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1.
The aim of the study was to investigate dynamics and pattern of natural regeneration and the influence of seedbed and light on germination and initial seedling survival of Scots pine (Pinus sylvestris L.) during three growing seasons. Four 5-m2 plots in a natural Scots pine forest in central Spain were randomly established at the beginning of the regeneration process. Germination and seedling survival were recorded in 100 regeneration sampling subplots (0.25 m2) per plot, while seedbed type and the relative light intensity reaching the forest floor was characterised in 1-m2 subplots. The spatial correlations between survival, light conditions and organic matter depth were analysed through cross-variograms. Germination and survival were highly variable both within and between plots (ranging from 0 to 89%) and affected by high summer temperatures (33–35°C) in 2003. The spatial pattern characterisation of survival by Ripley’s K function showed a dominant cluster distribution. Occurrence tended to be clumped when abundance was greater than 15 seedlings, whereas for lower densities, seedlings were randomly distributed. Seedlings and herbs occupied the same sites where environmental conditions were appropriate for them to live. Spatial association occurred frequently for light and organic matter depth. Results suggested that the Scots pine seedlings in our study in central Spain preferred moderate light conditions (Global Site Factor <0.40). Thus, if this forest is to be naturally regenerated with Scots pine, the shelterwood system (whereby some mother trees are left to provide shelter for at least 5 years), along with mechanical disturbance of the seedbeds, would promote seed germination, as long as a seed source is present. However, regeneration success is affected by year-to-year conditions.  相似文献   

2.
In Mediterranean pine forests, truffles and mushrooms generate greater profits than any other woodland products. However, there are no studies on Tuber melanosporum Vittad. associated with pines. For this reason, we have carried out a study of this truffle in mountain woods with Pinus sylvestris L. and P. nigra Arnold subsp. salzmannii (Dunal) Franco, in central Spain. Two hundred and eight Tuber melanosporum burns were monitored for 7 years in five different habitats within the same geographical area. An ANOVA test confirmed significant differences in carpophore production. In higher producing habitats, pines were less abundant. We also confirmed that in 433 burns, T. melanosporum was always unequivocally associated with the root base of Quercus or Corylus trees. Similarly, 14 truffle collectors confirmed that they had never found a single burn with carpophore production associated exclusively with pines. Nevertheless, soil analyses indicated that the soil of these pine woods was very favourable to Tuber melanosporum. We therefore conclude that at present Pinus nigra salzmannii and P. sylvestris are of little interest to Tuber melanosporum culture, as they hinder carpophore production. However, this study has also confirmed that Pinus nigra salzmannii and P. sylvestris mycorrhize easily with Tuber melanosporum, both in the laboratory and in natural environments. On this basis, we propose that pines may act as transmitters of T. melanosporum, although they do not induce fruiting. As a result, the commercial cultivation of Pinus nigra salzmannii and P. sylvestris seedlings mycorrhized with Tuber melanosporum is not recommended in truffle culture at the present time.  相似文献   

3.
To study the regeneration dynamics in mature black pine (Pinus nigra Arn.) plantations in the Middle Balkan Range, Bulgaria, a total of 48 research plots (sized 0.1 ha) were established; 26 plots were established in the foothills of the mountains (foothill zone) and 22 in the main massive (mountainous zone). Pine overstorey as well as advance regeneration in subplots were recorded. Analysis of variance and regression analysis showed that in general relative stand basal area (RSBA) was the most important predictor of deciduous advance regeneration density, top height as well as age span of the regeneration layer. In the mountainous zone soil depth (SD) and the presence of mature deciduous trees (MDT) explained a significant amount of the variation in regeneration features. Lower RSBA induced by earlier thinnings resulted in higher regeneration density, larger top heights and wider age span of deciduous species. Under the pine canopy oak regeneration can persist longer compared to natural broadleaved canopies. In the studied plantations pine regeneration was confined to few scattered seedlings and regeneration groups. Based on these findings silvicultural concepts for the further management of black pine plantations in the Middle Balkan Range are discussed.  相似文献   

4.
The regeneration of Japanese black pine (Pinus thunbergii) seedlings is inhibited in a black locust (Robinia pseudoacacia)-dominated area. We examined the presence of pathogenic fungi in Japanese black pine seedlings in the area in order to determine the effect of pathogenic fungi on the inhibition of regeneration. When Japanese black pine seedlings were planted in the soil obtained from a black locust-dominated area, all of the seedlings died under low-intensity light conditions, whereas 84% of the seedlings survived in the soil obtained from a Japanese black pine-dominated area under the same light conditions. One fungus was isolated from 48.7% of the dead pine seedlings and was identified as Cylindrocladium pacificum Kang, Crous & Schoch, based on the morphological characteristics, growth, and DNA analysis. This fungus was also isolated from 50% of the dead pine seedlings in 2005 and 66.7% of the seedlings in 2006—both were planted in a black locust-dominated area. The virulence of this fungus increased under high-nitrogen and/or low-intensity light conditions. These results reveal the possibility that the soil eutrophication and shading by the black locust are conducive to a severe damping-off disease and threaten the survival and regeneration of Japanese black pine seedlings.  相似文献   

5.
Pine wilt disease, caused by the pine wood nematode (PWN, Bursaphelenchus xylophilus), is a major threat to pine forests throughout East Asia. Nonetheless, its mechanism of invasion has not yet been described in detail. To better understand the pathology of this disease, it is important to examine the distribution of PWNs within pine tissue during the course of disease development. We attempted to stain nematodes with fluorescein-conjugated wheat germ agglutinin (F-WGA) as a means to locate and track the spread of PWNs. Although PWNs proliferated on Botrytis cinerea fungus were successfully stained only on their vulvas and spicule holes, PWNs extracted from inoculated Pinus thunbergii seedlings were stained on their surface. Stainability, or the percentage of PWNs stained with F-WGA over more than half of their surface, was about 20% by 1 day after inoculation, but increased to 80% at 10 days. The stainability of PWNs extracted from a 5-cm main stem segment that included the inoculation site was less than that of PWNs extracted from other parts of the main stem farther away (i.e., those that had dispersed). These results suggest that stainability is related to dispersal activity in time. Thus, to raise the stainability of PWNs at shorter timeframes after inoculation, PWNs with more than 80% stainability were re-inoculated into pine seedlings. This resulted in more than 70% stainability from 1 to 6 days after inoculation. In F-WGA stained thin paraffin sections of pine tissue of re-inoculated seedlings, PWNs brightly fluoresced under epifluorescence and were easily detected against the dark background of pine tissue. This staining technique with F-WGA is an excellent tool for detecting PWNs in pine tissue.  相似文献   

6.
Spatial pattern of recruitment is an important factor influencing population dynamics of plant communities. The spatial pattern is determined by seed dispersal and by the spatial variability of germination and initial survival. In the process of forest expansion following farmland abandonment, mid- and late-successional species are often dispersed in pioneer forests by birds. This could result in an aggregated spatial pattern of seeds that could influence the dynamics of these species in mixed pioneer forests. In the sub-Mediterranean area, mid- and late-successional species such as Quercus pubescens (downy oak) and Fagus sylvatica (European beech) are expected to replace pioneer Pinus species. Using a point sampling method we demonstrated that beech and oak seedlings (height <2 m) have a clumped distribution in the understorey of pine. This could result from an aggregated dispersal by jays (dispersal effect) or from preferential recruitment in particular habitats (habitat effect). To test these hypotheses we proposed a statistical analysis of spatial patterns of regeneration of beech and oak. Ground cover variables (i.e. cover by rock outcrops, herbs, box shrubs, mosses or pine) did not differ significantly around seedlings as compared with random sample plots. Likewise, clumped seedlings had growth similar to isolated seedlings, thus refuting the hypothesis of preferential location in the most favourable microsites. Aggregated dispersal seems to be involved in the process of regeneration. Since beech and oak seedlings have contrasting ecological demands, we discuss the implication of this pattern for the replacement dynamics of pine by these species.  相似文献   

7.
An observational study of early seedling establishment (first 1–2 summers after emergence) was conducted in four ponderosa pine (Pinus ponderosa var. ponderosa Dougl. ex Laws.) stands east of the Cascade Range crest in central Oregon, USA. Newly emerged ponderosa pine seedlings were identified at the start of summer and were monitored through their second summer; a subsequent cohort of seedlings was identified and monitored for one summer. About 3% of the viable seed that was produced resulted in new seedlings. Mortality was substantial immediately following emergence. Most seedlings emerged without shading from understory vegetation, but few survived. Most seedlings alive after one year had emerged beneath live cover; by the end of two summers only shaded seedlings were alive. 63–85% of new seedlings initiated in clusters attributed to rodent caching, and seedlings originating in clusters remained a substantial proportion of the cohort (60%) after two seasons. Results suggest that understory vegetation in these types of stands facilitates the recruitment of ponderosa pine seedlings, most likely by buffering them against environmental stressors during the seedling establishment phase. They also reveal that seed caching rodents are highly active in seed redistribution and can exert a lasting influence on seedling recruitment.  相似文献   

8.
Micro-environmental factors viz., soil moisture and light intensity are important factors that affect natural regeneration in forests. These factors vary spatially depending on the overhead canopy density of the forest. The present study focused on studying the effect of variation of soil moisture and light intensity on natural regeneration of sal species (Shorea robusta) under different micro-environments due to overhead canopy of varying forest density. Experimental plots of 40m× 40m size were laid under different overhead canopy densities in a small sal forested watershed in the foot hills of Himalayas in Nainital District of Uttarakhand State, India. The plots were monitored on a long term basis for soil moisture at multi depths, light intensity and natural regeneration of sal. The results of the study revealed that the natural regeneration was highest under C1 (up to-0.30) canopy followed by C2 (0.30–0.50), and C3 (0.50–0.70) canopies. The C3 canopy showed the dying back of sal shoots over 4 years of study. The highest R2 value of linear regression between incremental score of plot regeneration and average soil moisture content was obtained as 0.156 for average soil moisture content during non-monsoon months at 100 cm depth. The R2 value between incremental score of plot regeneration and annual average light intensity was obtained as 0.688 which indicated that the regeneration is largely dependent on the light intensity conditions during the year. The multiple linear regression analysis between the incremental score of regeneration and the average light intensity and average soil moisture content revealed that that about 80% of variation in regeneration is explained by both the factors.  相似文献   

9.
The introduced tree species, Robinia pseudoacacia (black locust), has spread extensively in many countries. Because of its active regeneration and rapid initial growth, R. pseudoacacia has not been successfully eradicated despite many efforts. To manage this species, developing information on the growth of R. pseudoacacia as a biomass resource is desirable, and this will motivate logging and could contribute to the extermination or sustainable use of this species. In the present study, the stand volume and growth of R. pseudoacacia forests in riverbeds along the Chikumagawa River in Nagano Prefecture in Japan were examined by tree ring and stem analyses. Seven plots were established in R. pseudoacacia riparian forests of varying ages, and 611 measurements of diameter at breast height and 386 measurements of tree height were made. Stand volumes, which were estimated using equations of stem volume curve based on the results of stem analysis of 47 sample trees, were almost the same as or higher than those of native broadleaf forests in Japan. Stand volumes continued to increase for more than 20 years. Current annual increments of four plots (aged 13–22 years) indicated that it could take <5 years after regeneration for the annual increment of R. pseudoacacia forests to reach a maximum level. Growth of R. pseudoacacia was comparable to or faster than native broadleaved species in Japan, showing the possibility of short rotation harvesting.  相似文献   

10.
We evaluated the survival and growth of Abies homolepis and Picea jezoensis var. hondoensis seedlings on Mt. Ohdaigahara, where the population of sika deer (Cervus nippon) is high and an experimental fence has been in place for 13 years. No significant differences were detected in the survival of small seedlings between fenced and unfenced plots. The growth of A. homolepis was significantly higher in the fenced plot, but growth of P. jezoensis var. hondoensis did not show significant differences between fenced and unfenced plots. Seedlings of height ≤5 cm in the forest floor vegetation of the unfenced plot were probably too small for deer to find and browse, so they survived.  相似文献   

11.
Techniques for encouraging natural tree regeneration are of increasing interest to managers of Scots pine Pinus sylvestris forests. We tested prescribed fire at a management scale, with deer present or excluded, as a means of increasing rates of Scots pine forest expansion on heathland. At a semi-natural pinewood in Scotland, ten experimental blocks were established, within range of pine seed-fall. Each block comprised four, 100 m2 plots. Two plots at each site were burnt and two fenced, allowing the effects of burning on pine regeneration to be measured at two levels of deer abundance. We monitored pine seedlings, seed-fall, deer dung and vegetation for 5 years following treatment. Differences in seedling detection rates between treatments were quantified using dummy seedlings, and analyses corrected accordingly. Mean new pine seedling establishment was 9.8 times higher on burnt ground than unburnt ground (confidence intervals 3.2–30). Differences were even more pronounced in a year of high seed-fall, and following fires with low rates of spread. Establishment rates varied strongly between experimental blocks. Exclusion of deer increased establishment rates, but only in the first 2 years after fire. There was evidence that both seedling survival, and cumulative recruitment of older (over 12 months) seedlings, were improved by prescribed burning. Our results support the use of prescribed fire as a tool to promote increased Scots pine seedling establishment. This technique is likely to give strong fine-scale variation in seedling densities, and so would most suit areas where a variable spatial pattern of regeneration is sought, for landscape or naturalness reasons. Electronic supplementary material  The online version of this article (doi:) contains supplementary material, which is available to authorized users.  相似文献   

12.
The aim of this study was to determine changes in crystallinity and crystalline unit cell type of heat-treated Scots pine (Pinus sylvestris L.) and Uludağ fir (Abies nordmanniana stev. subsp. bornmuelleriana Mattf.) wood samples by means of FT-IR spectroscopic method. Heat treatment was applied on the test samples in an oven at three different temperatures (120, 150, and 180°C) and for two different periods of time (6 and 10 h) under atmospheric pressure. It was designated that crystallinity of both Scots pine and Uludağ fir wood samples increased during heat treatment depending on the duration. However, monoclinic structure in crystalline unit cells of Scots pine and Uludağ fir wood samples converted to triclinic structure when heat treated. It was estimated that monoclinic structure was dominant in the crystalline unit cell. It was established that the crystalline structure of Scots pine wood samples was more affected by heat treatment than that of Uludağ fir wood samples.  相似文献   

13.
To assess and improve methods for the development of coastal forests in Hokkaido, northern Japan, this study examined the factors affecting mortality of planted Pinus thunbergii Parlat. saplings in a region with seasonal soil frost. The needles of pine saplings at the study site turn red in late spring, suggesting xylem embolism, which can lead to death. Sapling mortality was strongly correlated with the degree of cold just after planting and with the occurrence of repeated severe winters. Volcanic ash soil had been supplied to improve the soil nutrients, and the volcanic ash soil always thawed later than the original coastal sand. Saplings in volcanic ash were significantly more damaged than those in original coastal sand. This suggests that hydraulic stress caused by cuticular transpiration under frozen soil conditions during spring may also accelerate damage due to embolisms. Hence, volcanic ash soil should not be applied to soils in regions with seasonal soil frost. A covering of woodchips can help prevent frost from reaching greater soil depths, and, indeed, fewer saplings at the study site died in areas with such coverage. Planting location can also affect pine sapling mortality. For example, shading caused by wood fencing may affect the soil freezing profile during winter; therefore, planting close to a fence should also be avoided in regions with seasonal soil frost. These findings should be used to improve the development of forests in regions with seasonal soil frost.  相似文献   

14.
Heat treatment of Pinus pinaster and Eucalyptus globulus woods, two important species in Portugal, was performed in the absence of air by steaming, inside an autoclave, for 2–12 h at 190–210°C. Mass losses increased with treatment time and temperature reaching 7.3% for pine and 14.5% for eucalypt wood. The wood behaviour with moisture was improved. The equilibrium moisture content decreased by 46% for pine and 61% for eucalypt, the dimensional stability increased (maximum anti-shrinking efficiency in the radial direction of 57 and 90% for pine and eucalypt, respectively) and the surface wettability was lowered. In relation to mechanical properties, the modulus of elasticity was little affected (maximum decrease of 5% for pine and 15% for eucalypt) but the bending strength was reduced (by 40% at 8% mass loss for pine and 50% at 9% mass loss for eucalypt wood). The variation of properties was related to treatment intensity and mass loss but significant improvements could already be obtained for a 3–4% mass loss without impairing the mechanical resistance. The response of eucalypt was higher than that of pinewood. Heat treatment of eucalypt wood shows an interesting potential to improve the wood quality for solid timber products.  相似文献   

15.
The time processes of photosynthetic induction responses to various irradiances in Korean pine (Pinus koraiensis) seedlings grown in open-light environments and in understory of forest were studied in an area near the Research Station of Changbai Mountain Forest Ecosystems, Jilin Province, China from July 15 to August 5, 1997. The results showed that at 200 μmol·m−2·s−1 photosynthetic photon flux density (PPFD) and 500 μmol·m−2·s−1 PPFD, the induction time for the photosynthetic rates of understory-grown seedlings to reach 50% and 90% steady-state net photosynthetic rates was longer than that of the open-grown seedlings. The induction responses of open-growth seedlings at 500 μmol·m−2·s−1 PPFD were slower than those at 200 μmol·m−2·s−1 PPFD, but it was the very reverse for understory-growth seedlings, which indicates that the photosynthetic induction times of Korean pine seedlings grown in the understory depended on the sunfleck intensity. Biograph: ZHOU Yong-bin (1970-), female, associate professor of Shenyang Agricultural University, Shenyang 110161, P.R. China. Responsible editor: Song Funan  相似文献   

16.
Albizia adianthifolia (Schumach) W. F. Wright, a N-fixing legume tree, has a wide distribution in Africa, in Ghana occurring in high rainfall forests and in seasonally droughted forests, and is associated in the Ghanaian forest zone with dry, infertile sites. We hypothesised that A. adianthifolia hosted different rhizobial strains in different forest types, and that these different strains would show different growth responses to moisture stress and different motility and mortality in droughted soil. Three isolates, extracted from seedlings of A. adianthifolia growing in three forest types differing in seasonal drought, were identified as Bradyrhizobium elkanii and exposed to varying levels of osmotic stress. Growth responses varied between the three strains, one of which displayed clear signs of drought tolerance. A novel approach using soil leaching columns was used to test the effects of soil pore water (in terms of neck diameter) on both the survival and movement of wet and dry forest rhizobial isolates through soil columns. The responses of the isolates were significantly different, the pore neck diameter, marginally insignificant and the drought treatment insignificant. Thus the dry forest isolate survived better in all treatments, and showed less response to the treatments, than the isolate from the wet forest. The results offer preliminary evidence that Bradyrhizobium elkanii strains from A. adianthifolia in Ghana have evolved in response to local differences in seasonal water availability. These differences could assist the selection of A. adianthifolia provenances for agroforestry or land rehabilitation.  相似文献   

17.
Most of Pinus brutia (Ten.) Holmboe forests are grazed, as silvopastoralism is well adapted in the Mediterranean environment. However, little attention has been paid to the demographic dynamic of the understorey vegetation even though it is strongly affected by the absorbed radiation. The purpose of this study was to examine the adaptation of herbaceous plant species under a Pinus brutia canopy, in Northern Greece. Monocultures of four plant groups (annual and perennial grasses, annual and perennial legumes) were sown in experimental areas of 50%, 70% and 100% light intensity. The density of annual and perennial grasses and perennial legumes were decreased by the shade cast by the Pinus brutia canopy. Population density of the annual legumes was little affected by light intensity: the annual grasses Bromus mollis L. and Lolium rigidum Gaudin; the perennial grasses Dactylis glomerata L. var. palestine, Festuca arundinacea and Agropyron cristatum (L.) Gaertn; the perennial legume Medicago sativa L. var. romana; the annual legumes Medicago lupulina L. and all the tested varieties of Trifolium subterraneum adapted well to the 50% light intensity habitat.  相似文献   

18.
Trees of the family Dipterocarpaceae are the dominant trees in Southeast Asian tropical forests where they play an important ecological role and are also important commercially. An experiment was conducted to determine the effect of ectomycorrhizal fungi on the growth of dipterocarp species in peat soils. Seedlings of Shorea pinanga were inoculated with spores of two ectomycorrhizal fungi, Pisolithus arhizus and Scleroderma sp. were grown in pots containing sterilized peat soil for 7 months. The percentage of ectomycorrhizal colonization on S. pinanga exceeded 86%. Colonization of S. pinanga roots by ectomycorrhizal fungi resulted in increased shoot height, stem diameter, number of leaves, and shoot fresh and dry weight. Survival rates of S. pinanga were greater for inoculated seedlings than control seedlings. These results suggest that inoculation of ectomycorrhizal fungi can improve the early growth of S. pinanga grown in tropical forests and that this technique will accelerate the rehabilitation of degraded dipterocarp forests.  相似文献   

19.
UGPase gene related with wood cellulose synthesis was transferred into C. acuminata using the method of Agrobacte- rium-mediated genetic transformation, and an efficient transformation system was developed for C. acuminata on the basis of evaluations of several factors affecting Agrobacterium-mediated DNA transfer rate. The highest transformation rate was achieved when pre-cultttred leaf explants were infected with an Agrobacterium culture corresponding to OD600 （0.5） for 10 min, and cultured on explant regeneration medium for three days. The results of Southern hybridization showed that genomic DNA of the kanamycin-resistant shoots to an UGPase gene probe substantiated the integration of the transgene. Transformation efficiency （6%） was achieved under the optimized transformation procedure, This system should facilitate the introduction of important useful genes into C, acuminata.  相似文献   

20.
The conversion of anthropogenic into more natural, self-regenerating forests is one of the major objectives of forestry throughout Europe. In this study, we present investigations on permanent plots with different silvicultural treatment in NE German pine stands. Management of old-growth pine stands on acidic and nutrient-poor sandy sites differs in fencing, thinning, and planting of certain tree species. The investigations were carried out on the community, population, and individual level of the pine forest ecosystems. Thus, vegetation changes, size and height of tree populations, and height increment of tree individuals were observed over a time span of 6 years. Special attention was paid to short-lived tree species such as, e.g., Frangula alnus and Sorbus aucuparia, as well as to Fagus sylvatica as one of the most typical forest tree species of Central Europe. Vegetation changes are interpreted as a consequence of natural regeneration of formerly degraded forest sites, involving an increase in nutrient availability. High browsing pressure can be considered as a key factor for the inhibition of tree seedlings and growth of saplings. Some Sorbus aucuparia individuals, however, succeeded in growing out of the browsing height also in unfenced stands. Few found specimens of Fagus sylvatica proved that this species is able to establish spontaneously on these relatively dry, acidic sites under continental climate influence. Such natural regeneration processes, also including spontaneous rejuvenation of trees, can be integrated into silviculture as passive forest conversion management. An active management like thinning of stands, planting of trees, and fencing can accelerate forest conversion with regard to height growth and species number of trees.  相似文献   

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