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1.
? If selective breeding is to be successful, significant genetic variation must be present in the traits targeted for improvement (i.e. “objective traits”). ? This study aimed to quantify genetic variation in Eucalyptus globulus pulpwood and sawn-timber objective traits (rotation-age whole-tree volume, survival, whole-tree basic density, sawn-board Janka hardness and sawn-board internal checking) and estimate additive genetic correlations between these and inexpensively-assessed “selection traits”. ? Significant genetic variation was identified in all objective traits at the subrace and/or family within subrace level. ? Selection-age diameter at breast height (1.3 m, DBH) was strongly genetically correlated with rotation-age volume (0.78) and survival (0.82). Subrace and additive genetic correlations of selectionage Pilodyn penetration with rotation-age 12-×-12-mm-sample basic density (?0.70 and ?0.75 respectively) and whole-tree basic density (?0.83 and ?0.91 respectively) were also strong. ? No significant subrace or additive genetic correlation between wood-sample gross shrinkage and sawn-board internal checking was detected. However, subrace and additive genetic correlations of sawn-board Janka hardness with Pilodyn penetration (?0.75 and ?0.58 respectively) and sample gross shrinkage (?0.77 and ?0.73 respectively) were significantly different from zero. ? These findings suggest that genetic improvement of the examined objective traits is possible through selective breeding, although none of the assessed selection traits were strongly correlated with internal checking. 相似文献
2.
Context The development of multiple trait selection indices for solid (structure) wood production in the Scots pine ( Pinus sylvestris L.) breeding program requires genetic variances and covariances estimated among wood quality traits including stiffness. Aims Genetic control and relationships among Scots pine growth, fiber, and wood quality traits were assessed by estimating heritability, phenotypic and genetic correlation using a Scots pine full-sib family trial. Method Wood quality traits including clearwood and dynamic acoustic stiffness were measured using SilviScan and Hitman in a 40-year-old progeny trial and by sampling increment cores of 778 trees of 120 families. Genetic parameters were estimated using the mixed model by the ASReml software. Results Heritability ranged from 0.147 to 0.306 for growth, earlywood, transition wood and latewood proportion traits and from 0.260 to 0.524 for fiber dimension, wood density, MFA and stiffness traits. The highly unfavorable genetic correlation between diameter and whole core density (?0.479) and clearwood stiffness (?0.506) and dynamic acoustic stiffness (?0.382) was observed in this study. Conclusion The unfavorable genetic correlations between growth traits and stiffness indicate that multiple traits selection using optimal economic weights and optimal breeding strategies are recommended for the advanced Scots pine breeding program. 相似文献
3.
? Each annual ring in pines consists of earlywood and latewood with considerable difference in density and width. To get a better determination of the genetic regulation of total wood density in Scots pine (Pinus sylvestris L.), density and width of those ring sections were measured in annual ring numbers 12 to 21 of Scots pines in a full-sib progeny test. Tree height and stem diameter were also measured. ? Heritabilities for the annual ring sections increased with age for earlywood density from 0.08 to approximately 0.25; latewood density showed similar reductions. Heritability over all 10 annual rings was 0.25 for earlywood density, 0.22 for latewood density, 0.29 for height and 0.10 for stem diameter. Genetic correlations between earlywood and latewood density and growth traits were negative, while they were strongly positive between densities of adjacent annual rings (0.70–1.0). ? Despite the higher heritability of earlywood density, the strong positive genetic correlation between those traits indicates little benefit from focusing solely on earlywood density when selecting for wood density. Analysing earlywood and latewood separately does not benefit from including the width of the corresponding ring section as a covariate. Juvenile wood may possibly turn into mature wood 15–20 y from the pith. 相似文献
4.
? Variation in wood basic density and its correlation with tree growth were investigated at 13 years in a provenance/progeny test of Prosopis africana in Niger. The test included progeny from 256 trees sampled from 24 provenances in the Sahelian ecozone of Burkina Faso and Niger. ? Variation in wood density was significant due to provenances and families within provenances. Individual tree heritability was higher for wood density than for growth traits. Provenance means for wood density increased from the more humid to the drier parts of the sample region. Phenotypic correlations indicated that larger trees tended to have denser wood. ? Clines suggest that natural populations of P. africana from the drier parts of the sample region have the genetic capacity to produce denser wood, compared with populations from the more humid parts of the region. Correlations and heritability estimates suggest that selection of faster-growing trees may produce a small gain in wood volume but little (if any) gain in density in the subsequent generation. Multi-location provenance/tests are needed to confirm this tentative conclusion. 相似文献
5.
? The aim of this work was to study both the effects of genetic entry and competition by neighbouring trees on growth, yield, wood density traits and fibre properties of 20 Norway spruce clones grown in an experimental trial located in southern Finland. The material included 10 Finnish clones, 2 Russian clones and 8 provenance-hybrids clones, the latter ones representing crosses between Finnish and foreign parent trees. ? We found that, compared to growth and yield, wood density traits and fibre properties showed, on average, lower phenotypic variations. Moreover, significant differences could be observed among the clones regardless of the trait (p < 0.05). Conversely, on average, no clear differences could be found between Finnish, Russian and provenance-hybrids clones in most of the studied traits. ? The marked differences among the clones, and lack of clear differences among the provenances suggested that any ranking, regarding different traits, should be based on individual clones. The phenotypic correlations between different traits ranged from moderate to strong suggesting that selection based on one trait would affect other traits. Regardless of clone, the growth of trees (e.g. breast height diameter) decreased and the wood density increased with increasing competition by neighbouring trees, which was expressed by competition index. 相似文献
6.
? Direct assessment of modulus of elasticity (MOE) on standing trees is attractive for breeders to evaluate genotypes prior to selection: this can be done using the Rigidimeter, a bending-based measurement device. ? In this study, we tested its reliability to properly rank genotypes by relating trunk MOE with MOEs estimated with a vibrating analysis system (Bing) on different types of conditioned wood specimens from the same trees (boards and standardised 2×2×30 cm-clear-wood specimens). One hundred and ten trees from different genotypes of hybrid larch (Larix × eurolepis) were tested. ? Mean trunk MOE was 7 300 MPa with a similar value obtained for sawn boards. Clear-wood specimens MOE increased from pith to bark from less than 6 000 MPa to nearly 9 000 MPa. Moderate correlations (r = 0.48–0.61) were found at the individual tree level between trunk MOE and MOE of wood samples. ? Single specimen MOE was shown to be strongly related to a linear combination of trunk MOE and sample position. ? At the genotype mean level, trunk MOE was highly correlated with wood samples MOE (r = 0.80–0.91). Ranking of genotypes based on trunk MOE was mostly consistent with that based on standardised specimens. ? It was concluded that besides other operational advantages which are discussed, the Rigidimeter is a valuable tool for breeders to routinely evaluate and rank genotypes for stiffness prior to further selection. 相似文献
7.
? The physical properties of wood and the associated variations within and between trees were evaluated by analysing 770 small specimens of clear wood from 11 Pinus sylvestris L. trees thinned from 3 plantations. ? Within-tree variations in basic density or volumetric shrinkage increased with cambial age and decreased with increasing ring width. The effect of the height in the stem on wood properties was considered indirect and height was not included as an explanatory variable in the mixed models proposed to estimate basic density and volumetric shrinkage. ? The models had random components for the intercept parameter and explained 52.5% of the total variance in basic density and 56% of the total variance in volumetric shrinkage. Linear shrinkage in the direction of the grain was extremely variable. ? Between-tree variation and between-plot variation in the physical properties of wood were high, considering that all trees sampled were growing in similar sites and stands. It would then be desirable to predict physical properties of wood on living trees in order to use the quality of wood as a criterion for timber tree selection in thinnings. 相似文献
9.
? Dimensional stability, along with the natural durability and colour of the wood, is one of the most important characteristics of teak used as timber. However, it is very time-consuming to take measurements of this kind. For the purposes of selection for the production of improved varieties, the number of samples to be measured rapidly exceeds the capacity of a traditional laboratory. ? Near-infrared spectroscopy, based on a set of reference data, is a tool enabling many of the chemical properties of wood to be predicted and the number of laboratory measurements to be reduced exponentially. The issue here is a question of checking the effectiveness of NIRS tool to build models and predict the shrinkage and fibre saturation point of teak wood from Togo. ? The results show the possible use of NIRS to measure the dimensional stability of teak wood and that it is appropriate to choose the type of wood and type of surface to be measured by NIRS. The best prediction models for radial and tangential shrinkage and fibre saturation point give R 2 values of 0.72, 0.83 and 0.87 respectively with ratios of performance deviation of 1.8, 2.4 and 2.8. ? Consequently, after verification on other sets of teak samples, which may or may not be included in the prediction model, NIRS can be used to predict shrinkage and fibre saturation point values accurately for a large number of samples, making it possible to include these characteristics in the selection criteria for classifying wood and high throughput phenotyping. 相似文献
10.
? Sawn wood which acquires twist during drying is a substantial problem for the sawmill industry and is to a large extent caused by spiralled grain in the tree. ? In this study, the feasibility of using spiral grain angles as a selection trait in Norway spruce (Picea abies (L.) Karst) tree breeding and the extent of grain angle and its breeding values were investigated. Grain angles under bark and diameter were measured at breast height in three 28-year-old Norway spruce progeny trials and in their parent seed orchard. ? The mean grain angle value was 1.76° in the progeny trials and 1.84° in the seed orchard. Estimated genetic standard deviations for grain angle were almost 1°. Progeny trial heritabilities (H 2 > 0.3) were larger than the seed orchard heritability (H 2 = 0.24). Genotype by environment interaction among the progeny trials was negligible for grain angle. Progeny trial grain angles exhibited genetic correlation with seed orchard grain angles in the range 0.66 to 0.84. ? Thus, mature wood grain angles are also under considerable genetic control, raising the opportunity to reduce the occurence of left-grained trees, which are prone to producing twisted wood. 相似文献
11.
? Eight-year old trees from two Eucalyptus globulus Labill. clones planted across three different sites in Tasmania, Australia, were sampled for wood and kraft pulp/handsheet properties. ? Site had a significant effect on all measured properties. Compared with the poor site (Parkham) the wood from the good site (West Ridgley) had on average 11 % lower wood density. The poor site had also greater microfibril angles, shorter fibres at lower pulp yields. ? The handsheets produced with pulp from the poor site resulted in comparatively higher bulkiness, lower burst, lower tear and tensile indices, lower zero span tensile strength, but higher opacity, higher light scattering and higher surface roughness. Significant height effects were found with all wood properties, and also with tear index, zero span tensile strength and opacity. ? Discriminant analysis showed that for 76 out of 100 handsheets the raw material source, i.e. growth site, could be predicted correctly using a set of handsheet properties with tear index and bulk index being most prominent. ? This is unique evidence that site conditions are strongly reflected in handsheet properties produced from Eucalyptus pulp. 相似文献
12.
? Introduction There has been an increasing interest in very early selection of radiata pine to reduce the breeding cycle for solid wood products. For such selection, new approaches are required to assess wood quality in wood from very young stems. ? Methods Nursery seedlings of clones of radiata pine were grown in leant condition using two leaning strategies for 18–20 months. Opposite wood and compression wood were isolated from the leaning stems and tested for dynamic modulus of elasticity, density, longitudinal shrinkage, volumetric shrinkage and compression wood area using new methods evolved for testing small size samples quickly and reliably. The methods were tested for their efficiency in differentiating clones by their wood properties. ? Results Leaning of stems provided distinct opposite and compression wood for testing. Automated image analysis method used for compression wood area assessment was found to be a quick and effective method for processing large number of samples from young stems. Compression wood was characterised by high basic density, high longitudinal shrinkage and low volumetric shrinkage than that of opposite wood. Acoustic velocity in opposite wood had a strong negative association with longitudinal shrinkage. The study signifies the importance of preventing mixing of opposite wood with compression wood while assessing wood quality in young stems thus making leaning a critical strategy. The comparison of wood properties of opposite wood revealed significant differences between clones. Opposite wood of the clone with the lowest dynamic modulus of elasticity exhibited the highest longitudinal shrinkage. ? Conclusion Significant differences in measurable wood properties between clones suggest the prospects of early selection for solid wood products. 相似文献
13.
? In forests, the substitution of broadleaf species by conifers can reduce biodiversity because coniferous forests generally provide less diverse vascular understories than broadleaf forests. However, in some cases, former pine plantations might be useful for restoring native forests. We compared plant species composition on the plot scale in natural beech and mixed oak forests with that in plantations of Pinus radiata. Links between plant diversity and landscape parameters (patch size, fractal dimension and distance to the nearest patch of the same type) were investigated. ? The objective of this study was to evaluate the use of pine plantations for restoring native diversity in a zone where native forests are very fragmented. ? Similar to oak forests, plant diversity in pine plantations was high, mainly due to the presence of generalist species. Some species characteristic of oak forests also appeared in pine plantations, suggesting the onset of natural forest regeneration. ? These results suggest that pine plantations could be used to promote natural regeneration of original oak forests. Moreover, residual native stands should be conserved as important sources of native species and their seeds. 相似文献
14.
- ? It is essential to understand how characteristics are related to each other in breeding programmes to select wood properties, in order to avoid that, in selecting for one trait, we are negatively affecting another. Moreover, measuring wood properties is time consuming and expensive.
- ? This study assesses genetic and phenotypic correlations between wood density components and spiral grain of 46 half-sib families of Pinus pinaster in seventeen-year-old trees.
- ? Results showed that genetic correlations for all wood density components were higher than corresponding phenotypic correlations. Furthermore, all wood density components were highly genetically correlated with ring density, and also closely associated among themselves. Results showed a higher genetic correlation of ring density with earlywood density (r g = 0.96) than with latewood density r g = 0.79). A moderate to high positive genetic correlation was found between spiral grain and wood density characteristics (0.29–0.61).
- ? We conclude that ring density (overall wood density) can be improved by increasing either earlywood density, latewood percent, or both of these traits, and spiral grain can be modify in future plantations.
相似文献
15.
? Awareness of the shortage of fossil resources leads to an increasing demand for woody biomass. We investigated the feasibility of using fast-growing Gmelina arborea wood for material production. Gmelina arborea wood samples were collected from trees of varying cambium ages in Indonesia, from 3.5-, 7- and 12-year-old plantations. ? The lateral growth rate and the cambium age did not significantly affect the longitudinal released strain of the growth stress, xylem density, or microfibril angle at the outermost surface of the secondary xylem at any sampling site. However, fiber length in the 3.5-year-old plantation tended to be shorter in smaller diameter trees, whereas in larger diameter trees it was almost the same as that in trees from the 7- and 12-year-old plantations. This suggests that smaller diameter trees in the 3.5-year-old plantation had not yet produced mature wood. ? Xylem qualities had already reached values appropriate for harvesting, except in the smaller diameter trees from the 3.5-year-old plantation. This indicates that the larger diameter trees had already matured, regardless of their cambium age. These results suggest that the next step is to develop silvicultural treatments to increase the lateral growth rate during the early growing stage, in order to produce as much mature wood as possible, as quickly as possible. 相似文献
16.
? The objective was to optimise the strategy for maintaining the genetic balance in long-term breeding where the grandparents contribute equally to the breeding population, but the parents do not. ? The annual genetic progress under a budget constraint was compared for a number of scenarios. The factors considered were: genetic parameters, recruitment population size, mating number per grandparent, cost of plants and parents, selection age and time components. Phenotypic selection forward was assumed. Using more parents than grandparents affects the testing population only, thus, gene diversity loss and breeding population size remain constant. ? Using larger number of parents than grandparents was a superior strategy at all scenarios tested. The strategy with 6 parents per grandparent and 1% of the tested plants selected on their phenotype at age 15 for further mating is recommendable. The strategy to choose 6 parents per grandparent was robust over a heritability range 0.05 to 0.2. At a higher heritability, the optimum number of parents per grandparent was 2 to 4. ? Using larger number of parents than grandparents has the potential to improve annual genetic gain in the magnitude of 50% compared to the strategies currently used for Scots pine in Sweden. 相似文献
17.
? Environmental determinants of wood properties variation were examined in Eucalyptus globulus, a globally important hardwood plantation species, in southern Tasmania, Australia. ? Radial variation in wood properties, measured with the SilviScan system, were re-scaled from distance to time abscissa using stem radial growth data measured with dendrometers. With this re-scaled data it was possible to evaluate how water availability and temperature affected wood density, microfibril angle (MFA) and fibre and vessel transverse dimensions in irrigated and non-irrigated trees. ? Wood density, fibre radial diameter and MFA were sensitive to water availability. Wood density increased and fibre radial diameter decreased in response to reduced water availability. When high water availability was maintained, wood density was negatively correlated with temperature. Together, temperature and soil matric potential explained about 60% of temporal variation in wood density variation. In contrast MFA was not related to temperature but decreased with increasing water stress. Slower growing trees also had lower MFA than faster growing trees. Slower growing trees had a larger number of vessels per unit area of wood than faster growing trees within this even aged stand. However, vessel radius to the 4th power was significantly higher in faster growing trees than in slower growing trees. ? Overall, E. globulus wood properties were sensitive to temporal changes in environmental conditions (particularly water availability) and associated growth rates. The data provided support for the hypothesis that growth rates are hydraulically mediated. 相似文献
18.
? Wood used in outdoor conditions out of ground contact is susceptible to weathering, inducing both fungal decay and leaching of components to the environment. ? This paper presents a methodology to determine these two parameters for untreated, preservative-treated and modified wood. Therefore, the wood was first leached and subsequently exposed to fungal decay of the most prominent wood-rotting fungi. The crustacean Daphnia magna was exposed to the leachates to provide information on their impact on the environment. ? Combining both parameters reveals that preservative-treated wood and modified wood are capable of protecting the wood adequately for application under use class 3 conditions without posing a threat to the environment. ? This proves the suitability of the concept of combining efficacy and ecotoxicology for the evaluation of new types of wood treatments. 相似文献
19.
? The aim of this study was to investigate the effectiveness of using cellulose content, measured by the diglyme-HCl method, as a selection trait in breeding programs for kraft pulp yield in Eucalyptus urophylla. ? A total of 275 trees from sixty-two families were sampled from a thinned progeny trial of E. urophylla in northern Vietnam to evaluate cellulose content from breast-height increment cores. Among those, twenty unrelated trees were felled to evaluate cellulose content and pulp yield from breast-height disk samples. ? The regression of pulp yield of disk samples on cellulose content was strong either from disks (R 2 = 0.83) or increment cores (R 2 = 0.69). There was no significant difference in cellulose content between the provenances. The narrow-sense within-provenance heritability of cellulose content was 0.50 and the coefficient of additive genetic variation was 3.9%. Genetic correlations between cellulose content and growth (0.28–0.45) or wood basic density (?0.02) were not significantly different from zero. ? Breast-height increment core cellulose content measured by diglyme-HCl method is under strong genetic control and can be used to rank trees for pulp yield in E. urophylla plantations. Selection for increased cellulose content would have only minor effects on growth and wood basic density. 相似文献
20.
? Decay resistance of larch (Larix sp.) to fungi was evaluated on heartwood samples belonging to 3 species (L. decidua, L. kaempferi and their hybrid), 3 races of European larch (polonica, sudetica and alpine), 13 wood lots (populations) and 313 trees. ? Larch wood appeared, on average, as moderately durable although a high variability was observed. At the sample level as well as at the mean individual tree level, durability ranged from class 1 to 5 according to EN 350-1 standard. At the population level, larch wood varied from ‘durable’ to ‘slightly durable’. Genetics played a major role in decay resistance at the species, provenance and tree levels. Environmental factors such as the position of heartwood samples and the age of trees were also identified as a source of variability. ? The most durable wood was not necessarily from old native alpine stands of European larch: some young larches from faster growing lowland origins also produced durable wood. ? Genetic improvement of larch wood durability appeared therefore likely by the selection of the best populations for decay resistance as well as from the selection of individuals. 相似文献
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