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1.
Forest floor characteristics influence nutrient cycling and energy flow properties of forest ecosystems, and determine quality of habitat for many forest plants and animals. Differential crown recession and crown development among stands of differing density suggest that an opportunity may exist to control the input of fine woody litter into the system by manipulating stand density. The objective was to measure the rate of branch mortality among stands of differing density and to estimate the range in total per hectare necromass inputs. Although litter traps are reliable for estimating per hectare rates of litterfall, branch mortality dating on sectioned stems uniquely allows assessment of several other litterfall components: (1) individual tree contributions to total litterfall; (2) the amount of branch material released by mortality, regardless of whether the branches are shed to the forest floor; (3) the distribution of basal diameters characterizing the litterfall from a given tree and stand. Twenty-four trees were felled and sectioned on permanent plots that were part of a silvicultural study of stand density regimes in Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco.). Whorl branches were dissected out of bole sections to determine the dates of mortality, and a branch biomass equation was applied to estimate potential rate of litterfall. Periodic annual rates were expressed in four ways: (1) number of branches per tree; (2) mass of branches per tree; (3) mass of branches per unit of crown projection area; (4) mass of branches per hectare. For the growth periods investigated, larger trees and trees growing on denser plots tended to release a greater necromass through branch mortality. Average branch basal diameter generally decreased with increasing stand density. Annual branch mortality ranged from 33 to 430 g m−2 crown projection area for individual trees, and from 236 to 1035 kg ha−1 for individual plots. These rates approached the low end of the range of previously published fine litterfall rates for Douglas-fir. Rates on these plots were relatively low owing to the temporary delay in crown recession imposed by artificial thinning. A conceptual model of branch litter dynamics is presented to depict consistencies with crown development among stands managed under different density regimes.  相似文献   

2.
The aim of this study was to develop statistical models for first order branchiness in young planted forest stands of pedunculate oak (Quercus robur L.) and European beech (Fagus sylvatica L.), and to give an ecological and silvicultural interpretation to these models. The reported models focus on the lower most-valuable stem part (i.e. until 6 m height), and cover different tree development classes to capture the development of branchiness over time. For each species 30 study plots were selected spread over two nearby forests in Flanders (northern Belgium), minimising site and genetic variability. Branches were counted on a total of 399 oak and 376 beech trees. On a subsample of 30 trees per species (one tree per plot), detailed non-destructive branch measurements were performed, yielding data for 555 oak and 438 beech branches. For both species, models for tree self-pruning (i.e. total branch number and dead branch portion), branch mortality and branch architecture (i.e. branch diameter and branch insertion angle) were built. A generalised linear mixed modelling approach was adopted. The models for total branch number and dead branch portion may be interpreted in terms of four processes contributing to self-pruning: (1) stand and tree development, (2) tree competitive status, (3) stand density and (4) site humidity. The reported models reveal similar self-pruning rates in oak and beech, but with different driving factors: early branch dying and slow shedding for oak and the other way around for beech. Mortality of individual branches is further determined by branch position and branch dimension. Branch diameter and branch insertion angle of both species are mainly related to branch cord length and relative branch position. All modelled effects are consistent with known ecological and ecophysiological processes. Silvicultural implications for stand establishment and early tree selection are discussed. The reported models can be used to fine-tune operational silvicultural choices for quality timber production. This is a first step towards the integration of branchiness models for oak and beech into forest growth simulators.  相似文献   

3.
The stand density of a forest affects the vertical distribution of foliage. Understanding the dynamics of this response is important for the study of crown structure and function, carbon-budget estimation, and forest management. We investigated the effect of tree density on the vertical distribution of foliage, branch, and stem growth, and ratio of biomass increment in aboveground tissues; by monitoring all first-order branches of five trees each from thinned and unthinned control stands of 10-year-old Chamaecyparis obtusa for four consecutive years. In the control stand, the foliage crown shifted upward with height growth but the foliage quantity of the whole crown did not increase. In addition, the vertical distribution of leaf mass shifted from lower-crown skewed to upper-crown skewed. In the thinned stand in contrast, the foliage quantity of individual crowns increased two-fold within 4 years, while the vertical distribution of leaf mass remained lower-crown skewed. The two stands had similar production rates, numbers of first-order branches per unit of tree height, and total lengths of first-order branches. However, the mortality rate of first-order branches and self-pruning within a first-order branch were significantly higher in the control stand than in the thinned stand, which resulted in a higher ratio of biomass increment in branch. Thinning induced a higher ratio of biomass increment in foliage and lower in branch. The increased foliage quantity and variation in ratio of biomass increment after thinning stimulated stem growth of residual trees. These results provide information that will be useful when considering thinning regimes and stand management.  相似文献   

4.
长白落叶松人工林树冠形状的模拟   总被引：17，自引：3，他引：17  
李凤日 《林业科学》2004,40(5):16-24
以长白山地区 2 6a生长白落叶松人工林为研究对象 ,采用枝解析的方法 ,测定了 2 5株林木 (直径 10 5～2 4 9cm)的树冠变量 ,并建立了预测树冠外侧形状的冠形模型。基于枝条着枝深度 (DINC)和林木变量所建立的树冠形状模型包括 :基径、枝长、着枝角度和弦长等预估模型。对于大小相同树木的主要枝条来讲 ,这些树冠变量是随着DINC的增加而增大 ;而林木的胸径 (DBH)和树高 (HT)变量很好地反映了不同大小树木的冠形变化。冠形预测模型预测效果良好 ,充分体现了树冠结构的变化趋势 :树冠形状在树冠的中上部呈抛物线体 ,而在树冠的下部则为近圆柱体。文中所建模型 ,可以合理地描述长白落叶松人工林的树冠形状及其变化规律。  相似文献   

5.
目标树经营是一种通过降低邻木冠层竞争、增加目标树生长空间来提高单株木质量的营林技术, 可以看作是一种特殊的抚育间伐。文中从目标树选择标准、目标树密度、树冠重叠释放作业和作业时间等方面阐述了目标树经营体系, 并综述了目标树经营对单株目标树生长(直径、蓄积、树高、树冠)的影响和成本效益分析等研究进展, 以期为我国森林经营实践提供借鉴。  相似文献   

6.
[目的]研究不同强度采伐下马尾松的生长动态,筛选适宜的采伐强度,为马尾松人工林近自然经营提供技术支撑。[方法]2007年10月在14年生马尾松人工林(保存密度1 100株·hm~(-2))内进行采伐试验,设置4个采伐强度,即保留密度分别为225、300、375、450株·hm~(-2),以不采伐为对照;其后,自2008年开始连续8 a,每2 a测定1次马尾松的胸径、树高、枝下高和冠面积等生长指标,并计算单株材积和林分蓄积量,应用方差分析和Duncan多重比较分析生长指标对不同采伐强度的动态响应。[结果]表明:采伐强度显著影响林分生长,其中,林分平均胸径、单株材积、冠面积的年均增长量随保留密度增大而减小,但均显著高于对照(P0.05)。采伐后第1 3年,马尾松冠面积增长量显著高于采伐后期,胸径则在采伐后第3 5年最高,而不同采伐强度对林分树高生长影响不明显。保留密度显著影响林分枝下高和蓄积量的动态变化,其年均增长量随密度增大而递增。5个处理间林分蓄积年均增长量的差异随林龄的增大而逐渐缩小。[结论]马尾松人工林生长对不同强度采伐的动态响应以树冠最敏感,冠面积首先陡然增大,进而引起胸径的快速生长。树高和枝下高在采伐后年均增量变化相对平稳。4个采伐强度均显著促进单株材积生长,而仅保留密度为225株·hm~(-2)的采伐对林分蓄积增长量影响显著。综合比较林分的单株材积和林分蓄积连年增长量,建议在桂西南15年生马尾松人工林近自然经营中宜选择300株·hm~(-2)的保留密度进行采伐。  相似文献   

7.
We studied the combined effects of thinning on stand structure, growth, and fire risk for a Scots pine thinning trial in northern Spain 4 years following treatment. The thinning treatments were: no thinning, heavy thinning (32–46% of basal area removed) and very heavy thinning (51–57% of basal area removed). Thinning was achieved via a combination of systematic and selective methods by removing every seventh row of trees and then by cutting suppressed and subdominant trees in the remaining rows (i.e., thinning from below). Four years after thinning, mean values and probability density distributions of stand structural indices showed that the heavier the thinning, the stronger the tendency towards random tree spatial positions. Height and diameter differentiation were initially low for these plantations and decreased after the 4-year period in both control and thinned plots. Mark variograms indicated low spatial autocorrelation in tree diameters at short distances. Diameter increment was significantly correlated with the inter-tree competition indices, and also with the mean directional stand structural index. Two mixed models were proposed for estimating diameter increment using a spatial index based on basal area of larger trees (BALMOD) in one model versus spatial competition index by Bella in the other model. As well, a model to estimate canopy bulk density (CBD) was developed, as this variable is important for fire risk assessment. Both heavy and very heavy thinning resulted in a decrease of crown fire risk over no thinning, because of the reduction in CBD. However, thinning had no effect on the height to crown base and thus on the flame length for torching. Overall, although thinning did not increase size differentiation between trees in the short term, the increase in diameter increment following thinning and the reduction of crown fire risks support the use of thinning. Also, thinning is a necessary first step towards converting Scots pine plantations to more natural mixed broadleaved woodlands. In particular, the very heavy thinning treatment could be considered a first step towards conversion of overstocked stands.  相似文献   

8.
The effects of pre-commercial thinning on tree and branch characteristicswere analysed at two experimental sites in northern Sweden,where pre-thinning of Scots pine stands to 600, 1000 and 1800stems per hectares at heights of 1.5, 3, 5 and 7 m had beencombined in a factorial design. A total of 90 trees were cutand branches were measured in whorls sampled at 1-m intervals,starting with the whorl closest to 0.5 m above ground. Effectsof the treatments on tree and branch characteristics were analysedby a fixed-effect analysis of variance model, which was appliedseparately for each experimental site and whorl height. Diameterat breast height (d.b.h.) and living crown to height ratio decreased,while the height/d.b.h. ratio (stem form) increased with increasingstand density, and generally with increased height at thinning.No treatment effect was found on the number of branches perwhorl or branch angle, but branch diameter (BD) was significantlyinfluenced by both stand density after thinning and height atthinning. BD decreased with increases in stand density and decreasedwith increases in height at pre-commercial thinning. Relativebranch size (RBS), defined as the ratio between the sum of thediameters of branches in a whorl and stem perimeter under barkat the location of that whorl, was significantly influencedonly by stand height at the time of treatment. RBS decreasedby 4–6 per cent in whorls located below the current baseof the living crown with each step increase in stand heightat pre-commercial thinning. The results indicate that the numberof stems after pre-commercial thinning has a greater impacton tree size and shape than timing of pre-commercial thinning.Stand density after thinning also has a greater impact on branchsize, in the sense that it will affect branch size further upthe tree than the timing of pre-commercial thinning. However,by delaying the time for pre-commercial thinning it is possibleto reduce the RBS in the lower part of the stem. This mightbe an important measure to improve timber quality later in therotation period, which cannot be done solely by regulating standdensity.  相似文献   

9.
Recent advances in traditional forest growth models have been achieved by linking growth predictions to key ecophysiological processes in a hybrid approach that combines the strengths of both empirical and process-based models. A hybrid model was constructed for intensively managed Douglas-fir plantations in the Pacific Northwest, USA, by embedding components representing fundamental physiological processes and detailed tree allometrics into an empirical growth model for projecting individual tree and stand development. The simulated processes operated at a variety of scales ranging from individual branches to trees and stands. The canopy structure submodel improved predictions of leaf area index at the stand level when compared to allometric and other empirical approaches (reducing mean square error by 30–42%). In addition, the hybrid model achieved accuracy in short-term volume growth prediction comparable to an empirical model. Biases in 4-year stand growth predictions from the hybrid model were similar to those from the empirical model under thinning, fertilization, and the combination of these treatments; however, volume growth predictions in unmanaged plantations averaged approximately 36% less bias. These improvements were attributed to detailed information on crown structure (i.e. size, location, and foliage mass of primary branches), simple representation of key physiological processes, and improved site characterization. Soil moisture, temperature, and nitrogen mineralization predicted by the hybrid model also agreed closely with observed values from several previous studies. Overall, the model framework will be helpful for future analyses as it can lend insight into the influence of weather and site edaphic factors on growth, help identify mechanisms of response to silvicultural treatments, and facilitate the design of sound management regimes for Douglas-fir plantations across the Pacific Northwest region.  相似文献   

10.
It is recognized that estimation of internode length and maximum branch size is important for the prediction of clearwood in unpruned timber stands, as well as for evaluating the quality and value of logs in general. A review of existing branch models reveals a diversity of approaches as well as a tendency for results to be species specific. Here, a branch model is developed for Pinus radiata in New Zealand, capable of predicting successive internode lengths, the number of branches on each branch cluster, and the size of each branch up to the green crown (GC) at site index age 20. Inputs to the model include tree height and diameter at breast height (dbh) (both at age 20) and basal area per hectare of the top 100 stems. Further optional inputs are an ocular count of the number of branch clusters up to the green crown, and branch factor (BF)—the size of the biggest branch in the first cluster encountered above 6 m. The vertical distributions of internode length and maximum branch diameter are found to reach maxima around 0.3–0.4 of relative height. Internode length and the number of branches per whorl were found to be independent of tree size, site index and stand density. Stems per hectare is not required as an explicit predictor variable but it appears implicitly through tree dbh. Some model output is given and the results are discussed.  相似文献   

11.
抚育间伐对人工红松林生长效应的影响   总被引：2，自引：0，他引：2  
利用不同间伐强度后40 a的人工红松林,连续抚育3次的长期定位观测资料,分析了抚育间伐对人工红松单木胸径和材积、林分断面积和蓄积、林分枯损和总收获量的影响。研究结果表明:间伐可以增加单木平均胸径和材积的生长率,且不同间伐强度表现为相同的规律,即中度(33.2%)强度(43.4%)弱度(23.1%)对照(0.0%);间伐后林分的断面积和蓄积生长率,具有相似的自然规律,间伐样地的生长率均高于对照;林木枯损率随间伐强度的增加而降低;间伐后的林分总收获量平均增加11.6%,间伐林分之间差别不明显。在整个研究阶段,不同的抚育阶段表现不同的规律,抚育间伐可以延缓单木胸径和材积生长速度的降幅;而不同抚育阶段内,林分断面积和蓄积生长率均为增长,也就是说,在一定阶段内间伐可以有效地提高林分平均断面积和蓄积生长率;从林分枯损状况来看,应该减少间伐间隔期。  相似文献   

12.
文章以广东省湛江特呈岛高潮带、中潮带、低潮带白骨壤纯林为研究对象,调查了其树高、地径、冠长、冠幅以及分枝角度等,揭示了其生长及树冠特征.结果表明:不同潮带间白骨壤冠幅及树冠面积差异不显著(P＞0.05),而树高、地径、冠长及树冠体积均存在显著差异(P＜0.05),中潮带、高潮带与低潮带的树冠体积分别达31.522,15...  相似文献   

13.
Thinning of Korean pine (Pinus koraiensis Sieb. et Zucc.) is used to facilitate timber and cone production. The present study in Northeast China investigated the effects of thinning intensity on individual tree growth, temporal variation in cone yield, and seed quality in Korean pine plantation. In 2005, five thinning intensity levels (none, extreme, heavy, moderate and light) were set in 15 permanent plots in a 32-year-old Korean pine plantation at Mengjiagang Forest Farm, Jiamusi City, Heilongjiang Province. We recorded tree growth and seed cone production from 2013 to 2016, i.e., from 8 to 11 years after thinning. Except for height growth, thinning increased tree growth (diameter at breast height and crown size) and improved cone yield. The extreme thinning treatment (to 300 trees per hectare) resulted in the largest tree diameter, tree volume, crown size and 4-year cone production per tree. The highest cone yield per tree in the mast year (2014) was observed when stands were thinned to 500 trees per hectare (heavy thinning). Although the best cone and seed quality and the largest cone and seed mass per tree were recorded in the heavily thinned stand, no significant differences were found between heavy and moderate thinning stands (750 trees per hectare). At the stand level, the moderately thinned stand had the highest basal area, stock volume and seed cone production per stand. Our results suggest that thinning to 750 trees per hectare will improve timber and cone productivity in 40-year-old P. koraiensis stands.  相似文献   

14.
Models of Douglas-fir branch and whorl characteristics were developed from contrasting spacing experiments in southwest Germany. The dataset was based on 100 young (20–30 years old), unpruned and partially pruned trees from a 100, 200, and 1200 stems ha−1 spacing experiment on Douglas-fir that was replicated 3 times across the region. The material was used to predict (1) the number of branches whorl−1, (2) branch angle, (3) status (living/dead) of the branches within the living crown, (4) maximum branch diameter whorl−1, and (5) relative diameter of branches within a whorl. For each of these models (except branch status), both a linear and nonlinear, generalised hierarchical mixed effects equation was developed. The comparison of the linear and nonlinear approaches showed that both had a relatively similar level of bias, but the nonlinear equations generally performed better (reduction in mean absolute error of 1.1–69.5%). Overall, individual branch and tree properties were sufficient to give logical and precise predictions of the branch characteristics for the models across the range of sampled stand densities. In addition, the models showed a similar behaviour compared to models on Douglas-fir crown structure from the Pacific Northwest, USA. This suggests that the allometric relationship between tree size and branch characteristics for a given species may be relatively consistent across regions, even ones with highly contrasting growing conditions like in this study. The models performed well across a range of stand conditions and now will be further integrated into an individual tree growth and yield simulations system.  相似文献   

15.
Both crown and branch characteristics greatly affect tree growth and timber quality, and their development is closely related to planting density (PD). Here crown and branch attributes of Betula alnoides were investigated in a 14-year-old experimental plantation in southern China with five planting densities ranging from 500 to 3333 stems per hectare (sph). The results showed that high PD significantly reduced crown and branch sizes. The ratios of crown diameter to stem diameter at breast height at the stand level and for dominant and co-dominant trees were almost constant regardless of PD. PD also had no significant influence on the number and angle of branches, and only planting with the highest density significantly increased branch density and mortality. Insertion angle (IA) of dead branches was greater than that of live branches and these differences were significant at the two low planting densities (833 and 500?sph). Linear mixed models revealed that branch diameter (BD) was negatively correlated with its relative height in the crown and IA. The binary-logistic regression model indicated that branch status (live or dead) was codetermined by BD and PD. These findings can help farmers cultivate high-quality wood of B. alnoides and obtain a higher economic return.  相似文献   

16.
对秃杉人工林开展不同造林密度和抚育间伐试验研究 ,结果表明 :随秃杉造林密度增加 ,其胸径、冠幅和整齐度减少 ,枝下高增大。单株生物量随林分密度增加而减少 ,林分生物量及树干和根系占全部生物量比率随密度增大而增加 ,枝条和针叶所占比率随密度增加而减少 ,根据培育目标 ,用下层间伐及中度间伐可切实有效地提高林分生物量和生长量  相似文献   

17.
Converting coppices into high forests with continuous cover has often been established during the last decades as a management goal in hilly and mountainous Mediterranean areas to attenuate the negative effects that frequent clearcutting may have on soil, landscape and biodiversity conservation. The silvicultural tool usually adopted for this purpose is the gradual thinning of sprouts during the long span of time required to complete the conversion, that also allows the owner to keep harvesting some wood. This research compared the effects of various thinning intensities (three treatments plus control) on the stand growth and structure of a beech coppice with standards. The optimal density after thinning was assessed by expressing mean tree spacing as a function of main stand attributes like stand height and stand dbh. This system was preferred to the empirical evaluation of the percentage of basal area to be removed in order to give forest managers general reference guidelines to adapt to the varying environments of the Mediterranean mountains. Results confirmed that the positive effects of thinning on mean stem volume is due more to the higher diameter increment than to different height growth. The acceleration of crown growth in the thinned plots allowed canopy closure to be achieved 13 years after thinning. This reduced the negative effects of the opening of the stand overlayer and the elimination of most suppressed trees on soil protection. Under the conditions examined, the best thinning intensity proved to be a stand density 20% lower than normal prescribed by the yield tables elaborated for beech high forests in Central and Southern Italy.  相似文献   

18.
An individual tree, process‐based stand growth model is presented. It is based on the carbon balance, according to which tree growth depends on the activities of photosynthesis, respiration and senescence. A simple model is specified for each component of the carbon balance. Next, equations for the tree structure, in which e.g. pipe‐model theory is utilized, are presented. The growth model for dry‐weights of tree compartments based on the carbon balance is transformed using these equations to allow the expression of growth in terms of diameter and height. It is also possible to aggregate a number of physiological and biometrical coefficients into a small number of generalized coefficients of the dimensional growth model. Additional components, including the equations for recession of the crown base and tree survival that are necessary for a stand growth model, are specified. Comparison of the stand growth model with a yield table and a growth model for a sapling stand suggests that the model is capable of accounting for the basic features of stand growth. Furthermore, simulations with varying initial density and some of the model's coefficients indicate that the stand growth model is approximately consistent with the so‐called self‐thinning rule.  相似文献   

19.
Precommercial thinning is regarded as one of the most important measures for influencing timber quality in stands of Scots pine (Pinus sylvestris L.). The present study considers the influence of both thinning intensity and tree height at the time of thinning on diameter of the thickest branch and crown ratio. A total of 115 plots (either naturally regenerated, planted or sown, and either thinned down to 528–8000 stems?ha?1 or untreated) included in 20 sites in southern Sweden were analysed. The average tree height after thinning varied from 1.2 to 8.3?m. An increasing number of remaining stems resulted in a reduction in branch diameter, although the reduction appeared to be only minor if the number of stems after thinning was more than 3000 stems?ha?1. It was found that late thinning reduced the diameter of the thickest branch. The crown ratio decreased with stand height, number of stems after thinning and average height at thinning. The results were consistent for all trees and for the 500 thickest trees per hectare.  相似文献   

20.
2010年6月在普洱良种繁育基地用20月龄的云南红豆杉(实生苗、顶枝扦插和侧枝扦插)容器苗定植,采用单因素随机区组设计,2011年12月对其生长量(树高、地径和侧枝发枝数)进行观测记录和结果分析。结果表明:3个处理的树高、地径和侧枝发枝数各处理间均存在极显著差异,树高、地径和侧枝发枝数生长表现最好的处理均是处理A即实生苗,树高和地径表现最差的均是处理C即侧枝扦插,侧枝发枝数表现最差的是处理B即顶枝扦插。  相似文献   

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