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1.
We examined the short-term effects of group-selection harvestingwith seed tree retention on the diversity, abundance and establishmentof tree seedlings in a northern hardwood forest in the UpperPeninsula of Michigan (49 openings, 20 closed canopy referencesites). Three opening sizes were examined – opening radius0.5 x canopy height (267 ± 62 m2, n = 16), 0.75 x canopyheight (642 ± 85 m2, n = 17) and 1.0 x canopy height(1192 ± 155 m2, n = 16) (canopy height = 22 m). A singleyellow birch (Betula alleghaniensis Britt.) seed tree was retainedin the centre of each opening. Tree seedling density was significantlyhigher in the largest group-selection opening than at the closedcanopy reference sites (P < 0.05), the main factor for thiswas the increased proportion of yellow birch, red maple (Acerrubrum L.) and other minor species. Nevertheless, yellow birchwas still a minor component of the developing gap cohort, comprising5.9 per cent of the seedlings and 1.1 per cent of the saplings.Within openings, microsite variables, such as per cent coversof bare soil and coarse woody debris, were the best predictorsof yellow birch occurrence and density. Our results suggestthat microsite limitations and competing vegetation may greatlyreduce the efficacy of openings for ensuring the maintenanceof mid-tolerants. 相似文献
2.
Response to canopy opening does not act as a filter to Fagus sylvatica and Acer sp. advance regeneration in a mixed temperate forest 总被引：1，自引：0，他引：1
Blandine Caquet Pierre Montpied Erwin Dreyer Daniel Epron Catherine Collet 《Annals of Forest Science》2010,67(1):105-105
3.
Beech woodland can be restored by direct planting of beech (Fagussylvatica L.) seedlings in abandoned areas, but this methodis generally avoided in forestry because of the growth difficultiesof beech in full-light conditions. This study tested a methodthat consists of planting beech seedlings in full-light conditionswith silver birch (Betula pendula Roth.) or Scots pine (Pinussylvestris L.) as nurse trees. A total of 65 two-year-old beechseedlings surrounded by either 3, 4, 5 or 6 pine or birch seedlingsor without competitors were planted in bare-soil open-fieldconditions in Central France. Tree growth and light availabilitywere monitored at the end of each of the following 3 years,and soil moisture was also measured the last year. At the endof the experiment, aerial biomass measurements were made onthe beeches. Results showed that relative beech growth in termsof diameter was significantly reduced by the local neighbouringtrees and that this reduction was particularly noticeable inthe pine treatments where there was extensive pine seedlingdevelopment. In contrast, relative beech height did not exhibitany significant variations among the treatments. Light availabilitywas decreased by the neighbouring trees, especially in the pinetreatments. Light reduction was more pronounced in the last2 years and in the middle or lower parts of the canopy of thesubject beech. Soil water content was lower under the pine canopiesthan under the birch canopies, and was positively correlatedto beech relative diameter growth. Specific leaf area, height-to-diameterratio and crown length-to-crown width ratio weakly but significantlyincreased with competition from the neighbours. Beech biomasspartitioning was only weakly affected by the treatments. Thisstudy showed that nurse trees tend to reduce beech growth butimprove form although effects remained weak due to the insufficientlength of the experiment. Further studies are also needed toquantify the effects of the neighbourhood on the growth of competitiveherbaceous vegetation and on changes in the microclimatic conditions. 相似文献
4.
A plantation of lodgepole pine (Pinus contorta Dougl.), whichwas established in 1973 on a ditched minerotrophic fen at plantingspacings of 1.2 m, 1.8 m, and 2.4 m, was refertilized in 1985with control, P, PK and NPK treatments consisting of 60 kg ha–1phosphorus, 100 kg ha–1 potassium and 200 kg ha–1nitrogen. Although each seedling had been spot-fertilized in1973 with a mixture of 71 g urea, 99 g ground rock phosphateand 41 g potassium sulphate, height and needle nutrient assessmentsof the plantation in 1984 indicated poor growth, low concentrationsof needle nitrogen (1.26 per cent) and possible deficiency levelsof needle phosphorus (0.10 per cent) and potassium (0.36 percent). Between 1985 and 1991, height and diameter growth increasedsignificantly in all refertilization treatments. Growth responsewas greatest in the PK treatment and least in the P treatment.Needle weights, determined from 1987 to 1991, did not responduniformly to any of the refertilization treatments. Needle concentrationsof phosphorus increased to levels between 0.20 per cent and0.23 per cent following refertilization with P, PK, and NPK,but by 1988 had decreased to about 0.14 per cent. Needle concentrationsof potassium also increased sharply to 0.60 per cent in 1985following refertilization with PK and NPK, but by 1988 had decreasedto levels between 0.35 per cent and 0.45 per cent. Similarly,needle nitrogen concentrations increased to 1.84 per cent in1985 following refertilization with NPK, then decreased to levelsbetween 1.20 per cent and 1.33 per cent in 1986 and remainedat those levels each year thereafter. Although needle nutrientlevels fell sharply following an initial peak after refertilization,height and diameter growth did not decrease significantly, especiallyin the PK treatment, suggesting that nutrient levels remainedadequate for optimum growth. 相似文献
5.
The effects of thinning were investigated on naturally regeneratedFraxinus rhynchophylla Hance (Korean ash) grown in the understoreyof a Larix kaempferi (Lamb.) Carr. plantation, established ona site that previously supported old growth hardwood forests.The regeneration characteristics were evaluated after thinningof the plantation (about 60 per cent of relative density wasremoved). In the naturally regenerated and mixed stands, Quercusmongolica Fisch. was the most abundant tree species in the overstorey,but the understorey of the thinned plantations was dominatedby F. rhynchophylla with six other minor tree species. Fraxinusrhynchophylla was regenerated by four propagation types includingnatural seedling, stump sprout, root sprout and layering. Beforethinning, the difference of average height growth for each regenerationtype was insignificant. One year after thinning, the stump sproutsshowed the fastest growth among the regeneration types. Heightgrowth of advance regeneration was slow in the year followingthinning but then increased rapidly surpassing the other regenerationtypes 5 years after thinning. Thinning had an effect on thecomposition of regenerated trees also by seedling, stump sprout,root sprout and layering in the understorey. Advance regenerationsoccupied the upper understorey at 5 years after thinning. Alarge number of stump sprouts emerged in the plots 1 year afterthinning, while at 5 years after thinning the occupation ofstump sprouts decreased. Root sprouts and layers faded away.Considering the early growth and the composition of understoreyF. rhynchophylla, its advance regeneration has a high potentialto develop into overstorey trees at this site. 相似文献
6.
Little is known about broadleaf establishment within the nativepinewoods of Scotland, yet such information is critical to currentrestoration initiatives. A 2-year field trial was set up toassess the growth and performance of seedlings of four broadleavedspecies within different developmental stages of Pinus sylvestris(Scots pine) woodland in the Scottish Highlands. The developmentalstages included stem initiation (establishment stage), stemexclusion (thicket/pole stage) and canopy break up (old-growthcanopy), providing sites of varying shade and resource availability.It was predicted that differences in establishment between siteswould occur due to species-specific differences in ‘tolerance’of shade, low-nutrient availability and poor soil drainage.Analysis of trial results showed high survival but growth rates(relative stem height and basal diameter growth) declined rapidlyin year two. In year two, Ilex aquifolium achieved the highestmean growth rates in the stem exclusion stands (average 15 percent light) where there was a lower percentage of Phytomyzailicis (holly leaf miner) infection and frost damage. Sorbusaucuparia exhibited generalist behaviour with similar growthand performance across all stand treatments. Betula pubescensmaintained similar growth rates across stand types but was moresusceptible to insect damage in deeper shade. Overall, Alnusglutinosa grew more successfully than the other species andachieved highest mean growth rates in canopy break up (average44 per cent light) and stand initiation (average 71 per centlight) stages. Broadleaf establishment within pinewood standsis challenging on account of acidic nutrient-poor soil regimes,and in places, poor drainage, aggressive understorey competitionand canopies that restrict light availability and insect herbivory.Forest restoration techniques, including matching species tosite, accelerating establishment and using framework species(individuals of high field performance that are able to restoresite productivity and nutrient cycles), are discussed. 相似文献
7.
The influences of cambial age and ring width on density of Sitkaspruce (Picea sitchensis (Bong.) Carr) were analysed in relationto within-tree trends in tracheid diameter and cell wall thickness.Discs were sampled at breast height from a total of 24 trees,from seven stands at three contrasting sites in Wales, and atbreast height, 30 per cent and 60 per cent total tree heightfrom one of the stands. Across the juvenile wood, ring density decreased with ring numberfrom the pith while radial tracheid diameter increased. Theseoverall trends were considered to be inherent to tree growth,presumably associated with cambial ageing, since they occurredin all trees on all sites. In juvenile wood, density also variedwith site growth rate (as indicated by ring width) at similarcambial age, wider rings being associated with more rapidrateof change in tracheid diameter with ring number and with decreasein tracheid wall thickness. Consequently, on a site having treeswith high growth rate density decreased more rapidly acrossthe juvenile wood, down to a lower minimum value, than on siteswith a slower growth rate. In mature wood, the decrease in densitywith increase in ring width was associated with differencesin both tracheid diameter and wall thickness. Density was slightly(though not significantly) higher at breast height than in comparablerings at 30 per cent total height, associated with significantlythicker tracheid walls at breast height. Changes in radial tracheid diameter (with ring number, or withring width) were associated with greater differences in theearlywood than towards the latewood end of each growth ring,while variations in wall thickness with ring width were associatedwith rate of increase in wall thickness towards the latewoodend. This may account for some previously conflicting reportson influence of silvicultural management on density, for densityis likely to vary with influence of environment on the seasonalcycle of cambial activity. The extent of the juvenile wood as delimited by the inner coreof wide growth rings does not necessarily correspond to theregion of varying tracheid dimensions in Sitka spruce. 相似文献
8.
Ontogenetic variation in the relative influence of light and belowground resources on European beech seedling growth 总被引：1，自引：0，他引：1
We used height growth data from a 7-year field experiment with European beech (Fagus sylvatica L.) seedlings to test the hypothesis that the effects of above- and belowground resources on height growth depend on seedling size and age. Photosynthetically active radiation (PAR) was determined by hemispheric photography, and estimates of fine root biomass of the overstory trees were used as an inverse proportional surrogate for belowground resource availability. For recently germinated seedlings growing under the canopy of Picea abies (L.) Karst., belowground resource availability affected height growth more than light. During subsequent stages of seedling development, apart from initial seedling size, PAR increasingly determined seedling growth. Besides initial size, seedling age determined the effects of above- and belowground resources on seedling height growth. In seedlings identical in initial size but differing in age, the increase in height growth with increasing PAR was greater in older seedlings than in younger seedlings. The ranking of seedling height by year showed that small differences in size at the end of the first growing season resulted in continuously increasing differences during the following years. Mortality data indicated that the chances of a seedling surviving intraspecific competition was strongly determined by its dominance ranking within the first 5 years after establishment. 相似文献
9.
Characteristics of lowland broadleaved woodland being restocked by natural regeneration 总被引：3，自引：0，他引：3
A survey of 78 sites in southern England with approved managementplans for restocking by natural regeneration was made duringthe summers of 1993 and 1994. The following features were recorded:species, stocking, canopy cover and seed-bearing potential oftrees present in the overstorey; species and canopy cover ofthe understorey; ground cover; species, browsing damage, numberand heights of tree seedlings. In general, sites were poorlystocked with overstorey trees having an average of 135 stemsha–1 and a mean canopy cover of 36 per cent. Similarly,the understoreys were poorly developed with an average coverof only 23 per cent. Twenty-nine species of tree were foundin the overstorey, Quercus spp. and Fraxinus excelsior werethe most common. Many of the stems present had poor seed-bearingpotential. Cover of the ground flora often exceeded 75 per cent.Seedlings were present on most sites, with F. excelsior andBetula pendula being most abundant with mean seedling densitiesexceeding 10000 ha–1. Most seedlings were >20 cm tall,few exceeded 120 cm. About 30 per cent of seedlings were browsed.The results are related to current guidance and the future prospectsfor use of natural regeneration. 相似文献
10.
Post-thinning stump sprout response was assessed in a coast redwood [Sequoia sempervirens (D. Don) Endl.] stand from 5 to 9 years after five treatments were initiated. Nine years after treatment, leaf area index
(LAI) ranged from 5.9 to 14.1 and the percent above canopy light (PACL) ranged from 0.8 to 3.9. Sprout responses included
rapid self-thinning in all treatments with complete sprout mortality at low light regimes. Self-thinning of clumps increased
with greater overstory LAI and poorer light regimes. Leaf area of individual sprout clumps also declined in low light regimes.
The probability of sprout survival declined rapidly below 10 PACL and coincided with declining leaf area in sprout clumps.
A conceptual model is presented that shows the sensitivity of redwood sprouts to light regime. Results have implications for
management of multiaged stands and indicate these redwood coppice systems require relatively severe overstory treatments to
provide sufficient light for sprout vigor and growth. 相似文献
11.
Field experiments involving naturally occurring population levelsof larval European pine sawfly Neodiprion sertifer (Geoff. )and artificial defoliation in both field and glasshouse experimentswere used to investigate physiological effects of defoliationon lodgepole pine. Reductions in total projected leaf area,annual height and volume increments of 19, 33 and 32 per cent,respectively, were recorded on trees where larvae had consumedall mature foliage. Leader growth was up to 48 per cent lesson defoliated trees after two years reflecting a delayed growthimpact. Similar effects were found on trees which had been defoliatedby hand in a manner resembling sawfly feeding. Artificial defoliationof transplants also showed that this type of damage can haveserious consequences on below-ground growth, particularly offine roots. Root:shoot ratios were found to be 0.64 in undamagedplants and 1. 05 in defoliated plants. A high concentration(1300 ppm) of the insecticide fenitrothion, used to maintaincontrol treatments in the field, was found to have no significanteffects on the major growth parameters being assessed. 相似文献
12.
Tabari Masoud; Fayaz Payam; Espahbodi Kambiz; Staelens Jeroen; Nachtergale Lieven 《Forestry》2005,78(4):443-450
The response of nursery seedlings and wildings of oriental beech(Fagus orientalis Lipsky) to canopy gap size was studied ina climax beech forest in northern Iran with respect to seedlingsurvival, shoot growth and vitality. These parameters were followedfor 2 years after planting in gaps of 50, 200 and 600 m2, aswell as in the open field. Foliage coloration was used as acriterion for vitality. The survival rate of nursery seedlingsand wildings dropped rapidly with increasing gap size from 84per cent in 50-m2 canopy openings to 相似文献
13.
Development of water oak stump sprouts under a partial overstory 总被引：2，自引：0，他引：2
A 28-year-old water oak (Quercus nigra L.) plantation was thinned from below to either 254 or 462 stems per hectare to determine the influence of a partial canopy on oak stump sprout development. Sprout clump survival, number of living sprouts in a clump, and height and DBH of the dominant sprout in a clump were measured in years 1–5 and 7 after harvest. By year 7, sprout clump survival under the heavily thinned canopy was 23% higher than under the lightly thinned canopy. Within-clump sprout mortality was not influenced by overstory thinning level, but by year 2 height and diameter increment were 15% and 22% greater, respectively, under the heavy overstory removal. Positive height and diameter growth of dominant sprouts continued under both canopy conditions through year 7, but early benefits of the heavy overstory removal on sprout growth diminished. Additional overstory removal or sprout clump thinning would be necessary to sustain sprout clump survival and sprout growth over extended periods. 相似文献
14.
Growth of regenerating trees in different light environments was studied for the mountainous, mixed-species forests in the Carpathian Mountains of Romania. The primary species in these mixtures were silver fir (Abies alba Mill.), European beech (Fagus sylvatica L.) and Norway spruce (Picea abies (L.) Karst). Seedlings/saplings of these species were selected and measured in different stands from two different geographical locations. Regenerating trees were measured for height and diameter growth during the summer of 2002. For each seedling/sapling, percentage of above canopy light (PACL) and stand basal area (BA) were used to assess available and occupied growing space respectively. Regeneration growth was compared against these two variables and regression relationships were developed. Using these models, we predicted the dynamics of regeneration as both growth and species composition. Our results showed that in low-light environments (PACL<20–35%; BA>30 m2/ha), shade tolerant fir and beech clearly outcompeted the spruce. Therefore, in dense stands, spruce could be eliminated by the shade tolerant species. For intermediate levels of cover (PACL=35–70%; BA=15–35 m2/ha) the spruce grew at comparable rates as the beech and fir. All three species showed similar growth rates in open conditions (PACL>80–90%; BA<15–20 m2/ha) with the spruce having a tendency to outgrow the others. However, in terms of establishment, such conditions favor spruce and inhibit fir and beech. 相似文献
15.
We tested the effects of exponential nutrient loading and springtime carbon loading during nursery culture on the field performance
of black spruce (Picea mariana (Mill.) B.S.P.). Seedlings were grown from seed with a conventional, fixed dose fertilizer (10 mg N seedling−1) or an exponential nutrient loading regime (75 mg N seedling−1). The following spring, seedlings were exposed for two weeks to either ambient (370 ppm) or elevated levels of CO2 (800 ppm) and then planted in the field; seedling growth was followed for the next six years. Exponential nutrient loading
increased seedling height, stem diameter and leader growth, with the largest increases in height and leader length occurring
in the first three years after outplanting. Carbon loading increased seedling height and leader length, but only in seedlings
that had been exponentially nutrient loaded. A combination of carbon and nutrient loading increased shoot height 26%, stem
diameter 37% and leader length 40% over trees that received neither treatment. These results demonstrate that the growth enhancement
seen under exponential nutrient loading is maintained under field conditions for at least six years. Carbon loading just before
outplanting was a useful supplement to nutrient loading, but was ineffective in the absence of nutrient loading. 相似文献
16.
Henrik Hartmann Marilou Beaudet Marc J. Mazerolle Christian Messier 《Forest Ecology and Management》2009
In this study, we quantified the effects of local neighbourhood competition, light availability, and proximity to skid trails on the growth of sugar maple (Acer saccharum Marsh.) trees following selection harvest. We hypothesized that growth would increase with decreasing competition and increasing light availability, but that proximity to skid trails would negatively affect growth. A total of 300 sugar maples were sampled ∼10 years after selection harvesting in 18 stands in Témiscamingue (Québec, Canada). Detailed tree and skid trail maps were obtained in one 0.4 ha plot per stand. Square-root transformed radial growth data were fitted to a linear mixed model that included tree diameter, crown position, a neighbourhood competition index, light availability (estimated using the SORTIE light model), and distance to the nearest skid trail as explanatory variables. We considered various distance-dependent or -independent indices based on neighbourhood radii ranging from 6 to 12 m. The competition index that provided the best fit to the data was a distance-dependent index computed in a 6 m search radius, but a distance-independent version of the competition index provided an almost equivalent fit to data. Models corresponding to all combinations of main effects were fit to data using maximum likelihood, and weighted averages of parameter estimates were obtained using multimodel inference. All predictors had an influence on growth, with the exception of light. Radial growth decreased with increasing tree diameter, level of competition and proximity to skid trails, and varied among crown positions with trees in suppressed and intermediate positions having lower growth rates than codominants and dominants. Our results indicate that in selection managed stands, the radial growth of sugar maple trees depends on competition from close (≤6 m) conspecific neighbours, and is still affected by proximity to skid trails ∼10 years after harvesting. Such results underscore the importance of minimizing the extent of skid trail networks by careful pre-harvest planning of trail layout. We also conclude that the impact of heterogeneity among individual-tree neighbourhoods, such as those resulting from alternative spatial patterns of harvest, can usefully be integrated into models of post-harvest tree growth. 相似文献
17.
This study explores the indirect relationship between forest structural measures and initial seedling survival and growth
along a structural gradient between 64% to 92% canopy closure. The gradient was created by applying various levels of midstory
removal to fifty 0.05 ha areas located within a mixed-hardwood riparian forest corridor. Twelve yellow-poplar (Liriodendron tulipifera L.) and cherrybark oak (Quercus pagoda Raf.) containerized seedling pairs were underplanted within each area. Canopy closure was estimated using hemispherical photography;
height-to-canopy and basal area were recorded at each seedling pair. Survival, basal diameter, and height were monitored through
two growing seasons. Species-specific mortality and height growth models were developed for one and two growing seasons following
underplanting. The interaction of height-to-canopy and basal area along with canopy closure were found to be the most strongly
related to mortality. Height to the forest canopy and initial seedling size explained the most variance in height increment.
Although the height increment models possess limited predictive power (R2 range from 0.22 to 0.36), both mortality and growth analyses emphasize the importance of quantifying vertical canopy structure,
along with the more commonly considered horizontal measures of forest structure (basal area and stem density), when evaluating
seedling development beneath a forest canopy. 相似文献
18.
We compared shade tolerance of maple, ash and beech in the saplingstage from two sites with rich soils differing in water supply,growing in dense thickets underneath a beech shelterwood ofvarying canopy densities. Shade tolerance was described by twocomponents: mortality in shade and height growth in high light.At low light, beech showed the least mortality, maple the highestand ash in between on both sites. The decline with increasinglight was steepest in beech and more gradual with ash and maple.At 15 per cent above canopy light, all three species approachedzero mortality. Beech as the most shade-tolerant species hadthe highest survival rate under low light and the least lengthgrowth rate under high light (>17 per cent). Ash had a lowersurvival rate at low light than beech and a highest growth rateat high light. Maple showed a bit weaker trade-off with thelowest survival rate but a growth rate inferior to ash. On thebetter water-supplied site, height growth was significantlysuperior in all three species only under high light. On thebasis of these results, silvicultural conclusions are drawnwith respect to appropriate light levels and cutting types. 相似文献
19.
Yang Yonghui; Watanabe Masataka; Li Fadong; Zhang Jiqun; Zhang Wanjun; Zhai Jianwen 《Forestry》2006,79(1):135-147
To estimate the possible effects of site factors and climatechange on forest growth in the Taihang Mountains, northern China,we assessed the factors influencing forest growth by using forestinventory data from 712 forest sample plots. Meteorologicaldata from 77 meteorological stations in the region were usedto estimate temperature and precipitation at each site fromelevation and longitude. Analyses showed that temperature, aspect,precipitation and soil thickness all significantly influencedforest growing stock (FGS), i.e. stem volume. When temperaturerose, FGS was reduced, possibly because increasing temperatureincreased evapotranspiration. Precipitation had a positive effecton FGS. The effect of aspect on FGS was perfectly expressedas a cosine function, with south-west- and south-facing slopeshaving the lowest FGS and north-facing slopes having the highest.We developed multifactorial regression models to predict changesin FGS in the Taihang Mountains. Temperature, forest age, forestcover, soil thickness, precipitation and aspect were well relatedto FGS. The effects of a temperature decrease and a precipitationincrease on FGS would be 2.5–8 per cent per degree centigradeand 10 per cent per 100 mm, respectively. The combination oftemperature increase and precipitation changes under futureclimate change is likely to result in a decrease of FGS, thoughthis does not take account the effect of increasing CO2. Wealso used multifactorial regression models to analyse the effectsof site factors on FGS of Pinus tabulaeformis Carr. and Robiniapseudoacacia L., two major species used in afforestation inthe Taihang Mountains. Although site factors had similar effectson FGS, diameter at breast height and tree height of both species,prediction accuracy (regression coefficient) was improved greatlywhen we treated the species separately. 相似文献
20.
The study evaluates the establishment performance of four treespecies (Ficus carica, Fraxinus syriaca, Salix acmophylla, Platanusorientalis) 3 years after planting in the peatland in the HulehValley. To develop good tree establishment and avoid uprooting,two types of soils were tested as alternatives for the rhizosphere.For each tree 6 m3 of soil (2 x 2 x 1.5 m) were replaced withriver bed alluvium or terra-rossa, in which the tree was planted.Control trees were planted directly in the peat. Changing therhizosphere soil type to river bed alluvium significantly improvedthe establishment and performance of Salix acmophylla and Fraxinussyriaca (survival, height, diameter and canopy shade area).Ficus carica was not affected by changing the soil in the rhizosphere,while Platanus orientalis was affected mainly in canopy shadearea. Terra-rossa improved establishment only of Salix acmophylla;it did not affect the other species. 相似文献