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1.
Northern hardwood stands, notably those with American beech (Fagus grandifolia Ehrh), sugar maple (Acer saccharum Marsh.), and yellow birch (Betula alleghaniensis Britton), are abundant across the forested landscapes of northeastern USA and southeastern Canada. Recent studies have reported an increasing dominance of American beech in the understory and midstory of these forests. Beech is a commercially less desirable tree species due to its association with beech-bark disease, and because it commonly interferes with the regeneration of other more desirable tree species. We examined hardwood regeneration characteristics nine years after application of a 3 × 4 factorial combination of glyphosate herbicide (0.56, 1.12, and 1.68 kg ha?1) and surfactant concentrations (0.0, 0.25, 0.5, and 1.0% v v?1) to release sugar maple regeneration from beech-dominated understories using three stands that received shelterwood seed cutting in central Maine. Measurements nine years after treatment showed that glyphosate rate increased both absolute (AD) and relative density (RD) of sugar maple regeneration, but not its height (HT). In contrast, beech AD, RD, and HT were all significantly reduced with increasing glyphosate rate. Post-release browsing by ungulates and a high residual overstory basal area resulted in reduced sugar maple HT. Our results indicated that glyphosate herbicide applied in stands that have been recently shelterwood seed cut can significantly increase the abundance of sugar maple regeneration. However, subsequent browsing damage combined with the negative influence of the residual overstory cover can limit the longer-term benefit of understory herbicide treatments. Subsequent removal of the overstory and browsing-control measures may be needed to promote sugar maple regeneration over beech in similar northern hardwood stands.  相似文献   

2.

??Context

It is assumed that climate change will favour European beech (Fagus sylvatica L.) to Norway spruce (Picea abies [L.] Karst.) at its northern range margins due to climate change and induced disturbance events.

??Aims

An old-growth mixed forest of spruce and beech, situated near the northern beech margin, was studied to reveal effects of disturbances and response processes on natural forest dynamics, focussing on the understory.

??Methods

We carried out analyses on understory dynamics of beech and spruce in relation to overstory release. This was done based on a sequence of stand and tree vitality inventories after a series of abiotic and biotic disturbances.

??Results

It became apparent that beech (understory) has a larger adaptive capacity to disturbance impacts and overstory release (68 % standing volume loss) than spruce. Understory dynamics can play a key role for forest succession from spruce to beech-dominated forests. Disturbances display an acceleration effect on forest succession in the face of climate change.

??Conclusion

Beech is poised strategically to replace spruce as the dominant tree species at the study area. Due to an increasing productivity and a lower risk of stand failure, beech may raise into the focus of forestry in southern Sweden.  相似文献   

3.

Aims

In European Beech (Fagus sylvatica L.) large growth stresses lead to severe log end splitting that devaluate beech timber. Our study aimed at detecting relationships between growth stress and some morphology parameters in trees.

Methods

Growth stress indicators were recorded for 440 mature trees in nine stands from five European countries, together with morphology parameters.

Results

Most trees displayed an uneven distribution of growth stress around the trunk. Moreover, growth stress intensity varied largely between individual trees. Geometry of the trunk was a poor predictor of growth stress intensity. Crown asymmetry resulted in a larger stress dissymmetry within trees. Trunk inclination was not correlated to max tension stress, contrary to what is usually found in younger trees. In the case of small inclination, growth stress was close to expected from biomechanics of restoring verticality. Trees exhibiting a larger inclination probably evolved a different mechanical solution: a rather large crown, lower tree slenderness and a sufficient asymmetry in growth stress as to prevent a higher inclination due to growth.

Conclusion

A large slenderness is the best accurate predictor of a large growth stress, although variations in the ratio height/diameter at breast height explained only 10 % of the variability of growth stress. A large crown surface was the best predictor of a low level of growth stress. A large spacing between trees seems a good solution to lower the risk of growth stress in mature beech.  相似文献   

4.

? Context

The knowledge of how shrub–seedling interactions vary with summer drought, canopy opening, and tree species is crucial for adapting forest management to climate change.

? Aims

The aim of this study was to assess variation in shrub–oak recruitment associations along a south–north drought climate gradient and between two levels of canopy cover in coastal dune forest communities in a climate change-adapted forest management perspective.

? Material and methods

Mapped data of associational patterns of seedlings of three oak species with interspecific pooled shrubs were analyzed using a bivariate pair correlation function in 10 (0.315 ha) regeneration plots located in forest and recent gap sites along the climate gradient. An index of association strength was calculated in each plot and plotted against a summer moisture index.

? Results

The association strength increased with increasing summer drought from wet south to dry north and from closed forests to gaps.

? Conclusion

Consistent with facilitation theory, our results suggest that climate change may shift associational patterns in coastal dune forest communities towards more positive associations, in particular in canopy gaps. In a perspective of climate change, foresters may need to conserve understory shrubs in gaps in order to promote oak species regeneration.  相似文献   

5.

? Context

It is assumed that global change is already affecting the composition, structure and distribution of forest ecosystems; however, detailed evidences of altitudinal and latitudinal shifts are still scarce.

? Aims

To develop a method based on National Forest Inventory (NFI) to assess spatio-temporal changes in species distributions.

? Methods

We develop an approach based on universal kriging to compare species distribution models from the different NFI cycles and regardless of the differences in the sampling schemes used. Furthermore, a confidence interval approach is used to assess significant changes in species distribution. The approach is applied to some of the southernmost populations of Pinus sylvestris and Fagus sylvatica in the Western Pyrenees over the last 40 years.

? Results

An increase of the presence of the two species in the region was observed. Scots pine distribution has shifted about 1.5 km northwards over recent decades, whereas the European beech has extended its distribution southwards by about 2 km. Furthermore, the optimum altitude for both species has risen by about 200 m. As a result, the zone in which the two species coexist has been enlarged.

? Conclusions

This approach provides a useful tool to compare NFI data from different sampling schemes, quantifying and testing significant shifts in tree species distribution over recent decades across geographical gradients.  相似文献   

6.

Context

Excessive recruitment in post-fire regeneration of Pinus halepensis can require pre-commercial thinning. The 1994 Moratalla fire (Spain) and the thinning applied there since 2004 provided good conditions for testing pre-commercial thinning effectiveness.

Aims

To analyse pre-commercial thinning effects on tree size, reproductive potential, stem biomass and annual growth in 15-year-old saplings.

Methods

Twenty nine circular plots (5 m radius) were established based on (1) years since thinning and (2) aspect. Non-thinned plots were included as controls. Sapling variables were measured and pine cones were counted and classified according to maturity. Three saplings per plot were felled, five sample disks were extracted from each and tree rings were analysed with Windendro® software.

Results

Sapling size, annual growth and stem biomass were higher in thinned plots than in control plots 2–4 years after thinning. Annual growth declined five years after thinning. Cone counts were higher 2–5 years after thinning, depending on the cone type. Aspect influenced some variables.

Conclusion

Thinning accelerated stand maturity but positive effects were indiscernible until 2–5 years later. Serotinous cones, which are fundamental to regeneration after fire, increased in number after 5 years. The effects of aspect were inconclusive due to interactions with years since thinning.  相似文献   

7.

? Context

While historical increases in forest growth have been largely documented, investigations on historical wood density changes remain anecdotic. They suggest possible density decreases in softwoods and ring-porous hardwoods, but are lacking for diffuse-porous hardwoods.

? Aims

To evaluate the historical change in mean ring density of common beech, in a regional context where a ring-porous hardwood and a softwood have been studied, and assess the additional effect of past historical increases in radial growth (+50 % over 100 years), resulting from the existence of a positive ring size–density relationship in broadleaved species.

? Methods

Seventy-four trees in 28 stands were sampled in Northeastern France to accurately separate developmental stage and historical signals in ring attributes. First, the historical change in mean ring density at 1.30 m (X-ray microdensitometry) was estimated statistically, at constant developmental stage and ring width. The effect of past growth increases was then added to assess the net historical change in wood density.

? Results

A progressive centennial decrease in mean ring density of ?55 kg?m?3 (?7.5 %) was identified (?10 % following the most recent decline). The centennial growth increase induced a maximum +25 kg?m?3 increase in mean ring density, whose net variation thus remained negative (?30 kg?m?3).

? Conclusions

This finding of a moderate but significant decrease in wood density that exceeds the effect of the positive growth change extends earlier reports obtained on other wood patterns in a same regional context and elsewhere. Despite their origin not being understood, such decreases hence form an issue for forest carbon accounting.  相似文献   

8.

Context

Although adventitious shoots are produced in many tree species in response to injury, little is known about the effects of salinity on sprouting.

Aims

The main objective was to examine the sprouting capacity of Pinus leiophylla seedlings in relation to injury and physiological changes induced by NaCl.

Methods

Seedlings were grown in controlled-environment growth rooms and treated with 0, 100, 150, and 200 mM NaCl. Numbers of adventitious shoots were recorded and growth and physiological parameters measured after 64 days of treatments and following a 30-day recovery period.

Results

NaCl treatments triggered sprouting of adventitious shoots, mainly in the lower parts of the stems. However, fewer sprouts were induced by 200 mM NaCl compared with the lower concentration treatments. These changes were accompanied by needle necrosis, decreased chlorophyll concentrations, seedling dry mass, and stem diameter. Stomatal conductance, net photosynthesis, and root hydraulic conductance decreased with increasing NaCl concentrations and did not return to the control levels after 30 days of stress relief.

Conclusions

Pinus leiophylla has the regeneration potential when exposed to salinity. However, very high salt concentrations induce severe physiological impairments and, consequently, a decrease of this regeneration potential.  相似文献   

9.

Context

Projections of species distribution models under future climate are usually based on long-term averages. However, singular extreme drought events presumably contribute to the shaping of distribution limits at the retreating low-elevation xeric limits.

Methods

The objectives of this study were to set up a distribution model based on extreme drought events (EDM), which uses sanitary logging information as a proxy of vitality response of beech, and compare it with the results of classical species distribution models (SDMs).

Results

Predictions of the EDM for 2025 were in agreement with those of the SDM, but EDM predicted a more serious decline in all regions of Hungary towards the end of the century.

Conclusion

These results suggest that the predicted increase in frequency and severity of drought events may further limit the distribution of beech in the future.  相似文献   

10.

Context

Natural regeneration with broadleaved species and reforestation with coniferous trees are two widely practiced forest regeneration strategies after timber harvesting. They lead to different tree species composition and may cause different understory biodiversity, but the effects on ground bryophyte composition and diversity are not well-known.

Aims

We tested whether natural regeneration with broadleaved species and reforestation with spruce induced different diversities of the ground bryophyte populations 20–40 years after old-growth spruce forest clearcutting in the subalpine regions of southwestern China.

Methods

Differences between natural stands and plantations were compared through the analysis of 13 paired stands, with 78 plots, 390 shrub/herb quadrats, and a total of 1,560 bryophyte quadrats.

Results

Naturally regenerated forests were characterized by lower density and cover and lower tree height but higher herbaceous plant height, shrub cover, and bryophyte diversity. They also harbored many more ground bryophytes. The species richness of pleurocarpous mosses and fans, mats, and turfs were significantly higher in naturally regenerated forests. Frequency difference analysis demonstrated that more bryophyte species preferred ground habitats in naturally regenerated forests than in plantations (116 vs. 48 species). The canonical correspondence analysis indicated that stand structure attributes were more important determinants of ground bryophyte diversity and abundance.

Conclusion

Natural regeneration and reforestation resulted in large differences in ground bryophyte populations. A larger diversity was observed in the former case, and natural regeneration practices can be an effective measure for the protection of ground bryophyte diversity after clearcutting.  相似文献   

11.

Context

Pulsed food resources may strongly affect the population dynamics of several consumer species, with consequences on the ecosystem. One of the most common pulsed resources is forest mast seeding.

Aims

We analysed mast seeding in deciduous forests in a mountainous area of northern Apennines and its effect on population dynamics of wild boar (Sus scrofa L.).

Methods

We performed a quantitative, 20-year analysis on annual seed production in Turkey oak (Quercus cerris L.), beech (Fagus sylvatica L.) and chestnut (Castanea sativa Mill.) forest stands using litter traps. The wild boar population density was estimated by means of drive censuses and hunting bag records. The role of other biotic (density of predators) and abiotic (climate) factors potentially affecting wild boar mortality was also investigated.

Results

Turkey oak and chestnut showed high levels of seed production, whereas lower levels were found in beech. The pulsed resources of chestnut and Turkey oak positively affected piglet density. Analyses also highlighted the influence of snow cover and wolves on wild boar population dynamics.

Conclusion

Wild boar can be considered a pulse rate species, the management of which can be improved by annual monitoring of seed production.  相似文献   

12.

?Context

Selective logging followed by natural regeneration is rarely employed for restocking subtropical evergreen broad-leaved forests in East Asia compared with the use of clear-cutting.

?Aims

To clarify the succession of these forests, the effects of selective logging on stand structure, species diversity, and community similarity were studied in a mature and regenerating forest in Okinawa, Japan.

?Methods

Four study plots were established, and trees ≥1.2 m height were identified by species name, tree height, and diameter at breast height.

?Results

The results showed that the species composition of regenerating forest was similar to mature forest; however, the former had a greater species density and Shannon–Wiener index than the latter. Castanopsis sieboldii and Distylium racemosum, the predominant trees in the mature forest, continued to dominate the regenerating forest, with a broad layer distribution. High Sørensen and Jaccard community similarity indices for mature and regenerating forest indicated that the regeneration occurred in a progressive succession.

?Conclusion

The similar species composition and stand structure for both mature and regenerating forest, and the higher species diversity for the latter, provided no evidence of forest degeneration and suggested that the regenerating forest may develop into a stand similar to preselective logging forest.  相似文献   

13.

Context

Climate change is expected to increase forest vulnerability through disturbances such as windstorms and droughts. Forest managers are therefore investigating strategies to increase forest resistance and resilience, especially by promoting uneven-aged and mixed forests through group selection, and by reducing stand stocking and large trees proportion. However, there is little information on the long-term impacts of these two practices.

Aims

The objectives of this study were (1) to develop an original silviculture algorithm designed for uneven-aged management and (2) to use it to assess the effects of the above-mentioned management methods in long-term simulations.

Methods

We simulated individual and group selection techniques in order to study the effects of group size, harvesting intensity and their interactions on wood production, stand heterogeneity, and regeneration in mountain spruce–fir forests. We used the spatially explicit individual-based forest model Samsara2 to simulate forest dynamics.

Results

Our simulation results confirmed the positive effect of group selection practices on structure diversity and regeneration but not on spruce maintenance. Increasing harvesting intensity enabled forest destocking but decreased structure diversity and led to non-sustained yields for the most intensive scenarios.

Conclusion

As adaptation measure, we thus recommend moderate group selection harvesting creating 500 m2 gaps.  相似文献   

14.

Context

Old-growth trees play a very important role in the maintenance of biodiversity in forests. However, no clear definition is yet available to help identify them since tree age is usually not recorded in National Forest Inventories.

Aims

To develop and test a new method to identify old-growth trees using a species-specific threshold for tree diameter in National Forest Inventories.

Methods

Different nonlinear mixed models for diameter–age were generated using data from the Spanish Forest Inventory in order to identify the most appropriate one for Aleppo pine in its South-western distribution area. The asymptote of the optimal model indicates the threshold diameter for defining an old-growth tree. Additionally, five site index curves were examined to analyze the influence of site quality on these models.

Results

The Hossfled III mixed model was found to be the most appropriate to fit diameter–age curves for Aleppo pine trees. The overall diameter at breast height threshold for old-growth trees was 40.6 cm, although over a range of sites with increasing site quality, the threshold figure was 36.0, 38.0, 40.4, 43.1, and 46.3 cm, respectively.

Conclusions

This method allows the identification of old-growth trees and therefore of biodiversity hotspots, thus providing decision makers with a useful tool for management purposes.  相似文献   

15.

Context

In the context of a probable increase in intensity and frequency of extreme summer drought events, a better understanding of the key processes involved in water relations is needed to improve the theoretical foundations of predictive process-based models.

Aims

This paper aims to analyse how temperate deciduous trees cope with water shortage.

Methods

The exceptional summer drought of 2003 in Europe provided an opportunity to monitor stomatal conductance and twig water potential in European beech (Fagus sylvatica L.) at predawn and midday and to analyse variations with respect to leaf height within the canopy. By comparing our field measurements of twig water potential to values found in the literature, we confirmed the strong impact of soil water shortage on crown water relations.

Results

This paper shows that (1) the vertical gradient of stomatal conductance within the crown disappeared under extreme soil water depletion; (2) at maximum drought intensity, predawn twig water potential (ψ pd) reached ?2.3 MPa at a height of 14 m in the crown and ?2.0 MPa at a height of 10 m. The significant differences in ψ pd between the two measurement heights in the canopy may be due to night transpiration; (3) there was a close relationship between predawn twig water potential and relative extractable soil water; (4) as drought conditions intensified, there was a close relationship between canopy radiation interception and predawn water potential, as estimated daily from relative extractable soil water.  相似文献   

16.

Context

After wildfire, surviving trees are of major ecological importance as they can help in the post-fire regeneration process. Although these trees may be damaged, they may also benefit from reduced fuel hazard and competition. However, little is known about the long-term growth response of surviving trees.

Aims

This study aims to explain short- to long-term variations in the postfire growth of surviving black pines in an area burnt in 1994, focusing on levels of fire severity and tree sizes.

Methods

Relative basal area increments were used to detect time-course variations in postfire radial tree growth depending on fire severity. Linear mixed-effects models were used to describe the factors affecting postfire ring growth.

Results

In the short term, fire caused stronger reduction in growth in small trees with increasing bole char height. However, as time since fire increased, a positive effect of fire on growth due to reduced competition counteracted the short-term fire impacts. Indeed, small surviving trees demonstrated a surge in growth 15 years after the fire.

Conclusion

It was concluded that reduced competition might offset the short-term negative effects of fire in surviving black pines.  相似文献   

17.

Context

Reliable estimates of wood density (WD) within individual trees could maximize the value of Pinus massoniana for specific end-use.

Aim

We examined and quantified the axial patterns of WD in trees with different social status in the stands.

Methods

Wood disks were sampled at the bottom, breast height, and middle of each 1-m sections from 108 stems, harvesting from three social classes in subtropical forests. A mixed-effects model was designed to quantify axial variation.

Results

The WD at different height was significantly different from the whole-stem WD (WWD) except the relative height of 0.1. An overall decrease of 133.8 kg m?3 in WD was found from stem base to top. WD was significantly influenced by relative heights, tree age, and social class. WD of each relative height in mature trees was significantly higher than that of younger trees. Tree social class can affect WD development in the axial direction at age classes 2 and 3. Combining the fixed plus random effects, the final model explained 91 % of the observed variation in WD.

Conclusion

The WD development patterns in the axial direction vary considerably among tree age, diameter at breast height, and social class. To distinguish and supply timber for specific end-uses, we should use the axial variation in disk WD (DWD) instead of WWD directly. The accurate predictions of WD provided by the model could be used to optimally classify logs into different product classes and maximize economic benefits. We can use DWD at the relative height of 0.1 instead of WWD of a single tree.  相似文献   

18.

??Context

In the construction sector, wood is facing competition with other materials such as concrete, steel or plastics. Therefore, there is a need for more efficiency in the forest–wood chain by improving silvicutural management and wood processing technologies.

??Aims

The objective of the study is to analyse the influence of log diameter and quality to recovery rate, veneer quality and economic benefit.

??Methods

The trees used in the study came from a 30-year-old Pinus taeda L. thinning trial in Southern Brazil. In total, 57 logs (20.7 to 67.0 cm) were peeled following the standard industrial processing methods of the plywood mill.

??Results

Average recovery rate was 54 % ranging from 35 to 72.6 %, with a linear trend (R 2?=?0.48) of increasing recovery with an increment on the log small-end diameter. Results show that the gap between theoretically possible and real recovery was lower in the logs with bigger diameters, indicating their higher efficiency in industrial processing. Moreover, the economic analysis detected that the current prices for log assortments reflect only the industrial potential of low-quality pruned logs. An optimised pruning strategy would result in higher industrial efficiency, which would allow higher log prices.

??Conclusion

The results indicate that the recovery rate of bigger logs is higher in terms of volume of peeled veneer. The quality and therefore the value obtained from each log were negatively influenced by inadequate pruning strategies. Management of pines for higher value utilisation requires optimized thinning and pruning strategies in order to meet high growth rates and proportionally bigger dimensions of clear wood.  相似文献   

19.

? Context

A 20-year-old Nelder wheel planted with hoop pine (Araucaria cunninghamii Aiton ex D.Don) and Queensland maple (Flindersia brayleyana F.Muell.) in 18 spokes and 8 rings represents nominal point densities of 3,580, 2,150, 1,140, 595, 305, 158, 82, and 42 stems/ha and offers an opportunity to examine competition and spatial interaction between these two species.

? Aims

This study aimed to evaluate the intraspecific and interspecific competition between two contrasting tree species and to determine the distance over which competition can be observed.

? Methods

Competition was estimated using Hegyi’s index, implemented using the Simile visual modeling environment, and calibrated using nonlinear least squares with PEST.

? Results

Interactions were detected between pairs of stems closer than D ij ?<?40(d i ?+?d j ) where D is distance (in centimeters) and d is stem diameter (in centimeters diameter at breast height). F. brayleyana trees surrounded by A. cunninghamii trees experience negligible competition, whereas A. cunninghamii surrounded by F. brayleyana trees suffer strong competition.

? Conclusion

Forty times diameter offers a useful guide to the extent of competition in even-aged stands planted with these species. Competition can be observed empirically when pairs of trees are closer than 40 times the sum of their diameters, but the intensity of the competition may vary considerably with species.  相似文献   

20.

Context

Stem straightness is an important trait for growers and processors of Eucalyptus globulus logs for solid-wood products.

Aims

The aims of the study were to determine the extent of genetic variation in stem deviation from straightness in E. globulus and assess the utility of a six-point subjective scoring method as a selection criterion for stem straightness.

Methods

Two E. globulus progeny trials, grown under solid-wood product regimes, were studied. At age 9 years (post-thinning), stem straightness was measured using both image analysis and a six-point subjective scale. Diameter at breast height (DBH; 1.3 m) was measured at both age 5 (pre-thinning) and age 9 years.

Results

Significant additive genetic variation was observed. Strong, positive and significant additive genetic correlations were observed between the stem straightness assessment methods and between DBH at ages 5 and 9 years. Significant positive genetic correlations were shown between subjectively scored stem straightness and DBH at both ages 5 and 9 years.

Conclusion

The six-point subjective scoring method is a cost-effective selection criterion for stem straightness in E. globulus. The image measurement technique may be applied where objective estimates of stem straightness are required, for training purposes and to verify subjective scores.  相似文献   

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