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1.
Elena Marcos Leonor Calvo José Antonio Marcos Ángela Taboada Reyes Tárrega 《European Journal of Forest Research》2010,129(1):25-30
We aimed to study tree effects on the chemical properties of forest soils. We compared soil features of three types of forest ecosystems, each with four stands (replicates): beech forests (Fagus sylvatica), oak forests (dominated by Quercus pyrenaica) and pine plantations (Pinus sylvestris). Five samples from the top 10 cm of soil were taken per stand, from which pH, organic matter content (O.M.), total nitrogen (N) and available calcium (Ca2+), magnesium (Mg2+), potassium (K+) and sodium (Na+) were determined. Litter layer depth was measured at each soil sampling point. We also measured tree density and crown diameters at each stand. Our results indicated that soil samples from the four pine plantation stands were more similar while oak and beech stands were characterised by great variability in terms of soil properties and leaf litter depth. Although the identity of the dominant tree species significantly influenced several topsoil chemical properties (increase in pH and available cations in oak forests and higher organic matter and total nitrogen in beech and pine ecosystems), there were other important factors affecting soil features that may be taken under consideration. Differences between soil properties of the three types of forest ecosystems were mainly related to the characteristics of the litter layer and less related to the tree layer structure. Finally, the establishment of pine plantations in naturally deciduous tree areas made the topsoil features more homogeneous. 相似文献
2.
《Forest Ecology and Management》2005,204(1):23-36
Spiders (Araneae) and ground beetles (Coleoptera, Carabidae) were studied in a woodland of the Northwest German lowland. An ancient oak–beech stand (170 years old) growing on mineral soil as well as a 110-year-old Scots pine (Pinus sylvestris) and a 55-year-old spruce (Picea spp.) forest growing on peat were investigated by pitfall trapping. A total of 155 species (39 carabids, 116 spiders) and 16,887 individuals (5269 carabids, 11,618 spiders) was recorded. Beetle diversity was high in the oak–beech stand and the spruce forest, but significantly lower in the pine forest. In both conifer plantations the activity density of carabids was considerably lower. Spider diversity was significantly higher in the spruce forest when compared to the beech and pine stand, respectively. Analyses of assemblage similarity distinguished clearly the fauna of all three stands. For each forest type, indicator species were detected. Although both conifer plantations were planted on former bogs, spider species typical of bogs were present only in the pine stand, not in the spruce stand. In both animal taxa, species typical of deciduous forests were more numerous and abundant in the oak–beech stand when compared to the conifer plantations. Although they were in direct contact, the conifer stands on peat only to a very low extent serve as secondary habitats for the epigeic fauna of the autochthonous deciduous woodland. During the 1990s, various agricultural programmes in Central Europe promoted such conifer plantations – in contrast, such afforestation measures on extensively used or fallow land of former bogs are not supported by the results of this study. 相似文献
3.
Reyes Tárrega Leonor Calvo Ángela Taboada Elena Marcos José Antonio Marcos 《European Journal of Forest Research》2011,130(6):949-957
We compared the structure of the arboreal layer and the diversity and species composition of the understory vegetation of
three types of mature forest communities: oak (Quercus pyrenaica) and beech (Fagus sylvatica) forests and Scots pine (Pinus sylvestris) plantations. Our main aim was to determine whether differences in these variables existed and were due to the identity of
the dominant tree species. We selected four stands or replicates per forest type located geographically close and with relatively
similar conditions. We found no differences in the arboreal structure of oak and beech forests, which were characterised by
great variability in tree size, while in case of plantations, this variability was lower at both the intra-stand (estimated
by the coefficient of variation) and inter-stand (i.e. the four replicates harboured trees of similar sizes) scales. However,
the highest variability in the canopy layer of natural forests was not consistently linked to greater understory species richness.
Indeed, the lowest plant species richness was found in beech forests, while oak forests harboured the highest value at either
the sampling unit (per m2) or stand scales. The greatest negative correlation between plant diversity and the environmental variables measured was
found for litter depth, which was the highest in beech forests. The results obtained by the CCA indicated that the four replicates
of each forest type clustered together, due to the presence of characteristic species. We concluded that pine plantations
did not approach the environmental conditions of native forests, as plantations were characterised by singular understory
species composition and low arboreal layer variability, compared to natural woodlands. 相似文献
4.
Carbon and nitrogen stocks and their medium-term and readily decomposable fractions in topsoils were compared in relation to soil microbial biomass and activity along sequences from coniferous to deciduous stands. The study was carried out in the Ore Mountains and the Saxonian lowland, representing two typical natural regions in Saxony, Germany. In accordance with current forest conversion practices, the investigation sites represent different stands: mature conifer stands of Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) (type A); Norway Scots spruce and pine with advanced plantings of European beech (Fagus sylvatica L.) or European beech/Common oak (Quercus petreae Liebl.) (type B); and mature deciduous stands of European beech and European beech/Common oak (type C). The investigated forest sites can be grouped into three silvicultural situations according to the development from coniferous stands to advanced plantings and finally mature deciduous forests (chronosequence A–B–C). The organic layer (L, F and H horizons) and uppermost mineral soil (0–10 cm) were analysed for potential C mineralisation, microbial biomass, concentrations of total C and N (TOC and TN) and for medium-term and readily decomposable C and N fractions, obtained by hot- and cold-water extraction respectively. The results showed an increase in organic layer thickness and mass as well as TOC and TN stocks along the forest sequences in the lowland. Yet, underplanted sites with two storeys revealed higher organic layer mass as well as TOC and TN stocks as compared to coniferous and deciduous stands. Stocks of hot- and cold-water-extractable C and N in relation to microbial biomass and its activity revealed a high turnover activity in deeper organic horizons of deciduous forests compared to coniferous stands. The stand-specific differentiation is discussed in relation to microbial biomass, litter quantity and quality and forest structure, but also with respect to the site-specific climatic factors and water budget as well as liming and fly-ash impacts. Results indicate higher dynamics in deciduous stands in the lowland especially during the initial turnover phase. The elevated microbial activity in deeper organic horizons of deciduous litter-influenced sites in spring is discussed as a specific indicator for long-term C sequestration potential as besides C mineralisation organic compounds are humified and thus, can be stored in the organic layer or in deeper soil horizons. Due to liming activities, stand-specific effects on organic matter turnover dynamics have evened out today in the Ore mountain region, but will presumably occur again once base saturation decreases. Here, the stand-specific effect on microbial biomass can currently be seen again as Cmic in the L horizon increased from spruce to beech. Our study sites in the lowland revealed no significant fly-ash impact. Differences between sites were evaluated by calculating the discriminance function. TOC and TN as well as medium-term degradable C and N were defined in this study as indicators for turnover dynamics along forest conversion sites. 相似文献
5.
Using the physiological single tree growth model BALANCE, vitality of forest stands was simulated in dependence of the site-related
factors, climate and stand structure. At six level II plots in southern Germany with the main tree species beech (Fagus sylvatica L.), oak (Quercus robur L.), spruce (Picea abies [L.] Karst.), and pine (Pinus sylvestris L.), simulated results were compared to measured values (soil water content, bud burst and leaf colouring, diameter at breast
height, tree height and crown density) in order to validate the model. Sensitivity tests were done to examine the influence
and the interactions of the environmental parameters. The validation results show that BALANCE is capable of realistically
simulating the growth and vitality of forest stands for central European regions for medium term time spans (several years).
The validation of the water balance module produces mean absolute errors based on field capacity between 2.7 and 6.9% in dependence
of sites and forest stands. Senescence of foliage as well as crown density is reproduced with a correlation coefficient of
0.70 compared to measurements. Differences between measured and simulated diameter values were smaller than 1% for spruce
and smaller than 6.5% for beech after 7 years of simulation, and smaller than 1% for oak after 8 years of simulation. On the
other hand, the simulations for pine trees conform less with the measurements (difference: 22.6% after 8 years). The sensitivity
of the model on environmental changes and on combinations of these parameters could be demonstrated. The responses of the
forest stands were quite different. 相似文献
6.
7.
Tzvetan Zlatanov Ivaylo Velichkov Manfred J. Lexer Tomislav Dubravac 《New Forests》2010,40(3):289-303
To study the regeneration dynamics in mature black pine (Pinus nigra Arn.) plantations in the Middle Balkan Range, Bulgaria, a total of 48 research plots (sized 0.1 ha) were established; 26
plots were established in the foothills of the mountains (foothill zone) and 22 in the main massive (mountainous zone). Pine
overstorey as well as advance regeneration in subplots were recorded. Analysis of variance and regression analysis showed
that in general relative stand basal area (RSBA) was the most important predictor of deciduous advance regeneration density,
top height as well as age span of the regeneration layer. In the mountainous zone soil depth (SD) and the presence of mature
deciduous trees (MDT) explained a significant amount of the variation in regeneration features. Lower RSBA induced by earlier
thinnings resulted in higher regeneration density, larger top heights and wider age span of deciduous species. Under the pine
canopy oak regeneration can persist longer compared to natural broadleaved canopies. In the studied plantations pine regeneration
was confined to few scattered seedlings and regeneration groups. Based on these findings silvicultural concepts for the further
management of black pine plantations in the Middle Balkan Range are discussed. 相似文献
8.
This paper summarises the results from 35 years-observed thinning experiments on 256 permanent sample plots in 10–60 year-old
stands of ash, aspen, birch, oak, pine and spruce in Lithuania. Thinning enhanced crown projection area increment of residual
trees. The largest effect was observed in stands of aspen and birch (growth increase by 200%), followed by ash and oak (over
100%), and spruce and pine (about 80%). Thinning also promoted dbh increment, especially in younger stands, and the increase
of dbh increment was positively correlated with the thinning intensity. The strongest reaction was exhibited by oak and aspen,
while ash, birch and conifers reacted to a lower extent. Low and moderate intensities of thinning stimulated volume production
in younger stands while the opposite was observed in older stands with increasing removals. Spruce stands exhibited relatively
strongest increase of volume increment and pine, –the weakest, while the effect on deciduous species was intermediate. The
results demonstrate that significant increase in volume increment is achievable with thinning of only young forest stands,
e.g. 10–20 year-old pine, birch and ash, or 10–30 year-old oak, aspen and spruce. 相似文献
9.
Adoración Martínez Juan Carlos Iturrondobeitia Arturo Goldarazena 《Annals of Forest Science》2009,66(3):304-304
10.
Heterobasidion annosum sensu stricto is the most important damaging agent in Scots pine stands planted on the former agricultural lands in Poland. The routine action in pine stands which have lost stability because of H. annosum root rot is to change stand management, including species conversion. In many cases, the Fagus sylvatica is used for this purpose. This study was the first assessment of widespread infection by H. annosum in young F. sylvatica plantations. Disease symptoms included atrophy and yellowing of leaves, wilting and the presence of pathogen sporocarps around the root collars of young trees. Heterobasidion annosum s. s. was observed on both 4‐ and 17‐year‐old beech. Based on annual increments, the disease could be present for 3–4 years before tree death. A high incidence of H. annosum in pine stumps of previous stands (80–100%) and dry periods in recent years may be the main reasons for such common infection of F. sylvatica. This work also showed that mice and frost were not the main killing factors F. sylvatica in plantations. 相似文献
11.
Growing concern about increasing concentrations of greenhouse gases in the atmosphere, and resulting global climate change, has spurred a growing demand for renewable energy. In this study, we hypothesized that a nurse tree crop may provide additional early yields of biomass for fuel, while in the longterm leading to deciduous stands that are believed to better meet the demands for other ecosystem services. Ten different species combinations were planted, with two different stocking densities, at three different sites in Denmark. Significant differences, with regard to biomass production, were observed among the different sites (P?P??1?yr?1 more biomass. The additional biomass production was similar to what was obtained in stands with conifers only (Sitka spruce, Douglas-fir and Japanese larch), which produced 4.9–6.1?t ha?1yr?1 more biomass than the pure beech stands. No effects of initial planting density (P?=?.19), or of initial weeding (P?=?.81), on biomass production were observed. Biomass production of the broadleaved crop was in most cases reduced due to competition. However, provided timely thinning of nurse trees, the qualitative development of the trees will allow for long-term timber production. 相似文献
12.
Beech woodland can be restored by direct planting of beech (Fagussylvatica L.) seedlings in abandoned areas, but this methodis generally avoided in forestry because of the growth difficultiesof beech in full-light conditions. This study tested a methodthat consists of planting beech seedlings in full-light conditionswith silver birch (Betula pendula Roth.) or Scots pine (Pinussylvestris L.) as nurse trees. A total of 65 two-year-old beechseedlings surrounded by either 3, 4, 5 or 6 pine or birch seedlingsor without competitors were planted in bare-soil open-fieldconditions in Central France. Tree growth and light availabilitywere monitored at the end of each of the following 3 years,and soil moisture was also measured the last year. At the endof the experiment, aerial biomass measurements were made onthe beeches. Results showed that relative beech growth in termsof diameter was significantly reduced by the local neighbouringtrees and that this reduction was particularly noticeable inthe pine treatments where there was extensive pine seedlingdevelopment. In contrast, relative beech height did not exhibitany significant variations among the treatments. Light availabilitywas decreased by the neighbouring trees, especially in the pinetreatments. Light reduction was more pronounced in the last2 years and in the middle or lower parts of the canopy of thesubject beech. Soil water content was lower under the pine canopiesthan under the birch canopies, and was positively correlatedto beech relative diameter growth. Specific leaf area, height-to-diameterratio and crown length-to-crown width ratio weakly but significantlyincreased with competition from the neighbours. Beech biomasspartitioning was only weakly affected by the treatments. Thisstudy showed that nurse trees tend to reduce beech growth butimprove form although effects remained weak due to the insufficientlength of the experiment. Further studies are also needed toquantify the effects of the neighbourhood on the growth of competitiveherbaceous vegetation and on changes in the microclimatic conditions. 相似文献
13.
David I. King Scott SchlossbergRobert T. Brooks Michael E. Akresh 《Forest Ecology and Management》2011,261(1):10-18
Fire-dependent ecosystems include some of the most threatened ecosystems in the world, and where fuels are allowed to accumulate, they can present significant threats to human life and property. Fuel reduction activities can be effective in reducing the risk of wildfire, but these practices need to be evaluated relative to their effect on biodiversity. We surveyed birds in an inland pitch pine-scrub oak barren, a fire-dependent plant community, in which fuel reduction had been carried out via thinning of canopy trees to reduce the risk of running crown fires. We hypothesized that thinning pitch pine forest would negatively affect the abundance of mature forest birds and positively affect the abundance of scrub-shrub birds. Our results confirmed these expectations: several mature forest bird species were less abundant in thinned pitch pine than unthinned pitch pine, although most of these species were also present in mixed deciduous forest, and therefore regionally well represented. In contrast, another group of bird species was scarce or absent from unthinned pitch pine and mixed deciduous forest, but present in thinned sites and scrub oak stands. These were scrub-shrub species that do not nest in mixed deciduous or pitch pine forest but depend on shrubland or savannah habitats that cover ∼3% of the region. We conclude that fuel reduction by thinning canopy trees at this site provides habitat for high-priority scrub-shrub bird species at the cost of modest reductions in numbers of forest birds whose regional aggregate population is large. 相似文献
14.
Data describing the growth, survival and stem form of: a) Europeancontinental, and b) British provenances of trees native to Britainwere collated from Forestry Commission records. The growth ratesof European continental provenances of Scots pine (Pinus sylvestris),silver birch (Betulo pendula), sessile oak (Quercus petraea)and common alder (Alnus glutinosa) were inferior to those ofBritish provenances in 90 per cent of cases. European provenancesof these species also showed inferior survival. The growth ratesof continental provenances of beech (Fagus sylvatica) were superiorto British provenances in about 50 per cent of cases, provenancesfrom northern France, Belgium and Holland being the fastestgrowing. Relationships between the relative height growth offoreign provenances and latitude were demonstrated for Scotspine, silver birch and beech (relative height growth = the heightgrowth of a foreign provenance expressed relative to that ofBritish provenances at the same site). The continental provenancesthat grew best originated at 0–4° south of the latitudesof the planting sites. Relative height growth decreased bothnorth and south of these latitudes. In the case of oak, relativeheight growth did not vary widely between origins. The survivalof continental provenances of Scots pine was particularly pooron exposed upland sites. The stem form of British provenancesof Scots pine was intermediate between Fennoscandian provenancesand provenances from the rest of Europe. In oak and beech therewas generally little difference in stem form between Britishand continental provenances, but the best continental provenances(Belgian beech; French oak) were most reliable. The growth andsurvival of native trees is interpreted in the light of thepost-glacial history of these species. The implications regardingthe continued use of seed of native hardwoods imported fromcontinental Europe are discussed. 相似文献
15.
Investigations on root and crown status of spruce and beech were carried out on selected trees in the ‘Werdenfelser Land’ area (Bavarian Alps, Southern Germany). In addition, the association of fine root pathogens of the genera Phytophthora and Pythium with the trees’ rhizosphere was studied. In a variety of stands representing various site conditions, soil and root samples were taken from 12 spruce (Picea abies) pairs and eight beech (Fagus sylvatica) pairs. Each pair consisted of a healthy and a declining tree as indicated by crown transparency. The root status was characterized using a set of parameters, and correlations between crown and fine root status were observed. In spruce, most parameters decreased significantly with increasing crown transparency, whereas in beech, correlations were less pronounced. The total number of lateral roots per cm small root (diameter 2–5 mm) was significantly lower in both species for declining trees compared to healthy trees. Pythium spp. were isolated from 15 of 24 soil samples taken from under spruce, and from eight of 16 samples from under beech. Phytophthora citricola was found in two beech stands only. Among the isolated species, Pythium anandrum, Pythium inflatum and Pythium acrogynum were identified according to morphological features. After polymerase chain reaction‐restriction fragment length polymorphism analysis, residual Pythium isolates were assigned to four different groups. No crown transparency‐dependent differences in isolation frequency were found. In soil infestation tests, all species tested caused root damage on both young spruce and beech plants, with P. citricola being the most aggressive pathogen. Additionally, Pythium‐infected beeches showed severe leaf chloroses and necroses. Due to their low isolation frequency, Phytophthora spp. are not considered to play a major role in the decline of spruce and beech in the investigated area. Pythium spp., however, were isolated frequently, showed pathogenicity towards the fine roots of spruce and beech, and are therefore considered to be at least contributing factors in the decline of Bavarian mountain forests. 相似文献
16.
T. Jung 《Forest Pathology》2009,39(2):73-94
During the past decade, and in particular after the wet year 2002 and the dry year 2003, an increasing number of trees and stands of European beech (Fagus sylvatica L.) in Bavaria were showing symptoms typical for Phytophthora diseases: increased transparency and crown dieback, small‐sized and often yellowish foliage, root and collar rot and aerial bleeding cankers up to stem heights of >20 m. Between 2003 and 2007 134 mature beech stands on a broad range of geological substrates were surveyed, and collar rot and aerial bleeding cankers were found in 116 (86.6%) stands. In most stands the majority of beech trees were declining and scattered or clustered mortality occurred. Bark and soil samples were taken from 314 trees in 112 stands, and 11 Phytophthora species were recovered from 253 trees (80.6%) in 104 stands (92.9%). The most frequent species were P. citricola, P. cambivora and P. cactorum. Primary Phytophthora lesions were soon infected by a series of secondary bark pathogens, including Nectria coccinea, and wood decay fungi. In addition, infected trees were often attacked by several bark and wood boring insects leading to rapid mortality. Bark necroses were examined for their probable age in order to determine whether the onset of the current Phytophthora epidemic was correlated to rainfall rates recorded at 22 Bavarian forest ecosystem monitoring stations. A small‐scale survey in nine Bavarian nurseries demonstrated regular infestations of all beech fields with the same range of Phytophthora species. The results indicate that (1) Phytophthora species are regularly associated with beech decline and may also be involved in the complex of ‘Beech Bark Disease’, (2) excessive rainfalls and droughts are triggering the disease, and (3) widespread Phytophthora infestations of nursery stock might endanger current and future silvicultural projects aiming on the replacement of non‐natural conifer stands by beech dominated mixed stands. 相似文献
17.
The effects of microhabitat (shrubs and herbs), plant litter, and seed burial on the regeneration of Liaodong oak (Quercus wutaishanica Mayr) and Chinese pine (Pinus tabulaeformis Carr.) were studied in three typical stands (Liaodong oak forest, Chinese pine plantation, and grassland) in the Loess Plateau, China. We monitored the establishment and growth of seedlings of these two woody species in sown experimental plots, in which shrubs and herbs, plant litter, and seed burial were manipulated. In the grassland, shrubs and herbs facilitated Liaodong oak establishment, with no effect on the establishment of Chinese pine. In the two forest stands, shrubs and herbs primarily had an inhibition effect on the establishment of these trees. The effects of plant litter were facilitation or inhibition, depending on the target species and the habitat. Seed burial had a positive effect on seedling establishment. In all three habitats, shrubs and herbs had inhibition effects on seedling growth of both tree species. Plant litter and seed burial did not influence seedling growth in either species. Liaodong oak and Chinese pine use different regeneration strategies during early stages of succession and similar strategies during late stages of succession. 相似文献
18.
《Forest Ecology and Management》1998,106(1):35-44
In East Germany, there are a lot of areas covered by old pine stands. They are growing on soils, on which under natural conditions without anthropogenic impact mixed stands of oak (Quercus petraea (Matt.) Liebl.) and pine (Pinus silvestris L.), would form a natural forest. An important objective of ecological silviculture in these areas is to convert the pure pine stands into mixed oak–pine stands by using natural regeneration methods. A highly appreciated assistant in this connection is the European jay (Garrulus glandarius L.). The remarkable results of its work have been found by analyzing the natural regeneration in a few old pine stands in the forest district of Weißwasser in Saxony. Although mother trees of oak are very scarce there, natural regeneration of oak was found everywhere in the research area. The oak regeneration, undoubtedly created by the jay, amounted to at least 2000 oaks/ha. They were mainly distributed at random, while pine regeneration was aggregated in places where gaps in the pine canopy occurred. Oak regeneration was established much earlier than pine regeneration. Thus, oaks exceeded pines in age, height and diameter. The oaks were also superior to pines concerning height increment for the last three years. Thus, there is a good chance for oak to defend its prevailing role in the regeneration in the future. Probably, the next forest generation will be composed of oak and pine trees. The analysis of the quality of the oak regeneration shows that there is no substantial difference to artificially sown oak stands. This indicates that the European jay creates oak stands sufficiently both in number and quality. 相似文献
19.
The ecological effects of planting exotic Douglas-fir [Pseudotsuga menziesii (Mirb.) Franco] in Central Europe are still poorly understood. The aim of this study was to answer the question of whether Douglas-fir affects tree specific arthropod communities in different mature forest types (Douglas-fir, spruce and beech dominated) in Southern Germany. Therefore, arthropod communities of stem and tree crown strata of Douglas-fir and spruce (Picea abies L.) were sampled in the years 1999–2001 using arboreal photo-eclectors and flight interception traps. Statistical analysis was conducted for all species and focused on conifer specialists at three levels: (1) species diversity, (2) guild structure and (3) community structure. Within the stem stratum, species diversity was significantly higher on spruce than on Douglas-fir independent of year and stand composition. This could not be explained by a single feeding guild, rather by species changing strata during the vegetation period. In contrast, species diversity in tree crowns was approximately the same for both conifer species. However, communities in Douglas-fir crowns were conspicuously different from those in spruce crowns, especially in the Douglas-fir dominated stand type. While zoophagous insects exhibited higher activity on Douglas-fir in 2000, xylophagous beetles were more abundant on spruce in 2001. In European beech stands with widely spaced Douglas-fir trees, the site specific and broad-leaved tree related fauna might be maintained. In addition, Douglas-fir with its resource of Adelges cooleyi and crowns that overtop the broad-leaved tree canopy, offer additional resources for several aphidophagous and thermophile species. 相似文献
20.
Jakub Borkowski Dorota Dobrowolska Wojciech Dąbrowski Rafał Banul Dariusz Załuski 《European Journal of Forest Research》2017,136(5-6):787-800
There is little knowledge how ungulate pressure on forest regeneration may be mitigated by silvicultural methods. The knowledge is especially needed for artificially regenerated, deciduous tree species. We studied factors affecting browsing incidence by deer in the Pisz Forest District in Poland, an area where 10,000 ha of forest was damaged by a 2002 hurricane. In 2006, we established three experimental plots (in total, 22.6 ha), in which the main species was Scots pine (Pinus sylvestris) admixed with pedunculate oak (Quercus robur). The data on browsing were collected in 2008–2015. In general, oak browsing incidence was unrelated to oak planting density. On a plantation scale, it was significantly affected by the pine age. Although in each variant all the oaks were browsed for four consecutive years (2009–2012), in 2013 browsing incidence began to decrease. When the pines grew higher and formed a physical barrier, it was harder for deer—roe deer (Capreolus capreolus), red deer (Cervus elaphus) and moose (Alces alces)—to move through and locate the oaks. Moreover, within plantations, oak browsing incidence was higher in the patches with shorter pines. Browsing of individual saplings or small groups of saplings was also negatively affected by the height of neighbouring pine saplings. Oak density influenced deer selectivity depending on the tree height. In a low oak tree density, browsing incidence was unrelated to oak height, while in higher tree density, deer selected oaks of the height between 40 and 100 cm. We postulate that deciduous admixture in a coniferous (unattractive) stand can be planted with a few year delay. Older coniferous trees should impede locating of attractive tree species by deer and the browsing incidence. 相似文献