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1.
We investigated the selectivity of cod (Gadus morhua) and haddock (Melanogrammus aeglefinus) escaping from three different selection systems (a conventional diamond-mesh codend, a codend fitted with escape windows, and a codend fitted with a sorting grid) using the covered codend technique. By using a MultiSampler attached to the codend cover to partition the collection of escaping fish, we were able to determine the partitioned selectivity of each of these devices at different phases during the towing process: at depth, during the haul back, and at the surface.The percentages of escaping fish at depth from the diamond-mesh codend were significantly lower than those estimated for the escape windows and sorting grid codends: almost 62% of cod and 79% of haddock escaping from the diamond-mesh codend did so at depth, whereas approximately 75% of cod and 93% of haddock escaped from the other two systems at depth. The partitioned selectivity parameters, l50 and SR, were estimated and modelled for each of the three phases of the towing process and used to compare the selective properties of the three selection systems. A comparison was also made with a traditional whole haul selectivity analysis. Finally, bulk catch was added to the model as an explanatory variable and its effect upon selectivity was measured in both, partitioned and whole haul estimates.  相似文献   

2.
This paper presents a summary of all catch-comparison gear tests performed between 2002 and 2006 with Nordmøre-type sorting grids in the Swedish Nephrops trawl fishery in the Skagerrak and Kattegat. The overall aim was to minimise fish by-catch while maintaining catch rates for the target species, i.e. Nephrops. The results showed that a trawl fitted with a grid and a square-mesh cod-end, caught less small and marketable cod, haddock, whiting, hake and plaice than did a standard commercial trawl. For cod larger than 35 cm, the catch reduction was close to 100%. For marketable Nephrops no difference in catch amounts was detected. Results of three modified grid variants are also reported here. None of these showed convincing results in terms of better species-selectivity or superior handling characteristics than the standard grid. The results on fish by-catch reduction indicate that the Swedish grid may be a useful tool to reduce non-target mortality and to decouple Nephrops from cod and other demersal fish species under existing and future recovery plans.  相似文献   

3.
The selectivity of a 99 mm trawl codend was assessed using a codend cover fitted with a MultiSampler, which was acoustically triggered to take separate samples at three different phases of the haul. The first sample was collected during towing, the second during haul-up and the third at the surface. A total of 18 hauls were conducted with a commercial fishing vessel west of Scotland. It was demonstrated that escapes take place not only during the tow but also in the short period when the trawl is hauled back from the seabed and when the codend is at the surface. For haddock (Melanogrammus aeglefinus), whiting (Merlangius merlangus) and Norway lobster (Nephrops norvegicus), respectively, the mean percentages escaping at the surface were 16, 12 and 38% of the total escape while 17, 8 and 28% escaped during the haul-up phase. Compared to towing, the escape rate (no./min) increased for haddock by a factor 2.7 during haul-up and by a factor 1.7 at the surface, whereas the escape rates of whiting were similar for the three phases. The escape rate of Norway lobster increased by a factor of approximately 7 for both the haul-up and surface phases, compared to the towing phase. The selectivity parameters L50 (50% retention length) and SR (selection range = L75–L25) were estimated and compared for the three different phases and for the whole haul for haddock, whiting and Norway lobster. For all three species there was no significant (P > 0.05) difference in L50 between the three phases of the haul. There was also no significant difference for whiting and Norway lobster when comparing the SR of the three phases, whereas the SR was significantly lower for haddock when comparing the surface phase with towing and haul-up. The estimate of L50 when towing was about 6 cm lower for haddock and whiting and 9 mm for Norway lobster compared to the selection curve estimated conventionally for the whole haul. Finally, the effect of sea state, duration and codend catch on the selectivity parameters were estimated for the individual phases and for the whole haul. A significant effect of at least one variable was found in all phases.  相似文献   

4.
Discarding of fish species in the Kattegat-Skagerrak Nephrops directed fisheries remains at high levels. In this work we have tested four sets of codends pair-wise with the catch comparison technique under full commercial conditions to assess their potential in reducing the catch of undersized commercial species, in particular cod. We investigate the position of a 90 mm square-mesh panel (SMP), the effect of increasing the mesh size from 90 to 120 mm in the SMP, the effect of inserting a 90 mm SMP in an 80 mm codend and the effect of increasing the codend mesh size from 90 to 120 mm in a total of 89 hauls. Inserting the SMP 3–6 m from the codline compared to 6–9 m from the codline, reduced catches of cod above the minimum landing size (MLS) and Nephrops above and below the MLS. Increasing the mesh size in the SMP from 90 to 120 mm increased the catch of cod above the MLS in weight with an overall increase of 12% in the revenue. The effect of installing a 90 mm SMP 6–9 m from the codline in a nominal 80 mm codend had no effect on the catch of cod. Increasing the nominal codend mesh size from 90 to 120 mm reduced the catch of all species below the MLS, except monkfish. Catches of Nephrops above the MLS were, however, reduced by approximately one third and the total loss in revenue was 21%.  相似文献   

5.
Steel sorting grids are widely used in the North-east Arctic fisheries as excluding devices for juvenile fish. Sorting grids with four different bar spacings, 55, 60, 70 and 80 mm, were tested in sea trials during February–March 2006 and 2007. Cod and haddock were the target species. The study shows that there is little variation in the selectivity parameters when increasing bar spacing from 55 to 60 or 70 mm for haddock (2.7 cm in the l50 while the SR is fairly constant around 5 cm). For cod, no differences were found between the 55 and the 60 mm grids or the 70 and 80 mm grids, but the first two differed from the latter. The mean l50 increases from 56.08 to 73.33 cm and the mean SR from 7.46 to 14.28 cm when the bar spacing is increased from 55 to 80 mm. The selection curves move to the right and tend to lose sharpness, and the 95% confidence areas increase gradually as the bar distance of the grid is widened. The relationship between the l50 and grid bar spacing, based on this and previous studies, was determined to be linear for both cod and haddock.  相似文献   

6.
A substantial improvement in the bycatch selectivity of Norway lobster Nephrops norvegicus trawls is required, particularly with respect to cod Gadus morhua, whose stocks are at low levels in several areas. Conventional escape windows are not adequate to properly release cod and other bycatch species caught in the trawls. To address this issue, we developed a novel sorting box concept consisting of a four-panel section with a window on the top in order to improve the escape of cod and other bycatch species through an escape window while retaining the target catch of Norway lobster. The concept was tested on a commercial trawler in Kattegat and Skagerrak. Two different window mesh sizes and two different sorting box heights were tested using a traditional codend cover and a dual codend cover. We observed greatly reduced bycatches of both cod and other fish species compared to a standard codend. The reduction in bycatch decreased with decreasing mesh size and increasing height of the sorting box. Escape of Norway lobster through the escape window was limited. A modified version of the sorting box concept was implemented in the Kattegat fishery from 2009 onwards.  相似文献   

7.
The effect of trawling on fish size and on different quality parameters of cod (Gadus morhua) and haddock (Melanogrammus aeglefinus) was evaluated after conducting 16 valid hauls using two trawls in a double rig fitted with a traditional and a novel ‘T90’ codend, respectively. The total catch volume during the fishing period was 47.6 metric tons, with an average catch per codend of 1.5 (range 0.5–2.9) tons. The mean haul duration was 5 h. The catch was assessed according to fish size, mortality, external damage, initial white muscle pH and development of rigor mortis. Fillet quality (colour, blood spots, gaping) was assessed after 1 week of freeze-storage. Our results showed there was no difference between the two types of nets in terms of catch volume, but significantly slightly bigger fish were caught with T90 than with the traditional trawl net (p < 0.05). Haddock caught with the traditional trawl net had more external injuries related to the trawl gear than haddock caught with the T90 gear (p < 0.05). The gaping frequency for cod caught with the traditional trawl net tended to be higher than cod caught with the T90 gear, but the difference was not significant (p = 0.07). No other differences in fish quality between fish caught in the trawl nets were observed.  相似文献   

8.
The critical condition of the North Sea cod stocks has resulted in restrictions on not only cod, but also haddock and other species that are caught together with cod. Thus full exploitation of the haddock stock is unachievable unless cod can be excluded from the haddock catch. We designed a selective trawl based on the behavioral differences between haddock and cod as they enter a trawl, i.e., cod stay close to the seabed whereas haddock rise above it. The trawl's fishing line is raised ～60 cm above the seabed to allow cod to escape beneath the trawl while haddock are retained. To collect the escapees, three sampling bags were attached beneath the raised fishing line. The selective haddock trawl reduced the total catch of cod by 55% during the day and 82% at night, and 99% of the marketable haddock was caught during the day and 89% at night. Cod escape rates were highly length dependent: smaller cod escaped the trawl in greater numbers than did larger individuals. Whiting, saithe, lemon sole, and plaice were included in the analysis.  相似文献   

9.
《Fisheries Research》2007,87(2-3):262-267
Within commercial fisheries, particularly mixed fisheries, both target and non-target species are often discarded. Discarding represents a potentially significant loss to the productivity of fish stocks; it can have damaging ecological consequences, and is a potential cause of the failure of recovery plans. The Nephrops fishery in the North Sea is classified as a mixed fishery. Nephrops trawls are constructed with smaller meshes than trawls used to target whitefish; consequently, the bycatch of juvenile fish can be substantial. Several new Nephrops trawl designs have been tested in the North Sea. The data from these trials are used to investigate the potential impact of their implementation on cod, haddock and whiting stocks in the North Sea (including the Kattegat and Skagerrak).The model examines five trawl designs, and also the scenarios of a cessation of discarding in all North Sea fisheries and in just the Nephrops fishery. The model is deterministic, and evaluates the relative differences between scenarios assuming all other variables remain constant. If discarding of cod, haddock and whiting in the North Sea fisheries were eliminated, stocks would increase by 41%, 14% and 29%, respectively, within 10 years. Eliminating discarding in the Nephrops fishery alone would increase stocks by 2%, 1% and 13%, respectively, reflecting the relative proportion of catches of these species in the Nephrops fishery. For cod and haddock, the introduction of the Nephrops trawl with a grid with a square-mesh codend was the only scenario in which a notable increase in stock number was observed. This trawl design facilitates the escape of fish of all ages/sizes from the trawl, effectively making the Nephrops fishery a single-species fishery. For whiting, stock numbers and landings increased under all scenarios, but forecasted landings were lower than if current discard patterns continued in all except the no-discards scenario. The dependency of the results on the validity of the assumptions and on the accuracy of the input data is discussed.  相似文献   

10.
Gillnet selectivity parameters for the Atlantic sharpnose, Rhizoprionodon terraenovae, blacknose, Carcharhinus acronotus, finetooth, Carcharhinus isodon, and bonnethead, Sphyrna tiburo, sharks were estimated from fishery-independent catches in multi-panel gillnets with stretched mesh sizes ranging from 8.9 to 14.0 cm in steps of 1.3 cm, with an additional size of 20.3 cm. Mesh selectivities were estimated using a maximum-likelihood model, which fits a gamma distribution to length data for each mesh size using the log-likelihood function. The Atlantic sharpnose and finetooth shark exhibited the broadest selection curves. Peak selectivities for the Atlantic sharpnose were reached from 750 mm FL for the 8.9 cm mesh to 1150 mm FL for the 14.0 cm mesh in 50 mm FL increments per mesh. Peak selectivity for the finetooth shark was reached at 550 mm FL for the 8.9 and 10.2 cm meshes, increased to 650 mm FL for the 11.4 mesh, and 750 mm FL for the 12.7 and 14.0 cm meshes. Selectivity was highest at 1150 mm FL for the 20.3 cm mesh. The bonnethead and blacknose shark exhibited narrower selection curves, with peak selectivity occurring at 450 mm FL for the 8.9 cm mesh, 750 mm for the 12.7 cm mesh in 100 mm FL increments per mesh. Maximum selectivity for the 20.3 cm mesh was 950 and 1050 mm FL for bonnethead and blacknose shark, respectively. The θ1 values for blacknose and finetooth shark were most similar (140.58 and 141.25), whereas the value calculated for Atlantic sharpnose was the highest (211.95) and that for the bonnethead (131.77) was the lowest. Values calculated for θ2, a parameter that describes the variance of sizes by mesh, ranged from 27,259 for the bonnethead to 189,873 for the finetooth shark. Although gillnets used in this study were not directly constructed for use in estimation of gillnet selectivities, information on mesh selectivities estimated herein has direct applicability to commercial gillnets with meshes of similar sizes.  相似文献   

11.
杨炳忠  晏磊  李杰  王腾  张鹏 《中国水产科学》2023,30(10):1271-1281
大网目拖网是我国海洋渔业捕捞带鱼的重要渔具。为提高拖网网囊选择性, 本研究在南海区采用套网法对双船大网目拖网 4 组网目尺寸(30、35、40 和 45 mm)的网囊进行实验, 在双重拔靴法的架构下, 拟合选择性参数、比较选择性的差异、估算捕捞方式指标。结果表明, 随着网目尺寸的增大, 网囊对带鱼的 50%选择体长增大, 对带鱼幼鱼个体的选择率显著下降, 带鱼幼鱼的渔获比例也呈下降趋势。结合带鱼最小可捕规格(23.0 cm)管理规定, 结果还表明网目尺寸为 40 mm 的网囊选择性较差, 其对带鱼的抛弃率大于 93%。为了进一步提高带鱼拖网渔业的资源养护水平, 还需要开展更多的选择性实验。  相似文献   

12.
Mediterranean demersal trawl fisheries traditionally operate using small diamond-shape meshes in the codend, which tend to retain almost all animals. We investigated the effect of mesh configuration on the size selectivity of nine species (the scaldfish Arnoglossus laterna, the broad-tail shortfin squid Illex coindettii, the European hake Merluccius merluccius, the red mullet Mullus barbatus, the Norway lobster Nephrops norvegicus, the common pandora Pagellus erythrinus, the deepwater rose shrimp Parapenaeus longirostris, the Mediterranean horse mackerel Trachurus mediterraneus and the poor-cod Trisopterus minutus capelanus) commonly captured in the Mediterranean demersal trawls. Two codends having meshes with the same opening (ca. 38 mm) but different mesh configuration (DM38: diamond-mesh and SM38: square-mesh configuration) were fished daily and alternately on the same trawl. They demonstrated a substantial improvement in selectivity with square-mesh. Selectivity was measured using the covered codend technique, the cover being supported by circular hoops.The results were analysed taking into account the between-haul variation in selectivity. Two other important external variables were identified: the trawling depth and the codend catch, which for some species influenced between-haul variation, but there was no evidence of a coherent effect on selectivity parameters.The selectivity of 40 mm diamond-mesh codend has been reported to be rather poor because a large proportion of the codend catch is immature and smaller than the minimum landing size (MLS) or first maturity size. With the exception of the flatfish (A. laterna), the effect of a change of mesh configuration from diamond- to square-mesh on size selectivity positively affected the retention length at 50% (L50). However, square-meshes were found to be unsuitable for flat and/or deep-bodied fish as these escape more readily from diamond-meshes.The Council Regulation (EC) No. 1967/2006, concerning management measures for the sustainable exploitation of fishery resources in the Mediterranean, establishes the MLS of marine organisms. However, the increase in L50 with square-mesh would not avoid some of the existing contradictions in allowing the use of codend mesh which leads to lower L50 than the MLS.  相似文献   

13.
ABSTRACT:   The demersal multispecies trawl fishery in the western Mediterranean Sea has a poor selectivity; large numbers of juvenile fish are caught when using the legislated 40-mm diamond mesh cod end. The selectivity of a flexible sorting grid with 20-mm bar spacing (BS20) installed in front the conventional trawl cod end was investigated. The standard covered cod end method was used. Data was adequate for analyzing the selectivity of European hake Merluccius merluccius , poor cod Trisopterus minutus , greater forkbeard Phycis blennoides and Norway lobster Nephrops norvegicus . The selectivity of the 20-mm sorting grid was compared with the performance of 40-mm diamond (DM40) and square mesh (SM40) cod ends. The effect of a guiding funnel on the performance of the grid (BS20-f) was also explored. Finally, the potential changes in yield per recruit (Y/R) and biomass per recruit (B/R) after implementing a sorting grid were explored. An overall improvement in the 50% selection length ( L 50 ) with all four species was substantial when comparing the BS20 to the DM40. Higher improvement in L 50 s was achieved when the grid was equipped with the guiding funnel. When comparing the performance of the BS20-f to the SM40, there was no marked difference in L 50 s. The Y/R and B/R, however, are substantially higher with BS20-f for poor cod and about the same for European hake. BS20 compared to SM40 achieved lower Y/R and B/R for all four species; the smallest difference was observed for Norway lobster. Further development is required if the sorting grid is to be introduced into commercial fisheries.  相似文献   

14.
青岛斋堂岛海域春季双桩竖杆张网网囊网目的选择性   总被引：3，自引：1，他引：2  
2014年春季,采用平行作业法,以2a=16 mm(a为目脚长度)的菱形目网囊为对照网(CN),选取30 mm菱形目网囊和20 mm、30 mm、40 mm的方形目网囊(分别以30D、20S、30S和40S表示)作为实验网囊,对青岛斋堂岛附近海域的双桩竖杆张网进行了不同网目形状的网囊和不同网目尺寸方形目网囊的选择性比较试验。以多元统计分析方法进行种间选择性分析,以Logistic曲线作为选择性模型、采用极大似然估算法进行种内选择性分析。结果显示,与对照网相比,所有实验网囊均能有效释放玉筋鱼(Ammodytes personatus)、舒氏海龙(Syngnathus schlegeli)等低值渔获以及经济鱼类的幼体。比较不同网目形状网囊的渔获数据发现,方形目网囊较菱形目网囊具有更好的选择性,其中30S网囊对玉筋鱼和舒氏海龙的释放率(以尾数计)分别比30D网囊高4.61%和16.80%,对六丝钝尾虾虎鱼(Amblychaeturichthys hexanema)和日本枪乌贼(Loigo japonica)的50%选择体长(胴长)L50分别比30D网囊大38.47%和61.29%。比较不同网目尺寸方形目网囊的渔获数据发现,当实验网为20S和30S时,六丝钝尾虾虎鱼的L50分别为6.58 cm和7.20 cm,选择范围(SR)分别为3.02 cm和0.87 cm;当实验网为20S、30S和40S时,日本枪乌贼的L50分别为2.88 cm、4.03 cm和10.49 cm,SR分别为0.74 cm、0.52 cm和2.67 cm;不同尺寸的方形目网囊对这2种渔获物的L50随着网目尺寸的增大而增大。根据实验海区各种渔获物的经济价值、低值鱼类和幼鱼的释放比例以及分拣渔获物的劳动力投入等综合分析认为,30 mm方形目网囊在保护渔业资源的同时对渔业生产不会造成显著影响,建议予以推广。  相似文献   

15.
Tank respirometry was used to study the interactive effects of protein:lipid level (55%:11% vs. 42%:16%; both diets isoenergetic) and temperature (11, 6 and 2 °C) on the magnitude and duration of specific dynamic action (SDA) in juvenile Atlantic cod and haddock. The protein:lipid level did not affect any measured variable. However, numerous temperature and species effects were observed. For example, although the maximum post-feeding oxygen consumption (30–50% above routine metabolic rate; RMR) and SDA duration (∼55–85 h; SDADUR) were not affected by temperature, SDADUR g−1 of food increased from 11 to 2 °C (from ∼3 to 12 h g food−1). While absolute SDA (mg O2) decreased by ∼60–65% in cod and ∼75% in haddock from 11 to 2 °C, due to a concomitant decrease in food consumption from ∼2.0% to 0.6% body mass, SDA comprised between 3.3% and 5.2% of the dietary energy content at all temperatures. Finally, RMR at 11 and 2 °C and SDADUR at 2 °C were 25–35% and 25% greater in cod, respectively, as compared with haddock. These results suggest that feeding reduced protein diets at low water temperatures is unlikely to improve the growth of these species.  相似文献   

16.
The seasonal and geographical patterns of spawning for Georges Bank cod ( Gadus morhua ) and haddock ( Melanogrammus aeglefinus ) are estimated using composite distributions of stage I eggs derived from the Marine Resources Monitoring, Assessment and Prediction (MARMAP) data set (1977–1987). The inferred mean spawning locations are compared with patterns in particle residence times derived from a three-dimensional prognostic circulation model that estimates the tidal and seasonal mean circulation. The comparisons indicate that cod and haddock spawning occurs at times and locations characterized by model residence times in excess of 35 d. The results are discussed in the context of fish population regulation theory, particularly the member/vagrant hypothesis and the concept of abundance-dependent vagrancy.  相似文献   

17.
To examine the age, growth and reproduction of silky sharks, Carcharhinus falciformis, in the waters off northeastern Taiwan, 469 specimens (213 females and 256 males) were collected from August 2000 to January 2002 at the Nanfanao fish market, northeastern Taiwan. The relationship between body weight (W) and total length (TL) for both sexes combined was expressed as: W = 2.92 × 10−6 TL3.15 (n = 469, p < 0.01). The relationship between TL and vertebral centrum radius (R) for both sexes combined data was estimated as: TL = 25.979 + 18.197R (n = 250, p < 0.01). Growth bands (including translucent and opaque zones) in precaudal vertebrae formed once a year between December and January and were counted up to 11 and 14 for females and males, respectively. The von Bertalanffy growth function (VBGF) was used to model the observed length at age data. The sexes combined VBGF predicted an asymptotic length (L∞) = 332.0 cm TL, growth coefficient (k) = 0.0838 year−1, age at zero length (t0) = −2.761 year (n = 250, p < 0.01). Size at 50% maturity for males was estimated to be 212.5 cm based on the logistic curve, which corresponded to 9.3 years. Females matured at 210–220 cm, which correspond to 9.2–10.2 years. The length at birth was estimated to be 63.5–75.5 cm TL. The number of embryos per litter was 8–10 and sex ratio of embryos was 1:1.  相似文献   

18.
Abstract  The selectivities and catch compositions of conventional 30-mm diamond-mesh codends and a new square-mesh design made from 20-mm mesh hung on the bar for river and lagoon penaeid seines in south-eastern Australia were compared. The square-mesh design significantly improved the selectivities of the river seine for school prawns, Metapenaeus macleayi (Haswell), and the lagoon seine for greasyback prawns, Metapenaeus bennettae (Racek & Dall), by increasing their carapace length at 50% probability of retention ( L 50) and by decreasing between-haul variability in selectivity. The presence of weed reduced the L 50 for greasyback prawns caught in the conventional diamond-mesh codend during the lagoon-seine experiment. The differences among codend performances are discussed in terms of the differing methods of operation and composition of catches between the seines. These differences highlight the need to develop and manage modifications to improve the selectivity of fishing gears on a fishery-specific basis.  相似文献   

19.
Many jurisdictions have introduced management regimes prohibiting or limiting discarding, and a key objective of the European Union Common Fisheries Policy is the gradual elimination of discards. One way of reducing the catch of unwanted fish is to use more size‐selective gears. Gear‐based management options can best be explored using models that predict size selection across a wide range of variables related to gear design. Such a model is developed for haddock (Melanogrammus aeglefinus) through a meta‐analysis of 21 trials of codend selection and 19 trials of the combined selection of the codend and a square mesh panel. Individual‐haul estimates of the 50% retention length (l50) and the selection range (SR) are related to a set of explanatory variables through a structural model that describes the dual process of panel and codend selection. Codend l50 and SR are positively related to codend mesh size and negatively related to the codend twine diameter. Codend l50 is also negatively related to the number of open meshes around the codend circumference. Panel l50 increases with panel mesh size. The panel contributes more to gear selection as it is moved closer to the codline. The panel is most effective between November and January and least between May and July, periods which broadly coincide with peak and poor haddock condition. The results are illustrated for a typical trawler targeting haddock in the Scottish whitefish sector, and the utility of the model for gear design and legislation is discussed.  相似文献   

20.
《Fisheries Research》2007,83(1-3):111-118
The legally sized fish by-catch is an important economic component of many Nephrops fisheries. However, the small mesh size typically used by Nephrops trawlers results in many undersized fish being discarded. A selection grid was inserted in a two-tier cod-end in an attempt to separate the target species, Nephrops, from the fish by-catch and to retain the by-catch in the upper cod-end. Good separation would allow fish size selection to be improved by using a larger mesh size in the upper cod-end. The separation properties of grids with two different bar spacings and two lower gaps were investigated. Nephrops, haddock and dab were caught in reasonable numbers. Most small Nephrops passed into the lower cod-end, with the proportion passing into the upper cod-end increasing to between ∼40 and ∼70% at carapace lengths of 50 mm. The grids with the larger bar spacing allowed more Nephrops into the lower cod-end, but there was no significant difference in separation between grids for haddock or dab. The proportion of haddock passing into the upper cod-end increased from ∼60% for 10 cm haddock to ∼85% for haddock of 20 cm or more. The proportion of dab passing into the upper cod-end increased from ∼30% for 10 cm dab to ∼75% for dab of 20 cm or more.  相似文献   

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