. Associated transient responses at selected sites in the boreal and boreo-nemoral zones of Sweden are also simulated, using the forest gap model FORSKA2. The species-specific survival and regeneration constraints and growth responses of STASH modify the growth, establishment and mortality of trees in FORSKA2. The results obtained in this way differ sharply from the results of conventional forest gap models, where growth rates are assumed to decline to zero at minimum and maximum growing degree day limits. For example, towards the southern limit ofPicea abies (Norway spruce), STASH correctly shows no decline in productivity, but rather an abrupt cut-off corresponding to a chilling requirement during regeneration that is not met further south or west. In transient warming scenarios, this mechanism has the effect that natural regeneration can be blocked due to the warm winters even as yield is increasing due to the longer and warmer summers. STASH predicts drastic changes in species distributions in response to the large climate changes (especially winter warming) expected for northern Europe. Some of the common boreal species (e.g.,Picea abies; Pinus sylvestris; Alnus incana) are unlikely to survive in much of their present range, withdrawing to the far north. Other species already widespread may be able to occupy some of the few sites that are today unavailable to them (e.g.,Betula spp.;Corylus avellana). Other temperate deciduous species such asFagus sylvatica could have dramatic range expansions, potentially occupying large tracts of the present boreal zone. FORSKA2 transient simulations illustrate some of the possible routes towards different types of forest in a changed climate. Some sites in the north show little change in species composition, but sites towards the southern boundary of the boreal zone could develop a new suite of dominants. The degree of sensitivity of a particular site depends both on the climate change prediction and on the transient dynamics of the forest community. Many types of transient behaviour are shown to be possible. Coupled with uncertainties about the future role of dispersal and changes in disturbance rate, the complexity and variety of these transient responses imply a highly uncertain future for the north European boreal forests.  相似文献   

11.
Maintenance of old growth forest has been argued to be important for the future of many bryophyte species, but there has been no empirical test of this proposition. Bryophytes constitute much of the plant biodiversity in Tasmanian tall open-forests dominated by Eucalyptus regnans and Eucalyptus obliqua. These forests have a low level of protection in their old growth stage. We ask whether if there is a concentration of bryophyte species in particular forest age classes and assess its significance for biological conservation. The bryophyte species at 99 forest sites were recorded and related to young, mature and old growth age classes using ANOVA. Only two bryophyte taxa were confined, with statistical confidence, to one of these classes. Succession in bryophyte assemblages takes the relay floristics form, with a few highly frequent species in the young forest, most of which do not persist in later stages, a large number of species establishing in middle-aged forests, some dying out with their short-lived hosts by the time of the old growth forest stage, and liverworts becoming more prominent with age. Species concentrated in the old growth phase of these eucalypt forests have all been recorded from secure reserves in this and other vegetation types. If present patterns of land use continue, and no extensive wild fires occur in the reserved old growth forest, almost all bryophyte species of E. regnans and E. obliqua mixed forest are likely to survive. This may not be the case in other regions of the world, where reservation may be poorer than in Tasmania.  相似文献   

12.
人为干扰对高黎贡山社区森林树种多样性的影响   总被引:6,自引:0,他引:6  
研究人为干扰活动对高黎贡山社区森林树种多样性的影响结果表明 ,人为干扰对该社区森林有明显的负面影响 ,减少了群落树种多样性和破坏森林群落结构 ,降低森林资源量。当地居民对森林树种选择性的砍伐 ,使群落优势种组成发生变化 ,导致某些树种种群减小 ,物种生存受到极大威胁。并提出减少人为干扰的策略。  相似文献   

13.
Global conventions on biological diversity force governments to develop region-wide conservation strategies. Such strategies are difficult to design for all taxa because little is known about the important spatial scales. Here we applied additive partitioning of the diversity of saproxylic beetles in Bavarian forests in Southern Germany using a nested hierarchical design of five increasingly broader spatial levels: trap, strata, forest stand, forest site, and ecoregion. We consistently found a significantly higher percentage than expected by chance of between ecoregion diversity and significantly lower α diversity within traps. A significant proportion of β diversity was also found between stands. Analysis of species represented by <0.005% of all specimens in our samples and of species classified as threatened revealed similar results. Critical spatial scales for threatened species encompassed the critical levels of common species. Within habitat substratum guilds, the proportion of β diversity increased from species associated with fresh wood to those associated with rotten wood to those associated with fungi. Our results suggest that the most effective way to ensure saproxylic beetle diversity in a state-wide strategy is to add new conservation sites within different ecoregions and to establish new conservation areas in additional forest stands, rather than to enlarge reserves. Our findings further suggest that monitoring of saproxylic beetle diversity on a broader scale in European temperate forests can be restricted to “monitoring species”, i.e. a subgroup of families easy to identify, and that canopy sampling can be neglected without a substantial loss of information.  相似文献   

14.
This paper reviews the effects that windstorm-induced drastic changes (micro-climate, soil, vegetation, and ground structural heterogeneity) have on forest insect communities. In the current context of shady and CWD-deprived managed forests, windthrow gaps act as regional biodiversity hotspots by maintaining habitat continuity in a mosaic landscape, and by facilitating the breeding and population growth of clearing specialists and saproxylic species. Windthrow gaps are dead-wood islands where forest protection and habitat conservation goals may stand against each other. Besides the quantitative effect of dead wood on bark beetle outbreaks and saproxylic diversity, the latter is favoured by key dead-wood micro-habitats such as large logs, snags and sun-exposed coarse woody debris. The role of natural enemies and sanitation operations in regulating pest outbreaks is discussed. Heterogeneous openings provide many micro-habitats favouring flower-visiting insects, phytophages on saplings, on fallen tree crowns, and on diverse understory flora, as well as ground insects on specific micro-sites.  相似文献   

15.
金沙江干热河谷人工赤桉林群落结构   总被引:5,自引:2,他引:5       下载免费PDF全文
 为干热河谷地区的生态林建设提供理论依据,从反映群落稳定性的群落结构、物种多样性以及生长量和生物量等常用指标分析,金沙江干热河谷地区人工赤桉(Eucalyptuscamaldulensis)林密度普遍偏高,结构比较简单,群落稳定性较差。3种密度的赤桉林群落,随着林分密度的增大,林下植被层的群落分异性越大,林分中主要分布优势种(乔木为赤桉,林下植物为扭黄茅),其他种群的数量极少,林下植被层的结构比较简单,随着林分密度的增大,物种的丰富度、ShannonWiener指数、物种均匀度和生态优势度等4个指标不断减小,群落物种多样性在不断降低,结构合理性越差;群落生长量和生物量的变化,基本随赤桉林的密度增加而减小,密度最小的林分F1的生物量是其他2种高密度林分生物量的2倍以上。各林分林木生长缓慢,即使密度最小的林分F1的年平均胸径和树高生长量,分别只有0.72cm和0.78m。为了增加干热河谷地区人工乔木林群落的稳定性,使其逐步实现正向演替,发挥其应有的生态防护功能,应该考虑降低乔木层群落密度。  相似文献   

16.
We enhanced the forest patch model, Zelig, to explore the implications of 2×CO2 climate change scenarios on several forest regions in British Columbia and Alberta, Canada. In addition to the processes and phenomena commonly represented in individual-based models of forest stand dynamics, we added some species-specific phenology and site-specific frost events. The consideration of bud-break heat sum requirements, growing season limits, and chilling requirements for the induction of dormancy and cold hardiness slightly improved the ability of Zelig to predict the present composition of B.C. forests. Simulations of the predicted effects of future climatic regimes (based on the averaged predictions of four general circulation models) include some major shifts in equilibrial forest composition and productivity. Lowland temperate coastal forests are predicted to be severely stressed because indigenous species will no longer have their winter chilling requirements met. High-elevation coastal forests are expected to increase in productivity, while interior subalpine forests are expected to remain stable in productivity but will gradually be replaced by species currently characteristic of lower elevations. Dry, interior low-elevation forests in southern B.C. are likely to persist relatively unchanged, while wet interior forests are expected to support dramatic increases in yield, primarily by western hemlock. Northern interior sub-boreal forests are likewise expected to increase in productivity through enhanced growth of lodgepole pine. Conversely, the precipitous collapse of spruce stands in the true boreal forests of northeastern B.C. is expected to be associated with reduced productivity as they are replaced by pine species. Boreal-Cordilleran and Moist Boreal Mixedwood forests in Alberta are less likely to undergo compositional change, while becoming somewhat more productive. We believe these model enhancements to be a significant improvement over existing formulations, but the resulting predictions must still be viewed with caution. Model limitations include: (1) the current inability of climate models to predict future variation in monthly temperature and precipitation; (2) sparse information on the phenological behaviour of several important tree species; and (3) a poor understanding of the degree to which growth is constrained by different suboptimal climatic events.  相似文献   

17.
Investigations to determine stable or source-sink animal population dynamics are challenging and often infeasible for most species due to the time and expense of mark-recapture studies and the challenge of life histories attributes that result in low detectability and low recapture probabilities. Often, managers rely solely on occupancy or relative abundance patterns to assess a species’ sensitivity to environmental changes. Greater insight into population-level responses to environmental change can be gained by consideration of a combination of readily obtainable metrics, including occupancy, relative abundance, demographic structure and body condition. We examined how these metrics can improve our understanding of population-level responses to forest disturbance, using datasets for two exemplar species of terrestrial salamanders resident to the Pacific Northwestern USA. We compared population metrics for the Del Norte salamander (Plethodon elongatus) and the Siskiyou Mountains salamander (Plethodon stormi) across the seral continuum represented by four forest age classes: pre-canopy, young, mature, and old-growth. We compared these data with those collected from reference stands in mature (P. stormi) or old-growth (P. elongatus) forest containing robust populations. P. elongatus was twice as common as P. stormi. Both occupancy and salamander counts were lowest at pre-canopy sites for both species. Although there were numerous P. elongatus detections in young forests, higher proportions of these individuals were juveniles and sub-adults when compared to populations in late-seral forests. We found a negative relationship between the proportion of immature animals and total counts at a site, indicating that the high proportion of young animals in young forest stands is likely due to dispersal of young salamanders from nearby source populations and/or low survival of adult animals in young forests. We also found reduced body condition of P. stormi populations in young forests. Our results suggest that there are costs to populations occupying early seral forests, such as skewed age class structure and reduced body condition that are indicative of sink populations. Consideration of population-level metrics beyond occupancy and relative abundance can provide important insights when assessing a species’ sustainability in managed forest landscapes.  相似文献   

18.
In European forests, plants, fungi and invertebrates have been proposed as indicator species for assessing conservation value at the stand scale. To cover larger spatial scales, wide-ranging vertebrates could be added to that set of species. For resident forest birds, we (1) explored whether the occurrence of some species could be used to indicate high species richness and abundances, and (2) compared the results among four regions in the Baltic Sea drainage basin with a common species pool but different forest management intensities and varying proportions of deciduous and coniferous trees (south-central and southern Sweden, south-central Lithuania and northeastern Poland). Assemblages of deciduous forest birds in 100-ha landscape units were generally nested, suggesting that species richness within that group may be predicted based on the presence of a few species. Birds of coniferous forests, however, showed poorer conformity to nestedness in Sweden. Specialised species such as the middle spotted (Dendrocopos medius) and lesser spotted woodpeckers (D. minor) in deciduous forest and the three-toed woodpecker (Picoides tridactylus) in coniferous forest generally figured among the best indicators. In deciduous forest, there was high cross-regional consistency in the identity of the best indicators. Moreover, the sites where the best indicator was present also harboured higher relative abundances of most background species. For coniferous forest, however, such a relationship was not found. We conclude that an indicator species approach may be useful for resident birds of deciduous forests in hemiboreal Europe, emphasising that it should constitute one of many complementary tools for conservation management.  相似文献   

19.
Logging and intense shifting cultivation have caused major degradation of tropical forests and loss of biodiversity. Understanding the direct and indirect effects of those land uses on plant biodiversity is essential to the restoration of tropical forests. We compared the species diversity, community composition and basal area of all stems ?1 cm dbh among 18 1-ha tropical lowland and montane rain forest plots with a well-recorded long-term history of shifting cultivation and logging on Hainan Island, south China. We also explored the relative importance of disturbance and environmental factors in determining forest recovery. We found that the species density and diversity in old growth forests were higher than in shifting cultivation fallows (55 years old) but lower than in logged forests (35–40 years since logging). The species composition of shifting cultivation fallows was distinct from other forest types but logged forests were similar to old growth forests, especially in lowland forests. Disturbance intensity was the most frequently important factor in determining species composition, species density, diversity, and basal area accumulation. Soil nutrient availability explained some of the variation in species composition and diversity. Stem density was related to multiple factors including disturbance history, soil nutrients, and distance to old-growth forest. In general, we found that disturbance intensity was a better predictor of forest structure and diversity than edaphic environmental variables, highlighting the importance of human impacts in shaping tropical forest successional pathways.  相似文献   

20.
Higher densities of tree microhabitats in unmanaged forests may explain biodiversity differences with managed forests. To better understand the determinants of this potential biodiversity indicator, we studied the influence of tree characteristics on a set of tree microhabitats (e.g. cavities, cracks, bark features) on 75 plots in managed and unmanaged French forests. We hypothesized that the number of different microhabitat types per tree and the occurrence of a given microhabitat type on a tree would be higher in unmanaged than in managed forests, and that this difference could be linked to individual tree characteristics: diameter, vitality and species. We show that unmanaged forests contained more trees likely to host microhabitats (i.e. large trees, snags) at the stand level. However, at the tree level, forest management did not influence microhabitats; only tree characteristics did: large trees and snags contained more microhabitats. The number and occurrence of microhabitats also varied with tree species: oaks and beech generally hosted more microhabitats, but occurrence of certain types of microhabitats was higher on fir and spruce. We conclude that, even though microhabitats are not equally distributed between managed and unmanaged forests, two trees with similar characteristics in similar site conditions have the same number and probability of occurrence of microhabitats, whatever the management type. In order to preserve biodiversity, foresters could reproduce unmanaged forest features in managed forests through the conservation of specific tree types (e.g. veteran trees, snags). Tree microhabitats could also be more often targeted in sustainable forest management monitoring.  相似文献   

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