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1.
砧穗组合选配是苹果矮砧集约栽培模式的关键环节之一,幼树的早果性和易成形性是评判砧穗组合优劣的重要指标,这与幼树根系分布密切相关。以中国苹果生产上8种常见砧穗组合的3年生幼树为试材,采用壕沟法对幼树根系分布特征进行了调查分析。结果表明:8种砧穗组合幼树根构型可以分为M系自根砧、SH系自根砧、乔化实生砧木、M系中间砧和SH系中间砧等5大类。3年生幼树根系在水平和垂直方向分布的主要区域,均为0~60 cm范围,且随着距树干距离的增大而减少。不同砧穗组合,根系长度以直径2 mm级别的根系最长,其次是直径2~5 mm的,10 mm以上的最短。乔化实生砧木较无性系矮化砧木粗根多,而须根少;矮化砧木中,M系砧木根系构成以须根为主,而SH系和青砧系砧木须根均较少。砧穗组合幼树的地上分枝数、短枝比例和花芽数等早果性指标与细根(直径2 mm)根系数量、根长密度和根表面积密度呈显著正相关关系。通过对各砧穗组合易成花和易成形的综合评价,认为在渭北黄土高原有灌溉条件的地区,M系自根砧和中间砧组合早果性好和易成形较好,而SH系组合较易成形,但早果性较差,乔化组合早果性最差,这与它们的根系分布特征密切相关。  相似文献   

2.
In peach, xylem anatomical characteristics have been shown to be related to vigour of selected rootstocks. The goal of this research was to determine if xylem characteristics of a new set of rootstocks that exhibit a range of size-controlling potential and have a different genetic background from previously examined material would also exhibit similar differences in xylem characteristics. If so, then anatomical analysis of xylem may be a useful means of predicting the vigour control capacity of selected peach rootstock genotypes. Samples of xylem tissue were taken from roots, trunks and shoots of four new rootstocks that were derived from a genetic cross between ‘Harrow Blood’ and ‘Okinawa’ peaches and compared with tissue from ‘Nemaguard’, a vigorous control. Xylem samples were sectioned and analysed by optical microscope. The number and dimensions of vessels in recently developed xylem of each rootstock were measured and compared. The more dwarfing rootstocks had fewer large vessels and more small vessels than the more vigorous rootstocks. Weighted mean vessel diameter (Wm) and calculated hydraulic conductance (Kh) differed among rootstocks: more vigorous rootstocks had higher Kh and Wm than dwarfing rootstocks. Rootstock xylem vessel dimensions varied in relation to the vigour they imparted to a common scion cultivar (‘O’Henry’). After the ‘Nemaguard’ control, ‘HBOK 50’ was the most vigorous rootstock followed by ‘HBOK 10’, ‘HBOK 32’ and ‘HBOK 27’, respectively. Thus, as was seen in previous research with a separate set of rootstocks, the vigour-control capacity of this new series of peach rootstocks was strongly related to their xylem hydraulic characteristics and it appears likely that it would be possible to use xylem anatomical characteristics of shoots or roots of young trees to pre-select for size-controlling potential in a rootstock development program.  相似文献   

3.
The rates of sap flow and xylem vessel features were studied in two-year-old nongrafted and grafted avocado (Persea americana Mill.) trees. Daily sap flow rates were measured with heat and balance stem gauges in clonal Duke 7 (D7) and Toro Canyon (TC) trees and ‘Hass’ clonal scions grafted onto clonal D7 (H/D7) and TC (H/TC) rootstocks. Vessel features as size, number and total vessel area were determined histologically in the stem of the scion and rootstock and the roots of the grafted trees. Significant differences in the sap flow rate were found among the rootstocks, where D7 had a 29% higher sap flow rate than did TC (grafted and nongrafted trees). There were no differences among xylem vessel features in the stems of any of the varieties. However in the roots, D7 had wider and fewer vessels then TC do. Also, D7 had a 19% higher total vessel area than TC. These results suggest that the differences in water consumption of ‘Hass’ on different rootstocks may be associated with differences in the efficiency of the roots to absorb water across conductive tissue which may be linked to differences in the area of xylem vessels in the root.  相似文献   

4.
为了分析不同砧木苹果树细根的发生和周转动态,连续4年通过微根管技术研究不同砧木的5年生‘富士’苹果砧穗组合细根中的活根根长密度与死根根长密度的动态变化,以及细根年周转率和季节周转率。结果表明,乔砧树富士/八棱海棠的活根根长密度最大,矮化中间砧富士/M9/八棱海棠和富士/SH40/八棱海棠次之,矮化自根砧富士/M9、富士/SH40和富士/小金海棠最小,所有砧穗组合的活根根长密度随着树龄的增加而逐年减小。所有砧穗组合在夏秋季出现细根发生和死亡高峰期,乔砧树富士/八棱海棠的细根发生高峰期的活根根长密度和死根根长密度均最大。细根年周转率和季节周转率年度间差异大,矮化自根砧树细根的年周转率高于乔砧树。矮化自根砧和矮化中间砧树体ARLD低于乔砧树体可能与其致矮性相关。  相似文献   

5.
苹果紧凑型品种和矮化砧木内源激素的变化   总被引:15,自引:0,他引:15  
 以不同矮化程度的苹果品种、砧木和嫁接植株为试材, 研究了春梢旺长期茎尖和春梢停长期叶片的内源激素变化。结果表明, 与普通型品种相比, 紧凑型品种茎尖和叶片中的赤霉素(GA1+3) 含量显著降低而细胞分裂素(CTKs) 含量显著升高, 紧凑型品种的CTKs/ GA1+3约是普通型品种的4. 78~6. 24 倍。不同矮化程度的砧木间相比, 春梢停长期叶片中, M9 的IAA 含量比平邑甜茶降低了53. 7 %。以M26作中间砧, 显著降低了接穗叶片和基砧新根中的IAA 含量; 但以新红星作中间砧则影响较小。将M26中间砧与紧凑型品种组合, 与乔砧普通型植株相比, 春梢停长期叶片和新根中IAA 含量分别下降了46. 1 %和33. 3 %,CTKs/ GA1+3是其2. 8~3. 6 倍, 其变化幅度大于单一应用矮化砧、紧凑型品种的植株。因此, 紧凑型、矮化砧具有不同的激素调节机制, GA、CTK在紧凑型品种矮化中起重要作用, 而矮砧的矮化可能与IAA 密切相关。  相似文献   

6.
 以基砧为八棱海棠的4 种不同中间砧嫁接的苹果幼苗长富2 号/八棱海棠、长富2 号/M9、长 富2 号/M26、长富2 号/SH6 为试验材料,研究干旱胁迫对其导水特性的影响。结果表明:在干旱胁迫下, 4 种中间砧木嫁接苗的整体、冠层、茎干、根系叶比导水率均有减小,各器官叶比导水率基本趋势是乔 化 > 半矮化 > 矮化,其中矮化中间砧的变化幅度最大,乔化中间砧的变化最小。中间砧嫁接口导水阻 力表现为矮化砧比半矮化、乔化砧高,在正常水分条件下,八棱海棠、M9、M26 和SH6 中间砧嫁接区域 导水阻力在植株总体导水阻力中所占的比率分别为4.07%、6.60%、4.97%和5.11%,当受到干旱胁迫后, 嫁接区域所占比率均有不同程度减小。由于矮化苗有效导水率长期低下,根系吸水和运输水分的能力下 降,导致地上部分水分供给减少,从而影响树体的生长。  相似文献   

7.
Summary

The apple rootstocks M.9, M.26, and MM.106 were evaluated for their efficiency in bringing ‘Discovery’ apple trees into production. The experiment, carried out over a ten- year period compared two planting densities at 1666 and 3333 trees per ha. Tree vigour differences between rootstocks were measured in term of trunk growth, tree volume, weight of branches pruned off and final weight of the above-ground parts of the trees. Fruit production is presented both as total yield and as the weight of first class fruit. Fruit colour development is also shown. Cropping efficiency is calculated and presented in relation to the different vigour measurements. The results confirm that ‘Discovery’ is slow to come into production. M.9 was the most productive rootstock, but due to vigour differences MM.106 gave the same yields per tree, although the latter had the lower yield efficiency. M.26 performed poorly; its vigour was similar to M.9 but it produced the lowest yields.  相似文献   

8.
 以2 年生大田栽培矮化中间砧富士苹果(宫藤富士/SH6/平邑甜茶)幼树和乔砧富士苹果(宫 藤富士/平邑甜茶)幼树为试材,通过春季土施15N–尿素研究了SH6 矮化中间砧对苹果幼树N 素的吸收、 利用及贮藏的影响。结果表明:与SH6 矮化中间砧幼树相比,乔砧幼树长势强,净生长量大。树体各器 官的Ndff 值均表现为乔砧幼树大于SH6 矮化中间砧幼树;两种类型苹果幼树15N 分配率表现出一致规律, 即叶片中最高,新梢和粗根中次之,中心干最小,其中40% ~ 70%氮素分配给新生器官(新梢和叶);秋 梢停长期,乔砧幼树地上部新生器官N 肥分配率(63.66%)明显高于SH6 矮化中间砧幼树(57.68%), 乔砧幼树氮素利用率(14.32%)显著高于SH6 矮化中间砧幼树氮素利用率(8.55%);秋季落叶后,乔砧 幼树叶片中有33.11%的氮素回撤到树体内,而SH6 矮化中间砧幼树叶片有36.92%回撤到树体内,除细根 外,各个器官均有氮素回流贮藏,其中粗根和皮层是苹果氮素主要的贮藏部位,乔砧幼树地下部氮素增 量为8.34%,明显大于SH6 矮化中间砧幼树的增量6.85%。SH6 中间砧对苹果幼树氮素吸收及回流上均 有显著的阻滞作用。  相似文献   

9.
以SDC系苹果砧木为试材,连续9年观察测定了在其上嫁接的红星树早花、早果、产量、果实品质、树体矮化性及抗逆性等生物学特性.结果表明:(1)SDC系砧木嫁接树比M9、M7嫁接树提早1—2年开花结果,幼树期花量密度大,5年生树总花序数及单位树干面积花序数明显高于M9和M7.(2)果皮色素花青苷含量、果肉可溶性固形物含量和总糖量均高于M9和M7,果实外观及内在品质优良.(3)幼树矮化性状明显,树高/冠幅比值小,树冠开张.(4)叶片蒸腾量小.脱落酸(ABA)含量高,单位叶面积失水量小,抗旱性强.(5)嫁接树越冬期枝条自由水含量高,抽条率明显低于M9,越冬能力强.并对SDC系砧木的优缺点进行了讨论.  相似文献   

10.
A preliminary report is made of the results of growth studies on compound trees formed of all combinations, as scion and as rootstock, of three clones, chosen to cover a wide range of vigour, of apple, quince, and cherry.Under the conditions of the orchard experiment and with the clones used, no distinction was found between influence on tree size of any apple clone when present as the scion or when present as the rootstock. Major distinctions in influence as scion or as rootstock were, however, found in the case of certain quince and cherry clones.Evidence as to the precocity of flowering of the scions of compound trees indicates that precocity may occur independently of a dwarfing influence.  相似文献   

11.
不同砧木红星苹果树含水量的日变化呈V形曲线。以乔砧树最高,M7中间砧树次之,M9中间砧树较低。含水量的日变化与水势的日变化基本呈正相关。地上部含水量与砧木解剖构造有关,矮砧树地上部含水量较低是导致树体矮化的因素之一  相似文献   

12.
In composite (scion–rootstock) dwarfing fruit trees, an overgrowth at the graft union is often observed, the severity of which is correlated with degree of dwarfing. The graft union of dwarfing sweet cherry (Prunus avium L.) rootstocks may limit soluble sugar transport or starch mobilization, leading to localized accumulation. Soluble sugars and starch were measured in the tissues surrounding the graft union of young ‘Rainier’ (2002) and ‘Lapins’ (2003) sweet cherry trees on ‘Gisela 5’ (‘Gi 5’; dwarfing) and ‘Colt’ (vigorous) rootstocks. Separate rootstock shank, rootstock, graft union, and scion tissues were analyzed for both starch and soluble sugar content throughout the growing season in both years. Starch concentrations did not vary among locations within the graft union for ‘Rainier’ on either the dwarfing or vigorous rootstock, or for ‘Lapins’ on the dwarfing stock. However, for ‘Lapins’ on ‘Colt’, starch was highest in the rootstock shank and declined vertically (rootstock shank ? rootstock > union > scion). Soluble sugar concentrations were generally similar to or higher in scions on ‘Gi 5’ than on ‘Colt’, and were similar to or lower in the rootstock and rootstock shank on ‘Gi 5’ than on ‘Colt’. Results suggest that rootstock has a significant effect of localized accumulations of carbohydrates above and within the graft union of ‘Gi 5’ and below the graft union of ‘Colt’.  相似文献   

13.
Apple scions with diverse growth habits were grafted on various size-controlling rootstocks and morphological characteristics were measured after 6 years of growth in the field. Scion had more influence than rootstock on monthly growth rate. Across all rootstocks, scions with spreading growth habits grew rapidly in April and May and achieved most seasonal growth earlier than scions with upright growth habits that grew slowly early in the season. In all growth habits and rootstocks, growth rate slowed appreciably but did not cease by August and growth did not terminate earlier for any one scion–rootstock combination. Across all scions, the dwarfing rootstock, M.9, consistently had the lowest and seedling rootstock had the greatest tree height and trunk diameter. However, no one size-controlling rootstock consistently influenced dates of bud break and full bloom, shoot elongation rate, or duration of growth. Tree growth form was not fundamentally affected by rootstock. Significant interactions indicated that effects of size-controlling rootstock on components of shoot growth will vary with apple tree growth habit. These effects on phenology and development can be significant to growers and may assist breeders in developing new apple cultivars.  相似文献   

14.
Studies were carried out on the assumption that incompatibility and dwarfness of budded citrus trees result from impaired carbohydrate movement through the bud union. During autumn and winter, carbohydrates move from the tree tops to the roots and accumulate there as starch. The starch level can therefore serve as a measure of this movement.

This study included seven rootstock varieties all budded with Shamouti orange scion. Unbudded rootstocks were included for purposes of comparison.

The starch level was determined in the winter at three locations along the trunk, below and above the bud union.

No clear indication was found of impaired carbohydrate movement through the bud union of stock-scion combinations showing very low growth vigour.

The starch level in bark and wood in the rootstock portion of trunks of budded trees was found to be negatively correlated with tree size. This indicates that the level of starch in the trunk results from the growth vigour of the tree and is not a factor which affects it.

No relation was found between starch level in the scion and tree size. Based on these findings the conclusion was reached that incompatibility and low growth vigour in citrus do not seem to result from impaired carbohydrate movement through the trunk and starvation of the roots.  相似文献   

15.
SUMMARY

Basipetal auxin transport in shoots obtained from dwarfing (M.9) and less-dwarfing (MM.Ill) apple rootstocks was compared by measuring the distribution of 14C-[carboxy]-labelled indole 3-acetic acid supplied to the ends of shoot segments. Auxin was transported at a greater velocity in MM.111 than in M.9 in both woody and non-woody segments, and was effectively blocked by 2,3,5-triiodobenzoic acid. The velocities of transport were approximately 8 mmh-1 in M.9,and 13 mmh-1 MM.111. There was relatively little transport in the acropetal direction, and there were no significant differences between the two rootstocks in total auxin uptake. Chromatographic analysis suggested that the metabolismof the auxin supplied followed similar patterns in both rootstocks. It is proposed that the slower auxin transport in M.9 results from a lower capacity for auxin efflux from transporting cells. A limiting auxin transport capacity could explain the reduced cambial activity and subsequent xylem formation in the dwarfing M.9 rootstock, and thus make an important contribution to the dwarfing effect.  相似文献   

16.
Crown rot (Phytophthora cactorum) of apple came to prominence in Britain in the 1960s following the introduction of certain new types of clonal rootstock. Observations on crown rot outbreaks over the period 1976 to 1984 were as follows: only rootstocks MM.104 and MM.106 were affected; infection occurred in the bark of the rootstock axis below soil level; lesions caused girdling, and sometimes appeared above soil level and spread down the main roots to a limited extent; infection occurred during the growing season but disease development ceased with the onset of winter and did not resume the following year; trees on MM. 104 were susceptible throughout the life of the orchard while trees on MM. 106 were susceptible only during the establishment years; disease outbreaks occurred sporadically. Crown rot was induced experimentally on trees grafted on MM. 104 and on M.25 in field trials by irrigating for four to six weeks through inoculum applied to the base of the trees. No crown rot occurred on grafted MM.106 or on other grafted rootstocks in the same trials, although lesions developed above soil level on a few trees. The limited experimental data indicated that MM.104 was liable to attack throughout the growing season. The importance of environmental influences on the susceptibility of rootstocks to crown rot is discussed.  相似文献   

17.
Observations of respiration rate, catalase activity, peroxidase activity ("physiological activity”) and some growth characteristics of roots of citrus rootstocks were made. These were carried out on seedlings of six rootstocks (five months old) grown in a standard nutrient solution and on seven- to ten-year-old budded trees growing in the orchard. The orchard trees, budded to only one scion, were propagated from the same rootstocks as were used in the study of seedlings. A comparison was also made of five additional scions budded on one rootstock.

A technique was developed to obtain roots from orchard trees for the study of their physiological and morphological characteristics.

Significant differences were found in the “physiological activity” of the roots of the various rootstock seedlings. A significant negative correlation existed between the final length of the main root and the physiological activity of the roots.

Significant differences were found in the physiological activity of the roots of the various mature rootstocks budded with Shamouti sweet orange.

A clear and marked influence of the various scions on the physiological activity of the roots of sour-orange was observed.

An obvious positive correlation existed between respiration rate, catalase activity and peroxidase activity of the roots. The use of two different H-donors resulted in differences in the peroxidase activity measured.

The significance of the variations in physiological response of the roots is discussed.  相似文献   

18.
Summary

Yield and fruit quality parameters were studied during three years in a rootstock trial with ‘Mutsu’ apple. M.9 provided the highest total yield per tree as well as the highest quantity of intermediate and yellow fruit as compared with J.9, M.26 and B.9. Within similar colour categories, significant differences among rootstocks were also recorded for fruit starch degradation pattern (SDP), firmness, titratable acidity (TA), and soluble solids concentrations (SSC). Generally, the lowest SDP and highest fruit firmness was found in fruits from trees on rootstocks M.26 and B.9. The highest TA was found in fruits from trees on rootstocks M.26 and J.9 and the highest SSC in fruits from trees on rootstocks J.9 and B.9. However, considerable variations among years were also recorded. Correlation coefficients between rootstock and several yield and quality parameters revealed high correlation to SSC for green fruit and TA for medium and yellow fruit and some correlation to yield and number of fruits per tree.  相似文献   

19.
In horizontal apple stems extension shoots were usually produced only from buds on the upper side of the stem, while buds on the lower side remained dormant or grew into spurs, and the same tendency was shown in inclined stems bent so that the “upper” and “under” sides became reversed.

On horizontal stems lateral shoots showed a gradient of vigour, the longest shoots being produced by proximal buds. Xylem of horizontal branches was epitrophic. These responses appear to be due to effects of gravity on the distribution of endogenous growth regulating factors within stems. In trees grown horizontally and rotated, shoots and spurs grew from all sides of the stem and xylem developed concentrically.

In studies of lateral shoots of partly disbudded horizontal stems, including cincturing treatments, it was found that the vigour of basal lateral shoots was a function of bud position in relation to the apex rather than in relation to the roots.

A model for lateral shoot growth in horizontal branches is proposed, in which shoot vigour is related to the position of buds along a postulated inhibitory gradient.  相似文献   

20.
SUMMARY

Seedlings of Newton Wonder apple multiplied vegetatively on Malus sikkimensis rootstocks and inoculated with a mixture of chlorotic leaf spot, scaly bark and dwarf viruses showed varying degrees of sensitivity to these infections. Although none of the seedlings showed clearly recognizable virus symptoms, many showed a very significant reduction in vigour; the girth differences between the infected and control trees were still increasing three years after inoculation.

The infected trees showed signs of nitrogen deficiency, which were confirmed by foliar analysis ; leaf nitrogen as percentage of dry weight was 10% lower than that of the controls. Four years after inoculation 43% of the infected trees had cropped, compared with 58% of the controls.

The results suggest that some selection of varieties and rootstocks for virus tolerance has taken place and that the evaluation of new seedlings on their own roots may not be a true indication of their performance on infected rootstocks.  相似文献   

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