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1.

In a trenching experiment the impact of root competition by overstorey Norway spruce on growth and allocation of above-ground woody biomass of advanced planted beech was investigated in two stands over two growing seasons after trenching. Measurements of soil water potential were taken inside and outside trenched areas by tensiometers at 16 measuring points in one stand from May to September 1999. Height and diameter of the seedlings, as well as branch length and diameter, were measured of beech growing in trenched and untrenched conditions in September before trenching (1998) and afterwards (1999 and 2000). Tensiometers inside the trenches reached the expiring value of -0.085 Mpa 30-40 days later, or never, compared with those in untrenched areas. Whereas relative growth rate (RGR) in height was not affected by trenching, RGR both in diameter and in estimated dry weight of the main stem were substantially enhanced in the first growing season after trenching (1999). A different result was obtained in 2000, when RGR in seedling diameter and estimated dry weight of the main stem was slightly reduced by trenching. The only variable that showed increased RGR of the trenched seedlings in both years was the estimated dry weight of the branches, indicating different above-ground biomass allocation patterns of seedlings with and without root competition.  相似文献   

2.
Otsamo  Riikka 《New Forests》1998,16(1):71-80
Effect of removing Acacia mangium overstorey on the growth of underplanted Anisoptera marginata saplings was studied in a mixed tree plantation established on an Imperata cylindrica dominated grassland site in South-Kalimantan, Indonesia. At the time of overstorey removal the Acacia mangium trees were 80 months old (average height 21 m and DBH 20 cm) and the underplanted Anisoptera marginata saplings 68 months old (average height 4.2 m and DBH 3.1~cm), and the spacing of both species was 2 m × 4 m with such an arrangement that the saplings were located within the same lines as the nurse trees. Eight plots of 192 m2 in size were randomly designated to overstorey removal (canopy gap) and no removal (closed canopy) treatments. Removal of Acacia mangium overstorey was carried out by girdling and later cutting down the dead trees. The Imperata grass did not show regrowth in canopy gaps. The mean annual increment for height and DBH of Anisoptera marginata within 22 months after the treatment were higher in canopy gaps (1.3 m a-1 and 1.4 cm a-1) than under closed canopy (0.7 m a-1 and 0.7 cm a-1). Results indicate good prospects for diversifying the composition of fast-growing forest plantations on grasslands and integrating slow-growing valuable species in plantation programs. Studies on other establishment methods, such as planting in gaps or lines, and further management practices as well as their effect on available light, moisture and nutrients are needed.  相似文献   

3.
Growth of regenerating trees in different light environments was studied for the mountainous, mixed-species forests in the Carpathian Mountains of Romania. The primary species in these mixtures were silver fir (Abies alba Mill.), European beech (Fagus sylvatica L.) and Norway spruce (Picea abies (L.) Karst). Seedlings/saplings of these species were selected and measured in different stands from two different geographical locations. Regenerating trees were measured for height and diameter growth during the summer of 2002. For each seedling/sapling, percentage of above canopy light (PACL) and stand basal area (BA) were used to assess available and occupied growing space respectively. Regeneration growth was compared against these two variables and regression relationships were developed. Using these models, we predicted the dynamics of regeneration as both growth and species composition. Our results showed that in low-light environments (PACL<20–35%; BA>30 m2/ha), shade tolerant fir and beech clearly outcompeted the spruce. Therefore, in dense stands, spruce could be eliminated by the shade tolerant species. For intermediate levels of cover (PACL=35–70%; BA=15–35 m2/ha) the spruce grew at comparable rates as the beech and fir. All three species showed similar growth rates in open conditions (PACL>80–90%; BA<15–20 m2/ha) with the spruce having a tendency to outgrow the others. However, in terms of establishment, such conditions favor spruce and inhibit fir and beech.  相似文献   

4.
The reaction of young beech (Fagus sylvatica L.) and Douglas fir (Pseudotsuga menziesii (Mirb.) Franco) saplings on competition of two types of vegetation—(1) gramineous with mainly Agrostis capillaries, Calamagrostis epigejos, Deschampsia flexuosa, and (2) small shrubs with mainly Rubus fruticosus and R. idaeus—on clear cuts on two sites was studied for 2 years. Half the sample saplings were released from competing vegetation by repeated herbicide applications. This treatment significantly raised the diameter increment in both species at the site with higher competition intensity, and more strongly after the removal of small shrubs than after the removal of grasses. Sapling length increment was not significantly affected. After being released from small shrubs, saplings of both species developed a smaller specific fine root length (cm g−1 fine root biomass) than unreleased saplings during the second year which was characterized by low rainfall. Root nitrogen concentration significantly increased after weed control in both vegetation types. Sapling foliar content of main nutritional elements was negatively related to dry mass and total chemical content of surrounding ground vegetation. Based on these results, a release from ground vegetation could be a useful tool to improve growth of planted beech and Douglas-fir saplings on sites with well-developed small shrubs competition (mainly by Rubus fruticosus and R. idaeus), or under fairly dry conditions.  相似文献   

5.
Morphology and vertical distribution patterns of spruce and beech live fine roots (diameter ≤2 mm) were studied using a soil core method in three comparable mature stands in the Solling: (1) pure beech, (2) pure spruce and (3) mixed spruce–beech. This study was aimed at determining the effects of interspecific competition on fine root structure and spatial fine root distribution of both species. A vertical stratification of beech and spruce fine root systems was found in the mixed stand due to a shift in beech fine roots from upper to lower soil layers. Moreover, compared to pure beech, a significantly higher specific root length (SRL, P<0.05) and specific surface area (SSA, P<0.05) were found for beech admixed with spruce (pure beech/mixed beech SRL 16.1–23.4 m g−1, SSA 286–367 cm2 g−1). Both indicate a flexible ‘foraging’ strategy of beech tending to increase soil exploitation and space sequestration efficiency in soil layers less occupied by competitors. Spruce, in contrast, followed a more conservative strategy keeping the shallow vertical rooting and the root morphology quite constant in both pure and mixed stands (pure spruce/mixed spruce SRL 9.6/7.7 m g−1, P>0.10; SSA 225/212 cm2 g−1, P>0.10). Symmetric competition belowground between mixed beech and spruce was observed since live fine roots of both species were under-represented compared to pure stand. However, the higher space sequestration efficiency suggests a higher competitive ability of beech belowground.  相似文献   

6.
The form and magnitude of storm damage and stand disclosure patterns were assessed in 332 randomly chosen pure and regular stands of spruce (Picea abies L.) and beech (Fagus sylvatica L.) after storm lothar, within a region of the Swiss Midlands. This data was analysed in relation to maximal wind speed, measured with Doppler radar techniques and other influential factors such as relief, allometric characteristics, silvicultural history, and neighbourhood. In addition, storm damage, assessed from aerial photographs over an extended perimeter (about 70,000 ha) was considered. A storm of the magnitude of lothar (December 26 1999), with an average maximal wind speed of 45 m s−1 (160 km h−1) appears to have a highly chaotic wind field structure, with great spatial and temporal variation of wind gusts. Wind speeds were not a significant predictor for damage in spruce stands and only weakly influential for beech. The consequences of this high randomness were analysed to estimate the return time of such a storm at the stand level. It lies between 86 and 113 years for spruce, 357 and 408 for beech. Only a few independent variables were significant and the overall explanatory strength of the model was unexpectedly low (R 2=0.07 for spruce and 0.30 for beech). Among the more reliable predisposing factors were mixture and aspect combined with gradient. An admixture of 10% or more broadleaved tree species or wind-firm conifers like Douglas fir [Pseudotsuga menziesii (Mirb.) Franco] significantly reduced the vulnerability of spruce stands (by a factor of more than three). On wind-exposed aspects, damage was more than twice the average. Steeper slopes caused a significant reduction in susceptibility (by a factor of six for slopes over 50%, in comparison to gentle slopes <20%). Other factors such as height to diameter ratio of trees or time since last thinning did not appear to be significant predictors.  相似文献   

7.
Direct seeding and planting of European beech (Fagus sylvaticaL.) are two common techniques for the conversion of pure Norwayspruce (Picea abies [L.] Karst.) stands into mixed stands. Thisstudy tested whether the growth of sown beech seedlings differsfrom that of planted seedlings of two stock types. Thereforeat two experimental sites repeated measurements were made ofheight, stem and first order branch diameter on sown and plantedseedlings over a period of nine years. The results showed thatthe growth responses of planted and sown seedlings to the environmentalconditions below the canopy of overstorey Norway spruce wererather similar. No differences between planted and sown seedlingswere found in their diameter and their estimated abovegrounddry weight. For all three batches the ratio between the estimatedbranch dry weight and the estimated main stem dry weight (branch-shoot-ratio(BSR)) was increased with age. Due to intraspecific competitionBSR was lowest for the sown seedlings irrespective of theirage. Differences in growth dynamics between planted and sownseedlings were found. For example, differences in the heightor the estimated aboveground woody dry weight between sown andplanted seedlings at a given age were not constant. Height,diameter and estimated dry mass of the seedlings in relationto age could be modelled precisely by a second-order polynomialfunction in the hitherto studied period.  相似文献   

8.
The three nonparametric k nearest neighbour (kNN) approaches, most similar neighbour inference (MSN), random forests (RF) and random forests based on conditional inference trees (CF) were compared for spatial predictions of standing timber volume with respect to tree species compositions and for predictions of stem number distributions over diameter classes. Various metrics derived from airborne laser scanning (ALS) data and the characteristics of tree species composition obtained from coarse stand level ground surveys were applied as auxiliary variables. Due to the results of iterative variable selections, only the ALS data proved to be a relevant predictor variable set. The three applied NN approaches were tested in terms of bias and root mean squared difference (RMSD) at the plot level and standard errors at the stand level. Spatial correlations were considered in the statistical models. While CF and MSN performed almost similarly well, large biases were observed for RF. The obtained results suggest that biases in the RF predictions were caused by inherent problems of the RF approach. Maps for Norway spruce and European beech timber volume were exemplarily created. The RMSD values of CF at the plot level for total volume and the species-specific volumes for European beech, Norway spruce, European silver fir and Douglas fir were 32.8, 80.5, 99.0, 137.0 and 261.1%. These RMSD values were smaller than the standard deviation, although Douglas fir volume did not belong to the actual response variables. All three non-parametric approaches were also capable of predicting diameter distributions. The standard errors of the nearest neighbour predictions on the stand level were generally smaller than the standard error of the sample plot inventory. In addition, the employed model-based approach allowed kNN predictions of means and standard errors for stands without sample plots.  相似文献   

9.
In a field study, we measured saplings of beech, ash and maple growing in a fairly even-aged mixed-species thicket established by natural regeneration beneath a patchy shelterwood canopy with 3–60% of above canopy radiation reaching the saplings. Under low light conditions, maple and ash showed a slight lead in recent annual length increment compared with beech. With increasing light, ash and maple constantly gained superiority in length increment, whereas beech approached an asymptotic value above 35% light. A suite of architectural and leaf morphological attributes indicated a more pronounced ability of beech to adapt to shade than ash and maple. Beech displayed its leaves along the entire tree height (with a concentration in the middle crown), yielding a higher live crown ratio than ash and maple. It allocated biomass preferentially to radial growth which resulted in low height to diameter ratios, and expressed marked plagiotropic growth in shade indicating a horizontal light-foraging strategy. In addition, beech exhibited the highest specific leaf area, a greater total leaf area per unit tree height, a slightly greater leaf area index, and a greater plasticity to light in total leaf area. Ash and maple presented a “gap species” growth strategy, characterized by a marked and constant response in growth rates to increasing light and an inability to strongly reduce their growth rates in deep shade. In shade, they showed some plasticity in displaying most of their leaf area at the top of the crown to minimize self-shading and to enhance light interception. Through this, particularly, maple developed an “umbrella” like crown. These species-specific responses may be used for controlling the development of mixed-species regeneration in shelterwood systems.  相似文献   

10.
Oriental beech (Fagus orientalis Lipsky.) and Oriental spruce [Picea orientalis (L.) Link] are the two most common tree species in northeast Turkey. Their distribution, stand type and understorey species are known to be influenced by topographical landforms. However, little information is available as to how these changes affect litter decomposition rates of these two species. Here, we investigated the effects of slope positions (top 1,800 m, middle 1,500 m and bottom 1,200 m), stand type (pure and mixed stands) and purple-flowered rhododendron (Rhododendron ponticum) on litter decomposition rates of Oriental beech and spruce for 4 years using the litterbag technique in the field. Among these three factors, stand type had the strongest influence on litter decomposition (P < 0.001, F = 58.8), followed by rhododendron (P < 0.001, F = 46.8) and slope position (P < 0.05, F = 11.6). Litter decomposition was highest under mixed beech/spruce forest, followed by pure beech and spruce forest. Beech and spruce litter decomposed much faster in mixed bags (beech–spruce) than they did separately under each stand type. Purple-flowered rhododendron significantly reduced litter decomposition of Oriental beech and spruce. Beech and spruce litter decomposed much slower at top slope position than at either bottom or middle position. Differential litter decomposition of Oriental beech and spruce was mainly due to adverse conditions in spruce forest and the presence of rhododendron on the ground which was associated with lower soil pH. Higher elevations (top slope position) slowed down litter decomposition by changing environmental conditions, most probably by decreasing temperature as also other factors are different (pH, precipitation) and no detailed investigations were made to differentiate these factors. The adverse conditions for litter decomposition in spruce forest can be effectively counteracted by admixture of beech to spruce monoculture and by using the clear-cutting method for controlling rhododendron.  相似文献   

11.
This research investigates how species in the sapling phase differ in growth and survival depending on light availability (as estimated by canopy position) by means of tree-ring analysis and modelling mortality. We harvested 120 live and 158 dead saplings in self-thinning communities consisting of Silver birch (Betula pendula Roth.), Scots pine (Pinus sylvestris L.), Japanese larch (Larix kaempferi Carr.) and Douglas fir (Pseudotsuga menziesii Mirb. Franco) in the Netherlands. Results are evaluated within the framework of a trade-off between high-light growth and low-growth survival. Radial growth, measured at ground level, generally declined over time. In addition, a decreasing light availability further reduced growth in all species except Douglas fir. Trees died when radial growth was reduced to about 0.5 mm year−1. Mortality in all species except Scots pine was significantly related to recent growth, but mortality curves were not different. The light-demanding Silver birch and Japanese larch differed from the shade-tolerant Douglas fir in both high-light growth and low-growth mortality, in line with a growth-survival trade-off. The light-demanding Scots pine did not fit this pattern as it was unable to transfer high radial growth into height gain, leaving it in suppressed canopy positions. This indicates the importance of height growth in the growth-survival trade-off. Differences in mortality probabilities affect the potential for coexistence, however, in all species also fast-growing individuals died suggesting additional factors causing mortality during self-thinning, other than direct competition for light.  相似文献   

12.
The aim of this study is to investigate the effects of forest conversion on forest floor vegetation. ‘Ecological’ forest conversion, as defined by an interdisciplinary southern Black Forest project group, describes the transformation of even-aged spruce (Picea abies L. H. Karst.) stands to structured continuous cover forests consisting of spruce (Picea abies), silver fir (Abies alba Mill.) and beech (Fagus sylvatica L.). In order to analyse the conversion process, four conversion stages were defined in a conceptual forest development model. Four forest districts deemed to be representative of the southern Black Forest region were selected for the study. The ground vegetation was initially classified independently from the stand structures. Subsequently, the relationship between stand structures, as determined by development stage, and ground vegetation was analysed. It was revealed that forest conversion modified the ground vegetation. The main factors influencing ground vegetation were the influence of broadleaves, predominately beech (F. sylvatica), on the canopy composition and litter coverage on the one hand; and the canopy coverage of spruce, the proportion of needle litter and the associated light penetration on the other. The prevalence of moss and vascular plant species preferring acidic sites found in spruce forests decreased during the transition process, whereas, species requiring a moderate base supply increased in abundance. The continuous cover forest representing the final stage of conversion increasingly contained a mixture of ground vegetation species normally associated exclusively with either conifer or deciduous forest. Due to the fact that there is an associated ground vegetation specific to the different stages of forest conversion in stands dominated by Norway spruce or European beech and a mixture in the latest conversion stages, large-scale forest conversion should be avoided in favour of management promoting a diversity of silvicultural goals and treatments in neighbouring stands. Only a variety of treatments ensures the maintenance of floristic diversity in the long-term.  相似文献   

13.
Establishing white spruce (Picea glauca (Moench) Voss) by planting it under established aspen (Populus tremuloides Michx.), stands has substantial potential as a technique for regenerating boreal mixedwood stands. The presence of an aspen overstory serves to ameliorate frost and winter injury problems and suppresses understory vegetation that may compete with white spruce. In this study we examine the growth of white spruce during the first 10 years after being planted underneath a 39-year-old stand of trembling aspen following thinning and fertilization. Results indicate successful establishment and reasonable growth rates of white spruce planted under thinned and unthinned aspen stands, even with aspen basal area of 51 m2 ha−1. Thinning of overstory aspen to 1000 or 2000 stems ha−1 did not increase light reaching seedlings, but did result in improvements in light above the shrub layer and in diameter and height growth of the underplanted seedlings. However, these increases in growth of underplanted spruce may not justify the expense of thinnings. Fertilization of these stands prior to planting had no effect on spruce growth. Growth of spruce underplanted at this site near Fort Nelson was similar to that at two other stands near Dawson Creek, B.C.  相似文献   

14.
Natural regeneration measurements are the main silvicultural objective in overaged protective forests of the Bavarian Limestone Alps. While manifold problems with these stands, especially the impact of browsing, are widely recognised, the regeneration niches of Norway spruce (Picea abies [L.] Karst.) are insufficiently known. The purpose of this study was to determine favourable combinations of site factors for the development of spruce in small, unfenced canopy gaps, located on Aposerido-Fagetum caricetosum albae forest sites. We recorded the occurrence of spruce saplings (as dependent variable) and of six site factors (as independent variables) on 480 0.5 m2-subplots. In addition, we estimated the coverage of six acid adapted plant species to determine correlations with the humus depth. A binary logistic regression analysis was used to predict the probability of the occurrence of a spruce sapling in dependency of the different site factors. Supported by other studies, we assumed that the supply of solar radiation was adequate for the sufficient regeneration of spruce within the canopy gaps. Other site factors significantly determined the regeneration niches of spruce saplings. More spruce saplings were found near hindrances and on rough surfaces than would be expected from a random occurrence of saplings. These microsite types may have characteristics, especially protection against snow gliding that promotes spruce establishment. A calculated “hindrance index”, which accounted for the number, and the distance of surrounding hindrances might be a good specific value to describe the influence of hindrances on steep slopes. The sapling establishment decreased on thin humus layers. Our assumption for the sites was that thick organic layers might represent a good seedbed for spruce. Decayed dead wood was scarce, but was exceedingly favoured by spruce saplings. Results obtained suggest that the natural regeneration establishment of spruce on steep slopes can be successfully influenced by site factors which inhibit the influence of snow gliding. According to a “positive microsite” concept, we recommend for artificial regeneration measurements with spruce, microsites close to hindrances (e.g. stumps, downed trees) and Vaccinium myrtillus as a predictor for thick, acid humus layers.  相似文献   

15.
On the regeneration and infection of roots of Douglas fir (Pseudotsuga menziesii [Mirb.] Franco) by fungi present in the soil. 167 roots in 4 stands were severed 40–60 cm from the base of the tree. At the proximal cut ends of the thick roots of the 40 years old trees on loamy sand, either replacement roots or callous tissue were formed (only a small percentage of those roots was invaded by rot causing Basidiomycetes), or no new roots and no callus were formed and root rot fungi (Fomes annosus, Coniophora puteana, Resinicium bicolor) had invaded the root sections from the soil. On the root sections of the 35 and 55 years old Douglas firs on sand, with the exception of 2 roots, neither replacement roots nor callus were formed; Fomes and Coniophora had invaded the thicker roots of the older trees from the soil.  相似文献   

16.
Investigations on root and crown status of spruce and beech were carried out on selected trees in the ‘Werdenfelser Land’ area (Bavarian Alps, Southern Germany). In addition, the association of fine root pathogens of the genera Phytophthora and Pythium with the trees’ rhizosphere was studied. In a variety of stands representing various site conditions, soil and root samples were taken from 12 spruce (Picea abies) pairs and eight beech (Fagus sylvatica) pairs. Each pair consisted of a healthy and a declining tree as indicated by crown transparency. The root status was characterized using a set of parameters, and correlations between crown and fine root status were observed. In spruce, most parameters decreased significantly with increasing crown transparency, whereas in beech, correlations were less pronounced. The total number of lateral roots per cm small root (diameter 2–5 mm) was significantly lower in both species for declining trees compared to healthy trees. Pythium spp. were isolated from 15 of 24 soil samples taken from under spruce, and from eight of 16 samples from under beech. Phytophthora citricola was found in two beech stands only. Among the isolated species, Pythium anandrum, Pythium inflatum and Pythium acrogynum were identified according to morphological features. After polymerase chain reaction‐restriction fragment length polymorphism analysis, residual Pythium isolates were assigned to four different groups. No crown transparency‐dependent differences in isolation frequency were found. In soil infestation tests, all species tested caused root damage on both young spruce and beech plants, with P. citricola being the most aggressive pathogen. Additionally, Pythium‐infected beeches showed severe leaf chloroses and necroses. Due to their low isolation frequency, Phytophthora spp. are not considered to play a major role in the decline of spruce and beech in the investigated area. Pythium spp., however, were isolated frequently, showed pathogenicity towards the fine roots of spruce and beech, and are therefore considered to be at least contributing factors in the decline of Bavarian mountain forests.  相似文献   

17.
An overview is presented of the silviculture of pedunculate oak (Quercus robur L.) and sessile oak (Q. petraea Liebl.) in Germany. This presentation is confined to less dry to moist and stagnic gleysol sites, where the intention is to produce primary timbers in long rotation periods. Incorporation of a shade tolerant species as an admixture species is indispensable to the suppression of epicormic branches. The most frequent and recommended admixture species for this purpose is beech (Fagus sylvatica L.). Because beech is competitively stronger than the oaks on these sites, silvicultural measures must be taken to keep it in check. According to the classic silviculture which has taken shape in the Spessart and Pfälzer forests, the desired goal of a two-storeyed stand with oak in the overstorey and beech in the understorey is achieved by means of heavy seeding beneath an open and rapidly cleared canopy. Because this procedure resembles clear cutting and entails its recognized disadvantages, trials have recently been undertaken to regenerate oaks in a silviculture with permanent canopy cover. Trials with young oaks show that they still achieve satisfactory growth at 15–20% of full light. Accordingly, successful regeneration is possible in beech stands under an open canopy or in gaps, and under an approximately closed canopy cover in pine stands due to their more penetrable crowns. The problematic aspect of this, especially in beech stands, is that young beeches become competitively stronger than oaks as canopy cover increases. This requires great effort in restraining the beech during cleanings and thinnings. Furthermore, browsing by wild animals must be minimized as it exclusively affects oak. As a result, however, there will be significantly fewer oaks and more beeches in the dominant layer than when using classic methods.  相似文献   

18.
Douglas fir (Pseudotsuga menziesii (Mirb.) Franco) was first introduced to Europe from North America more than 150 years ago, was then planted on a large scale and is now the economically most important exotic tree species in European forests. This literature review summarizes the current knowledge on the effects of Douglas fir on soil chemistry, plants, arthropods and fungi. Douglas fir shapes its abiotic environment similarly to native tree species such as Norway spruce, silver fir or European beech. In general, many organisms have been shown to be able to live together with Douglas fir and in some cases even benefit from its presence. Although the number of species of the ground vegetation and that of arthropod communities is similar to those of native conifer species, fungal diversity is reduced by Douglas fir. Special microclimatic conditions in the crown of Douglas fir can lead to reduced arthropod densities during winter with possible negative consequences for birds. The ecological impacts of Douglas fir are in general not as severe as those of other exotic tree species, e.g., Pinus spp. in South Africa and Ailanthus altissima, Prunus serotina and Robinia pseudoacacia in Europe. Nonetheless, Douglas fir can negatively impact single groups of organisms or species and is now regenerating itself naturally in Europe. Although Douglas fir has not been the subject of large-scale outbreaks of pests in Europe so far, the further introduction of exotic organisms associated with Douglas fir in its native range could be more problematic than the introduction of Douglas fir itself.  相似文献   

19.
Silvicultural treatments creating large canopy openings failed to restore regeneration of balsam fir (Abies balsamea (L.) Mill.) due to browsing pressure from white-tailed deer (Odocoileus virginianus Zimmermann). Consequently, we tested two alternative silvicultural treatments aimed at improving balsam fir establishment on Anticosti Island (Québec, Canada). In 1998 and 1999, we set up shelterwood seed cutting using three harvest intensities (0, 25 and 40% of basal area) and strip clearcutting with scarification using three different strip widths (15, 30 and 45 m), both with fenced and unfenced regeneration plots, in balsam fir stands. After 8 years, shelterwood seed cutting did not allow the establishment of new balsam fir seedlings, nor the development of unbrowsed balsam fir seedlings. In the strip clearcutting, deer browsing suppressed growth of palatable species in all strip widths. This favoured the development of unpalatable species, especially white spruce (Picea glauca (Moench) Voss). Our study demonstrates that the use of silvicultural treatments alone is unlikely to restore balsam fir regeneration on Anticosti Island, as long as the deer population remains higher than 20 deer/km2.  相似文献   

20.
In even-aged, 120-year-old Norway spruce stands with underplanted beech (in 1995) four permanent research plots were established, each 100 × 100 m in size. Twelve subplots were selected on each plot along a light gradient from complete canopy closure to open light conditions. On each plot, photon flux density (PFD) was measured continuously, and during same time interval (August 2008, 2009), color digital hemispherical photographs were taken. Cumulative PFD values for measuring points were compared with potential PFD radiation obtained from hemispherical photograph analysis for different angle of hemisphere and different parameters from hemispherical image analysis (gap fraction, total openness and direct, indirect and total amount of radiation). Cumulative and average daily values for the plots were compared; 120° hemispherical photograph angle, gap fraction and total openness were the variables that explained the largest proportion of variance in light transmittance. Determination coefficients between direct and total light component were highest for the total potential radiation and lowest for the diffuse light component. Comparison between potential and instantaneous light measurements for radial, height increment and SLA of young beech showed that instantaneous radiation measurements explained height increment best; 120° of hemisphere proved to be the best explaining angle.  相似文献   

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