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1.
Using a historical reconstruction, we discovered a potential trophic cascade involving cougar (Puma concolor), mule deer (Odocoileus hemionus), and California black oak (Quercus kelloggii) in the Yosemite Valley of Yosemite National Park in California. Our objective was to determine whether large deer populations in the absence of a top-level carnivore were suppressing tree regeneration. As human visitation increased in the early 1900s and cougar became increasingly scarce, the mule deer population irrupted in the 1920s. In August 2006, we undertook a retrospective study of black oak recruitment (i.e., growth of seedling/sprouts into tall saplings and trees) by inventorying all black oaks in stands accessible to deer and greater than 0.5 ha in size (18 stands, 2921 trees). We similarly inventoried oaks within sites representing refugia from deer browsing (4 stands, 481 trees). While significantly diminished oak recruitment has occurred since the 1920s in stands accessible to deer (p < 0.01), continuous recruitment of oaks was found in refugia sites. We also found less oak recruitment in areas of high human activity near the park’s visitor center (p < 0.01), possibly due to behaviorally-mediated effects of lower cougar and higher deer densities. Overall our results are consistent with trophic cascade theory involving large predators, herbivores, and plants. The long-term lack of oak recruitment is also an indicator of a probable loss of biodiversity.  相似文献   

2.
The appropriate management of forest reserves is debated; two major alternatives are succession to ‘wild’ state, or management to produce semi-open stands. For temperate conservation stands, there are no strong experiments replicated at landscape level. In each of 22 forests rich in oaks (Quercus spp.) in Sweden, we set up a closed-canopy wild plot (1 ha), and a cutting plot (1 ha) to produce semi-open conditions, studying them before and after cutting. About 25% of the tree basal area was cut (large trees and dead wood retained) and harvested as bio-fuel, a CO2 - neutral energy source. We examined the response of beetles and trapped 59,000 individuals (1174 species; 100 red-listed species). For both the guild of herbivorous beetles (222 species) and of saproxylic beetles connected to oak wood (267 species), species richness increased by about 35% in the harvested plots, relative to the wild reference plots. Species composition within the groups changed, though not strongly. Thirteen saproxylic species of 50 analysed, and three herbivores of 12 increased in cutting plots. For red-listed saproxylic beetles, species richness did not change significantly. Regression analyses suggest that more open cutting plots disfavour the red-listed beetles of this forest type. Thus, partial cutting increased species diversity of two beetle groups, probably due to changed microclimate and increase in herbaceous plants, but some red-listed saproxylic beetles may be disfavoured. A hands-off alternative may through storms and other disturbances produce open patches, more dead wood, and favour some species. Combinations of these alternatives, carefully planned at the landscape level, need to be considered.  相似文献   

3.
Dehesa ecosystems are open woodlands with scattered oak trees as their main component. As a result of differing land-uses, the structure of vegetation found within dehesas varies between: (i) oak trees and intercropped cereals (cropped), (ii) oak trees and native grass vegetation (grazed), and (iii) oak trees with abundant understorey shrubs (encroached). The aim of this study is to investigate whether land-use influences the water dynamics of dehesas by measuring available soil water content (AWC) in the upper 250 cm of the soil at different distances from tree trunks (maximum 30 m) at four Quercus ilex dehesas in Central–Western Spain. The technique used was Time Domain Reflectometry and the study was undertaken between May of 2002 and December of 2005. Leaf water potential (Ψ) was also measured on trees at one site by mean of a pressure chamber. Within the upper meter of the soil, it appears that trees, grasses and shrubs extracted soil water resources in a similar way from both beneath and beyond the tree canopy. However, encroached plots in general showed lower average AWC values than cropped or grazed plots (3.7, 5.6, and 6.2% in encroached, cropped and grazed, respectively). Cereal crops do not compete more strongly than grasses with trees for available soil water resources. The similar Ψ values found at cropped and grazed plots supported these results. From our results, it could be hypothesized that ploughed dehesas could facilitate soil re-watering in the plots with pronounced slopes. The decrease of AWC values at encroached plots with respect to the cropped and grazed plots was found mostly beyond the tree trunk at deeper soil layers, indicating that shrubs use water partly not accessible to trees. The presence of an understory of shrubs seems to have slightly increased the water constraints on trees during the summer period (Ψd values of − 0.5, − 0.5, and − 0.8 MPa in cropped, grazed, and encroached plots, respectively). In cropped and grazed plots, an important amount of water seems to have remained unused for trees and grasses.  相似文献   

4.
This three year study was conducted to evaluate the suitability of municipal solid waste (MSW) compost as a soil amendment or a mulch to suppress weeds in the production of field grown nursery stock. In the soil amendment experiment, compost was applied at the rates of 0, 56, and 112 t/ha and incorporated into the soil prior to planting. In the mulch/weed control experiment, 224 t/ha of compost was applied to one set of plots as a mulch after planting and compared to plots treated with herbicide and untreated controls. Each plot in both experiments contained four rows with one each of red maple (Acer rubrum L.), sugar maple (Acer saccharum Marsh.), white pine (Pinus strobus L.), and pin oak (Quercus palustris Muenchh.). Mortality and weed counts were determined during the first growing season. Annual measurements of crown height, crown width, and caliper (at the 1 foot level) were made the following three winters. Two inches of compost mulch (224 t/ha) provided adequate weed control only in the first year. Mowing suppressed the weeds that invaded the plots beyond the first year. For maples, first year mortality in plots mulched with 224 t/ha compost or amended with 112 t/ha compost was significantly less than unamended controls. In subsequent years, the mulched maples grew more than trees in the unamended controls. In the plots amended with 56 or 112 t/ha compost, sugar maples had increased height, canopy, and caliper growth compared to the unamended controls. The hearty pin oak saplings survived transplanting and there was a significant increase in caliper growth in plots amended with 112 t/ha (1 inch) of compost. Mortality of white pine increased when mulched with 224 t/ha compost. Increased mortality was most likely due to high concentrations of ammonium and soluble salts in the immature compost. Beyond the first year, there were no negative impacts on the growth of white pine from any of the compost treatments.  相似文献   

5.
This study describes the changes in height and diameter in Quercus ilex trees grown on agricultural land in a semi-arid Mediterranean climate, in response to different soil management practices: mechanical weeding (MW), chemical weeding (CHW) and no treatment (NT); and compares trees grown from 1-year-old saplings and from acorns. After 6 years, trees that had been mechanically weeded were largest; for trees grown from saplings the average height was 120 cm and diameter was 3.83 cm, for trees grown from acorns average height was 100 cm and diameter was 2.56 cm. Soil organic matter increased in all managed plots while phosphorus, nitrate, and water content oscillated between years. The no treatment plot had the highest values for nitrate and organic matter while the chemical weeding plot had the highest level of phosphorus. Annual mean on C/N ratios were higher in the mechanical weeding (12.6 ± 3.3) followed by the chemical weeding (12.0 ± 1.6). Also mechanical weeding treatment showed the lowest soil water content throughout the study. An old tillage-induced compaction zone appeared in all plots at a depth of 12.5–17.5 cm with mechanical weeding causing the greatest compaction. However, no relationships were found with tree heights, neither with spatial compaction zoning. All trees showed the same growth pattern, whether raised from saplings or acorns and irrespective of soil management system. The height and diameter of trees were linked to the sum of height and diameter ratios from previous years. Our study shows that soil management does influence tree growth rate. Mechanical weeding should be considered for afforestation of agricultural lands with Q. ilex in semiarid Mediterranean lands.  相似文献   

6.
Polypore communities were compared between mature managed, overmature managed, and old-growth spruce-dominated forests in southern Finland. A total of 85 polypore species, with 6000 records, were found in 16 sample plots, each 4 ha in size. Old-growth stands had on average 80% more species than mature stands, and 38% more species than overmature managed stands. Variation in polypore species richness was best explained by diversity of dead wood and the number and volume of dead trees. The best predictor for the number of threatened polypore species was the number of cut stumps. Threatened species were practically confined to old-growth forests and to stands in which the amount of dead wood exceeded 20 m3/ha. This figure appears to represent a stand-level threshold value for the amount of dead wood, below which the persistence of threatened species becomes unlikely. Our results suggest that a significant increase in the amount of dead wood (e.g. by leaving large retention trees and even by killing trees) is needed in managed forests before they become suitable habitats for threatened polypores.  相似文献   

7.
The appropriateness of using seedling responses to predict mature tree responses to ozone is questionable considering the long life span of trees. To determine if northern red oak (Quercus rubra L.) seedlings and mature trees respond similarly to ozone, 30-year-old trees and two-year-old seedlings were fumigated with subambient, ambient, and twice ambient concentrations of ozone for three growing seasons using an open-top chamber facility. Ozone reduced leaf physiological function of mature trees. However, stem diameter growth and leaf mass of mature trees were not significantly reduced by ozone. Photosynthesis in seedling leaves was slightly reduced by ozone, but seedling growth and biomass were unaffected. Mature tree fine-root biomass responded negatively to increasing ozone, while seedlings exhibited increases in biomass. Carbon retention was enhanced in leaves and branches of mature trees by ozone treatment, while seedlings were unresponsive. Differences in ozone uptake, recurrent flushing and carbon dynamics rather than microclimate may explain differences in ozone sensitivity between seedlings and mature trees. These results indicate that seedlings may not reflect the ozone sensitivity of larger more physiologically mature trees.  相似文献   

8.
Many soils of the semi-arid tropics are deficient in P, and under such adverse conditions, the establishment of pearl millet seedlings is a critical step to achieve satisfactory crop stands. Phosphorus fertilizer is expensive for small holder farmers and is only applied at low rates, insufficient to give satisfactory crop stands. Methods are needed to enhance productivity at low rates of application. Here, we tested the hypothesis that a minute application of P at early seedling stage, equivalent to 125–500 g P/ha, would enhance the plant establishment under P-limiting conditions. We measured the minimum application of P needed to elicit a response of different genotypes. Pot experiments were conducted with pearl millet ( Pennisetum glaucum (L.) R. Br.) hybrids to measure the response to P placed close to the root system, 5 days after sowing (DAS), compared with a non-limiting P control (DAP). The placement treatments were 0, 0.25, 0.50, and 1.0 mg soluble P per seedling, applied as KH2PO4 solution. The localized placement of P increased biomass in all three soils tested. In one soil, plant biomass at 1000 μL (1 mg P) reached about 50% of the non limited control. If applied later than 19 DAS the placement had no effect on the plant biomass. Hybrid ICMP 451-P8 was more responsive than hybrid 81B-P6. Placement to 20 inbred lines of pearl millet increased biomass by an average of 105% compared with no placement with large genotypic variation. Although this work was not intended to be a way of applying P fertilizer to pearl millet under field conditions, it showed that applying minute amount of P to pearl millet seedling (equivalent to 125–500 g P/ha) enhanced their establishment and led to improved growth for at least 5–6 weeks after sowing. Further work is in progress to develop a feasible technology for field crops based on the results of this study.  相似文献   

9.
Bryophytes growing on siliceous boulders (diameter 50-200 cm) were studied in two forests in east-central Sweden to investigate the influence of different canopy tree species on the bryophyte species richness. Granite boulders lying below crowns of Ulmus glabra (elm), Fraxinus excelsior (ash), Acer platanoides (maple), Quercus robur (oak), Betula pendula (birch), and Picea abies (Norway spruce) were studied in two mixed stands. Both boulder area and within-boulder habitat diversity were positively related to species richness. Boulders below the base-rich deciduous trees F. excelsior, U. glabra and A. platanoides held roughly twice as many species as those below P. abies, with Q. robur and B. pendula as intermediates. The rank order among trees was as expected from bark pH and litter decomposability in the literature. We also used sample plots to investigate species richness at a smaller scale. Within plots, intermediate levels of litter, inclination and exposed rock promoted species richness, and also the covering tree species had an effect. Individual plots were on average not more species-rich on large boulders than on small ones. This leads us to conclude that population extinctions on the plot-level, and re-colonisations from other parts of the boulder, are less important for species richness than within-boulder habitat diversity. Since species richness was lower under P. abies it is important to favour a diversity of tree species and to include base-rich deciduous trees in otherwise homogeneous Picea forests.  相似文献   

10.
Enrichment planting with climax trees such as dipterocarps is required to rehabilitate degraded tropical forests in Southeast Asia, because these forests have been damaged by various anthropogenic activities. Severely degraded tropical secondary forests are the most important targets for enrichment planting in the region, because secondary forest trees potentially provide moderate conditions for planted seedlings by preventing strong sunlight and high temperatures. In an enrichment planting site located in Sarawak, Malaysia, we explored how variation in secondary forest structure (biomass and stand density) and soil (nutrient and water) conditions influenced the mortality and growth rates of planted dipterocarp seedlings (Parashorea macrophylla, Wyatt-Sm. ex P.S. Ashton) over the period from 2000–2007. Experimental plots were set up on different topographical features (slopes and valleys). Using the data collected, we demonstrated the ways in which environmental factors, such as soil nitrogen content, relate to climax tree seedling growth and mortality in degraded tropical secondary forests dominated by the genera Macaranga, Ficus, and Glochidion. Stand density and total aboveground biomass of secondary forest trees were significantly lower in the valley plots than on the slopes, whereas soil total nitrogen and water contents were significantly higher in valley plots. Over seven years, the total biomass of trees increased, whereas the density of secondary forest trees decreased in all plots. Nutrient stocks (nitrogen, phosphorus, potassium, calcium, and magnesium) in the soil (0–5?cm) also decreased over seven years in all plots. Height, diameter, and mortality of planted seedlings were higher in the valley plots than on slopes. Multiple stepwise regression demonstrated significant positive effects of total soil nitrogen and water content on the relative growth rate of seedlings over seven years. There were no significant relationships between seedling mortality and any of the environmental factors measured, including soil conditions. We conclude that (1) seedling growth was enhanced by soil total nitrogen rather than by available phosphorus or potassium, and (2) planted seedlings grew faster and died more often in the valley plots than on the slopes.  相似文献   

11.
The flood regime in the middle course of the Ural River has been studied on the basis of data obtained at hydrological stations from 1912 to 1990. Its effect on the growth and productivity of oak stands on different elements of the floodplain is discussed. The particle-size distribution and the reserves of physical clay, clay, humus, and available potassium and phosphorus under natural (three plots) and artificially planted (three plots) oak stands have been determined. It is proved that natural oak stands are allocated to levees with considerable content of physical clay (<0.01 mm) in the soil profile, which provides penetration of the tree roots to a depth of 6 m and more, down to the ground-water level during the summer low-water period. The role of water infiltration into the banks in the periods of low floods is shown to affect the ground-water level at a distance of up to 700 m from the river bed. The artificial cultivation of oak stands on the floodplain requires the proper selection of particular ecological niches. The successful growth of artificially planted oak stands is only possible near the lakes within the central floodplain. Problems of the expansion of oak stands onto the high elements of the central floodplain, the replacement of oak by other tree species, the natural regrowth of oak trees, and the shift of their southern boundary onto the floodplain in the lower course of the Ural River are discussed.  相似文献   

12.
蝴蝶兰种苗自动化切割可降低组培苗染病几率,提高种苗品质。为实现蝴蝶兰种苗自动化切割,该研究针对种苗结构特性提出基于拟合直线的切点定位方法,并配合切割方法设计了弹性切割末端,搭建了基于视觉伺服的蝴蝶兰种苗切割系统。首先,采用深度学习模型对采集到的图像进行目标检测;然后,根据检测结果使用基于几何规则的切割点定位算法计算切点;最后,将切割坐标传输给切割执行机构完成切割作业。目标检测试验中,ShuffleNet v2-YOLOv5模型检测精度达96.7%,权重文件大小1.3 MB,平均检测时间0.026 s。种苗切割试验中,切割合格率高于86%,单株平均切割时间小于18 s。该系统能有效完成蝴蝶兰种苗切割任务,为蝴蝶兰组培苗自动化生产提供新思路。  相似文献   

13.
Many bird species respond to forestry, even at moderate intensities. In New Brunswick, Canada, the Brown Creeper exhibits a negative, threshold response to harvesting intensity. This study aimed to determine whether (a) the threshold found in Brown Creeper occurrence is lower than eventual thresholds in its nesting requirements, and whether (b) the conservation of this species could be achieved through moderate-intensity harvest systems. Creepers are particularly sensitive to forestry because they nest on snags with peeling bark and they mainly forage on large-diameter trees. In northern hardwood stands, we compared habitat structure at local- (r = 80 m) and neighbourhood-scales (r = 250 m) around nest sites and sites not used by creepers. Over two years, we found 76 nests, 66 of which were paired with unused sites for comparison. At the local scale, densities of trees 30 cm dbh and snags 10 cm dbh, and the probability of presence of potential nest sites were significantly higher near nests than at sites where no creepers were detected. At the neighbourhood scale, the area of untreated mature forest was significantly higher around nests. Variance decomposition indicated that habitat variables at the local scale accounted for the majority of explained variation in nest site selection. We also found significant thresholds in the densities of large trees (127/ha) and snags (56/ha), and in the area of mature forest (10.4 ha). The conservation of breeding populations of Brown Creepers may thus require densities of large trees nearly twice as high as those associated with its probability of presence. Such a target seems to be incompatible even with moderate-intensity harvesting.  相似文献   

14.
The negative impacts of soil compaction on crop yields can often be alleviated by subsoiling. However, this subsoiling operation is often conducted at unnecessarily deep depths wasting energy and excessively disturbing surface residue necessary for erosion control and improved soil quality. A corn (Zea mays L.)–cotton (Gossypium hirsutum L.) rotation experiment was conducted over 4 years on a Coastal Plain soil with a hardpan in east-central Alabama to evaluate the potential for site-specific subsoiling (tilling just deep enough to eliminate the hardpan layer) to improve crop yields while conserving energy. Seed cotton yield showed benefits of subsoiling (2342 kg/ha) compared to the no-subsoiling treatment (2059 kg/ha). Averaging over all years of the study, site-specific subsoiling produced cotton yields (2274 kg/ha) statistically equivalent to uniform deep subsoiling at a 45 cm depth (2410 kg/ha) while not excessively disturbing surface soil and residues. Significant reductions in draft force were found for site-specific subsoiling (59% and 35%) as compared to uniform deep subsoiling at a 45 cm depth in shallow depth hardpan plots (25 cm) and medium depth hardpan plots (35 cm), respectively. Calculated fuel use for site-specific subsoiling was found to be reduced by 43% and 27% in the shallow and medium depth hardpan plots, respectively, as compared to uniform deep subsoiling in these same plots. Producers in the Coastal Plains who can determine (or who know) the depth of their root-impeding layer and perform site-specific subsoiling can have comparable cotton yields to traditional uniform depth subsoiling with reduced energy requirements.  相似文献   

15.
The objective of this work was to evaluate the C and N stocks and organic‐C fractions in soil under mixed forest stands of Scots pine (Pinus sylvestris L.) and Sessile oak (Quercus petraea [Matt.] Liebl.) of different ages in NE Germany. Treatments consisted of pure pine (age 102 y), and pine (age 90–102 y) mixed with 10‐, 35‐, 106‐, and 124‐y‐old oak trees. After sampling O layers, soils in the mineral layer were taken at two different depths (0–10 and 10–20 cm). Oak admixture did not affect total organic‐C (TOC) and N stocks considering the different layers separately. However, when the sum of TOC stocks in the organic and mineral layers was considered, TOC stocks decreased with increasing in oak age (r2 = 0.58, p < 0.10). The microbial C (CMB) was not directly correlated with increase of oak age, however, it was positively related with presence of oak species. There was an increase in the percentage of the CMB‐to‐TOC ratio with increase of oak‐tree ages. On average, light‐fraction C (CLF) comprised 68% of the soil TOC in upper layer corresponding to the highest C pool in the upper layer. CLF and heavy‐fraction C (CHF) were not directly affected by the admixture of oak trees in both layers. The CHF accounted on average for 30% and 59% of the TOC at 0–10 and 10–20 cm depths, respectively. Despite low clay contents in the studied soils, the differences in the DCB‐extractable Fe and Al affected the concentrations of the CHF and TOC in the 10–20 cm layers (p < 0.05). Admixture of oak in pine stands contributed to reduce topsoil C stocks, probably due to higher soil organic matter turnover promoted by higher quality of oak litter.  相似文献   

16.
采用3种不同深度的50孔穴盘进行断根嫁接西瓜苗试验.试验证实嫁接苗龄20 d时,处理期间株高、茎粗、叶面积、叶片数等生物学特征无显著性差异,与根际营养体积无显著性相关关系.嫁接苗龄25 d之后,3个处理之间表现出株高、地下部干重、叶面积指数、壮苗指数及根系活力与根际营养体积间呈显著相关关系,78.4、82.9 cm3的孔穴体积抑制了秧苗的生长和根系生理活性.分析研究认为孔穴为78.4 cm3和82.9 cm3的穴盘苗嫁接后最佳苗龄为20 d,最长不宜超过25 d;孔穴为131.6 cm3的穴盘苗嫁接后苗龄可达30 d.  相似文献   

17.
SOME EFFECTS OF PARTIAL STERILIZATION ON MINERAL NITROGEN IN A LIGHT SOIL   总被引:3,自引:0,他引:3  
Field plots on loamy sand with and without the soil fumigant ‘D-D’ (dichloropropane-dichloropropene mixture) were sampled at 0–12, 12–24, and 24–64 cm depths during May 1966–8. All plots received 125 kg/ha nitrogen fertilizer early in March each year and were cropped with sugar beet. Fumigation increased the total amount of mineral (NH4++ NO3?) nitrogen to a depth of 64 cm by 67 kg/ha on average, largely by increasing NH4+-nitrogen. In unfumigated plots some mineral nitrogen leached below 12 cm every year but there was at least 125 kg/ha in the surface 64 cm in 1966 and 1968; during the wet spring of 1967 much leached below 64 cm. Sequential sampling in 1968 confirmed that some of the fertilizer nitrogen moved down the profile during spring. Plots partially sterilized either one or two years previously and not again, contained the same quantity of mineral nitrogen as plots that had never been fumigated. Fumigation increased the mineral nitrogen in plots by a similar amount whether or not they had been fumigated previously. Thus, on this soil, partial sterilization with ‘D-D’ seemed to have little residual effect on mineral nitrogen.  相似文献   

18.
为进一步优化全膜双垄沟播玉米生产技术,2020 — 2022年在甘肃省中部半干旱雨养农业区研究了玉米减穴增株高产高效栽培技术。试验采用全膜双垄沟播技术,设每穴留苗1株(对照)、留苗2 株、留苗3 株、留苗4株4个处理,观察各处理下玉米经济性状、生育期和产量。结果表明,在密度相同的情况下,全膜双垄沟播玉米每穴留苗3株、4株主要性状明显优于每穴留苗1株。每穴苗4株玉米折合产量为7 723.48 kg/hm2,较每穴留苗1株增产723.48 kg/hm2,增产率10.34%;每穴留苗3株为7 549.24 kg/hm2,较每穴留苗1株增产549.24 kg/hm2,增产率7.85%,增产效果明显。该种植模式适宜在海拔2 000 m以下、年降水量350~500 mm的半干旱雨养农业区应用。  相似文献   

19.
采用3种不同深度的50孔穴盘进行断根嫁接西瓜苗试验。试验证实:嫁接苗龄20 d时,处理期间株高、茎粗、叶面积、叶片数等生物学特征无显著性差异,与根际营养体积无显著性相关关系。嫁接苗龄25 d之后,3个处理之间表现出株高、地下部干重、叶面积指数、壮苗指数及根系活力与根际营养体积间呈显著相关关系,78.4、82.9 cm3的孔穴体积抑制了秧苗的生长和根系生理活性。分析研究认为:孔穴为78.4 cm3和82.9 cm3的穴盘苗嫁接后最佳苗龄为20 d,最长不宜超过25 d;孔穴为131.6 cm3的穴盘苗嫁接后苗龄可达30 d。  相似文献   

20.
Sources of competition for limited soil resources, such as nitrogen (N), include competitive interactions among different plant species and between plants and soil microorganisms (microbes). To study these competitive interactions, blue oak seedlings (Quercus douglasii) were grown alone or grown together with an annual grass, wild oats (Avena barbata) in pots containing field soil. We injected 15N-labeled ammonium, nitrate or glycine into the soil of each pot and harvested plants 5 days later. Plant shoots and roots, soil microbial N and soil KCl-extractable N were analyzed for 15N content. When oak and grass were grown together, 15N recovery from the inorganic N treatments (NH4+, or NO3) was 34, 9 and 4% for the grass, microbes and oak seedlings, respectively, and only 1% remained as KCl-extractable N. 15N recovery from the glycine treatment was 18, 22, 5% for the grass, microbes and oak seedlings, respectively, and 4% remained as KCl-extractable N. When oaks were grown alone, 15N recovery by soil microbes was 21, 48 and 40% in the NO3, NH4+ and glycine treatments, respectively. N forms had no effects on 15N recovery in oak seedlings (7%) and in KCl-extractable N pool (13%). In general, total N recovery by the grass was much greater than by oaks. However, on a fine root surface area or length basis, oaks exhibited higher N uptake than the grass. Our results suggest that the high rooting density and rapid growth rate of the annual grasses such as Avena barbata made them superior competitors for available soil N when compared to blue oak seedlings and to microbes. Soil microbes were better competitors for organic than inorganic N when annual grasses were present, but preferred NH4+ when competing only with oak seedlings.  相似文献   

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