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1.
This paper addresses the factors that constrain the assembly of earthworm communities in temperate land use systems. An occurrence matrix containing 44 localities and 20 species was build from data sets of different studies carried out in Haute Normandie (France). The whole data set included species lists from the three main landscape units and from a range of land use types from unmanaged forests and grasslands to intensive pastures and annual crops. We used species pool, multivariate (correspondence analysis) and null model analyse to explore how species occurring in a given community are sorted from the regional species pool. Special attention was paid to identifying non-randomness in co-occurrence and morpho-ecological patterns and to describing community patterns in response to habitat constraints. Twenty species of Lumbricidae were identified in the total species pool. We found no significant effect of landscape units on species richness and density, while land use type had a significant impact on both variables, with the lowest values in crops and forests, and the highest values in grasslands. Correspondence analysis highlighted how broad land use types, soil gradients and land use intensity influence patterns of community composition and ecological coherence. Non-random patterns in guild proportion were found in a few situations, with differences between grasslands and forests. This shows that environmental filters select species at specific scales according to particular ecological traits, resulting in a convergence of these traits between species of a given community. Local species richness never exceeded nine species and species co-occurrence was significantly lower than expected by chance and much lower than described in the literature for other taxa. Although no significant patterns of limiting similarity were observed at the scale of our study, we conclude that earthworm communities are also highly structured by competition. Our study thus agrees with the idea that both habitat and competitive constraints operate simultaneously to determine how community assembly takes place.  相似文献   

2.
Disturbances favour plant invasions because they provide a pulse of resources for seedling establishment and growth. However, little attention has been paid to the importance of the different disturbance regimes in the extent of invasion. In this paper we asked to what extent invasion depends on spatial and temporal patterns of disturbance. By using the spatially explicit model Fateland, we simulated how Cortaderia selloana, a large-wind dispersed perennial grass invading Mediterranean grasslands, responds to different medium-term (up to 55 years) disturbance regimes. The different scenarios combine disturbance frequency, extent of disturbance and temporal and spatial disturbance variability, and they roughly mimic real situations of habitat management and disturbances in cultural landscapes.The model predicted that the proportion of invaded area would increase with disturbance frequency. The extent of the disturbance overcame the importance of frequency. Aggregated disturbances randomly distributed in space had a more positive effect on invasion than scattered disturbances at repeatedly fixed locations. Whether the disturbances regularly occurred in time or were irregularly distributed did not influence the percentage of invaded landscape. Our simulation analysis unambiguously supports the hypothesis that disturbances are essential for the persistence and expansion of Cortaderia sellona. The high amount of seed rain from a few individuals (e.g., gardens) make this species very sensitive to landscape changes produced by different disturbance patterns (e.g., land abandonment, habitat management or rubbish dumping). Our results can contribute to redirect land management options.  相似文献   

3.
Long transient times in response to decreasing habitat area and increasing isolation may cause the present plant species distribution to reflect the historical rather than the present landscape configuration, resulting in a so-called extinction debt. To investigate how plant species richness is shaped by both present and historical landscape configuration and local environmental conditions in fragmented calcareous grasslands, 64 sites in Southern Belgium were surveyed for their total species richness and environmental characteristics. Species were subdivided in specialist and generalist groups and the former were clustered into emergent groups (EGs) according to 16 relevant life-history traits. Four specialist emergent groups were derived: orchids, rosette species, annuals and half-rosette species. Both specialist and generalist species richness increased with present fragment area, while present fragment connectivity only affected the number of specialist species. This trend was maintained for most EGs, although habitat area was mostly more important than connectivity. All species groups responded to an insolation gradient, except EGs of rosette and annual species. Differential response to fragmentation among emergent groups could be interpreted in terms of dispersal and persistence traits. No relation between species richness and historical area or connectivity was found, making an extinction debt unlikely. Hence, present-day variation in plant species diversity seems to be no legacy of past landscape configuration. Our results clearly indicate that plant species may be quickly lost in response to new fragmentation events. To restore plant diversity, management should focus both on mitigating landscape fragmentation and restoring habitat quality.  相似文献   

4.
The impact from transportation corridors on surrounding habitat often reaches far beyond the edge of the corridor. The altered disturbance regime in plant communities along corridor edges and vehicle traffic facilitate the spread and establishment of invasive non-native plant species. We compared the frequency of non-native plant species along highways and railways and the ability of these species to invade grasslands and dense forests along these corridors. We measured the frequency of several non-native plant species along transects 0-150 m from the edge of highways and railways in grasslands and forests, as well as at control sites away from corridors. Both transportation corridors had higher frequency of non-native species than respective control sites. Grasslands had higher frequency of non-native species than forested habitats, but the frequency did not differ between the highways and the railways. The frequency of non-native species in grasslands along highways and railways was higher than at grassland control sites up to 150 m from the corridor edge, whereas the frequency in forested habitats along corridors was higher than at forested control sites up to only 10 m from the corridor edge. There was a significant decrease in the frequency of non-native species with increasing distance from both corridors in the forest, while grasslands showed no significant change in non-native species frequency with distance from corridors. This suggests that corridor edges and grassland habitats act as microhabitats for non-native species and are more prone to invasion than forests, especially if disturbed. Our results emphasize the importance of minimizing the disturbance of adjacent plant communities along highways and railways during construction and maintenance, particularly in grassland habitats and in areas sensitive to additional fragmentation and habitat loss.  相似文献   

5.
I apply a comparative, functional group approach to coastal sandplain grassland taxa in order to examine whether rare plant species share certain aspects of rarity and life history characters that are distinct from their more common, co-occurring congeners in these habitats. I compiled a comparative data set containing 16 variables describing biogeographic distributions, level of imperilment, habitat specialization, vegetative versus sexual reproduction, seed dispersal, and dormancy of 27 closely-related pairs of plant species that contrast in their abundance (infrequent versus common) in coastal sandplain grasslands. Frequent and infrequent species were paired within genera (or closely related genera) and thus distributed equivalently across families to control for phylogenetic bias. Paired comparisons revealed that infrequent species were intrinsically rarer range-wide, and exhibited a narrower range and more habitat specialization than their common relatives. A classification tree distinguished infrequent species from common species on the basis of higher habitat specialization, larger seed size, smaller plant height, less reliance less on vegetative (colonial) reproduction, and tendency toward annual or biennial life history. Research and management steps to reduce competition from larger-satured, colonial, perennial species are recommended for these infrequent species. Basic research involving more species and more data on ecophysiological characters, demography, and competitive interactions are needed to identify critical life history traits that will influence responses to particular management regimes.  相似文献   

6.
Calcareous grasslands are an important habitat for floral and faunal communities in the UK and Europe. Declines due to changes in management, scrub invasion and agricultural improvement have left much of the remnants of this habitat in a degraded and fragmented state. Grazing, by cattle or sheep, is one of the main management practices used to maintain and improve the floral and faunal quality of calcareous grassland. The long-term impacts of different grazing regimes, however, are poorly understood, particularly in terms of the invertebrate communities. This study contrasted the impacts of recently introduced and long-term sheep or cattle grazing on beetle communities present on one of the largest areas of calcareous grassland in Europe, the Salisbury Plain military training Area, UK. No effects of grazing management on beetle abundance, species richness or evenness were found, but plant diversity and overall percentage cover of grasses did influence beetle diversity. Proportions of the total number of individuals and overall species richness within beetle guilds (predatory, phytophagous, flower/seed feeders, root feeders and foliage feeders) were strongly influenced by both the duration and type of grazing animal. At the species level, beetle community structure showed significant differences between ungrazed, long-term cattle and long-term sheep grazing treatments. Changes in plant community structure were found to influence beetle community structure. The significance of these results is discussed in terms of the long-term impacts of grazing on beetle community structure, and the benefits of different grazing regimes for the conservation management of calcareous grasslands.  相似文献   

7.
Conservation biology often relies on the protection of (semi)natural habitat remnants. However, the ever increasing human population is taking over natural resources and habitats. Here, contrary to most other studies, we ask how human-associated severe changes in the environment can be used to enrich local biodiversity. We tested if industrial activity (gravel excavation) leads to the creation of habitats that support grassland butterflies and how these areas add to the richness of local species when compared to typical semi-natural habitats (grasslands). We also identified key factors affecting the richness, abundance, diversity and commonness of butterfly species to provide practical recommendations. Species richness, diversity index and the occurrence of rare species were higher in gravel-pit shores than in grasslands. The richness of butterfly species and their abundance were positively affected by the richness of plant species, shrub density and age of the gravel-pit but negatively by the cover of water reservoirs in the surrounding area and the isolation of gravel-pits from grasslands. Butterfly diversity was positively influenced by the richness of plant species and proximity of human settlement but negatively by area of the shore and isolation. Our study is the first one to show the high value of gravel-pits for the conservation of butterflies. We recommend the inclusion of gravel-pits in a system of ecological networks and management of their surroundings to improve the colonization rate of rare species. We suggests that directing interest to the possible positive effects of industrial development on biodiversity may support conservation efforts.  相似文献   

8.
探究干旱半干旱区土地利用结构动态变化特征，评估并预测生境质量，可为区域生态规划和恢复提供决策性依据。本研究以搭载OLI和ETM+传感器遥感影像和气象数据、社会经济数据等为基础，采用土地利用转移流概念、InVEST模型和CA-Markov模型研究2000-2030年宁夏中部干旱区土地利用和生境质量时空变化特征及规律，对未来情境进行模拟和预测。研究结果表明：宁夏中部干旱区土地系统结构变化与生态建设规划高度一致，具有黄河流域两侧集中分布的特征，土地系统综合动态度逐期减小，活跃度具有稳中有变、部分较活跃的特征。土地系统信息熵特征值呈逐年降低趋势。随着时间变化土地系统受人类活动干扰强度低，自我调整程度高，修复能力强，系统向稳定状态转化。2000-2015年研究区生态环境质量呈“U”型特征，以优秀等级为主。2015-2030年土地系统结构将发生显著变化，生境质量优秀、差和良好等级面积逐渐扩大，生态系统呈现稳定向好的趋势。本研究的多模型集成应用可为区域土地规划和生态恢复建设提供理论依据和支撑。  相似文献   

9.
Both disturbance history and previous land use influence present-day vegetation and soils. These influences can have important implications for conservation of plant communities if former disturbance and land use change species abundances, increase colonization of nonnative plant species or if they alter soil characteristics in ways that make them less suitable for species of conservation interest. We compared the plant species composition, the proportion of native and nonnative plant species, and soil biogeochemical characteristics across seven dominant land use and vegetation cover types on the outwash sandplain of Martha’s Vineyard that differed in previous soil tillage, dominant overstory vegetation and history of recent prescribed fire. The outwash sandplain supports many native plant species adapted to dry, low nutrient conditions and maintenance of native species is a management concern. There was broad overlap in the plant species composition among pine (Pinus resinosa, P. strobus) plantations on untilled soils, pine plantations on formerly tilled soils, scrub oak (Quercus ilicifolia) shrublands, tree oak (Q. velutina, Q. alba) woodlands, burned tree oak woodlands, and sandplain grasslands. All of these land cover categories contained few nonnative species. In contrast, agricultural grasslands had high richness and cover of nonnative plants. Soil characteristics were also similar among all of the woodland, shrubland and grassland land cover categories, but soils in agricultural grasslands had higher pH, extractable Ca2+ and Mg2+ in mineral soils and higher rates of net nitrification. The similarity of soils and significant overlap in vegetation across pine plantations, scrub oak shrublands, oak woodlands and sandplain grasslands suggests that the history of land use, current vegetation and soil characteristics do not pose a major barrier to management strategies that would involve conversion among any of these vegetation types. The current presence of high cover of nonnative species and nutrient-enriched soils in agricultural grasslands, however, may pose a barrier to expansion of sandplain grasslands or shrublands on these former agricultural lands if native species are not able to outcompete nonnative species in these anthropogenically-enriched sites.  相似文献   

10.
The conservation of biodiversity critically depends on the value that humans attach to it. Apart from an ecological and economic value, an aesthetic value has often been assigned to biodiversity. However, it is not known whether lay people appreciate the diversity of species and not just certain individual species or nature as a whole. We studied in a series of experiments and field studies people’s perception and appreciation of species diversity. We presented meadow-like arrays of different species richness and evenness but random species composition to lay people and asked them to estimate plant species richness and rank the arrays by attractiveness. The experiments were complemented by two larger-scale field studies using natural meadows. Both in the experiments and the field studies the mean perception of species richness by people increased with true species richness, but was slightly overestimated at low and increasingly underestimated at high diversity levels. Lay people’s aesthetic appreciation of both the experimental grassland arrays and the natural meadows increased with true species richness. Communities consisting of the same number of species were perceived to be more species-rich and were appreciated more when their evenness was high. Our results demonstrate that plant diversity in itself is attractive to humans. The current reduction of the diversity of grasslands due to intensification of management may thus reduce the attractiveness of regions where grasslands are a dominant feature of the landscape. This could have negative consequences for tourism and may add an economic argument for the conservation of biodiversity in grasslands.  相似文献   

11.
The effectiveness of measures targeted at the restoration of populations of endangered species in anthropogenically dominated regions is often limited by a combination of insufficient restoration of habitat quality and dispersal failure. Therefore, the joint prediction of suitable habitat and seed dispersal in dependency of management actions is required for effective nature management. Here we demonstrate an approach, which links a habitat suitability and a seed dispersal model. The linked model describes potential species distribution as a function of current species distribution, species-specific dispersal traits, the number of successful dispersal events, dispersal infrastructure and habitat configuration. The last two variables were related to water management actions. We demonstrate the applicability of the model in a strategy analysis of hydrological restoration measures for a large fen area in which still numerous endangered plant species grow.With the aid of the linked model, we were able to optimise the spatial planning of restoration measures, taking into account both the constraints of water management practices on abiotic restoration and the effects of habitat fragmentation on dispersal. Moreover, we could demonstrate that stand-alone habitat suitability models, which assume unlimited dispersal, may considerably overestimate restoration prospects. For these reasons, we conclude that linked habitat suitability and dispersal models can provide useful insights into spatially differentiated potentials and constraints of nature restoration measures targeted at the sustainable conservation of endangered plant populations whose habitats have been deteriorated due to undesirable effects of land and water management on abiotic conditions. These insights may contribute to the design of cost-effective nature restoration and conservation measures.  相似文献   

12.
Since the middle of the last century agricultural intensification within Europe has led to a drastic decline in the extent of botanically diverse grasslands. Whilst measures to enhance the diversity of agriculturally-improved grasslands are in place, success has often been limited. One of the primary factors limiting success is the paucity of sources of propagules of desirable species in the surrounding landscape. The restoration of two contrasting grassland types (lowland hay meadow and chalk grassland) was examined using a replicated block experiment to assess the effectiveness of two methods of seed application (hay strewing and brush harvesting) and two methods of pre-treatment disturbance (power harrowing and turf stripping). The resulting changes in botanical composition were monitored for 4 years. Seed addition by both methods resulted in significant temporal trends in plant species composition and increases in plant species richness, which were further enhanced by disturbance. Power harrowing increased the effectiveness of the seed addition treatments at the lowland hay meadow site. At the chalk grassland site a more severe disturbance created by turf stripping was used and shown to be preferable. Whilst both hay strewing and brush harvesting increased plant species richness, hay strewing was more effective at creating a sward similar to that of the donor site. Soil disturbance and seed application rate at the recipient site and timing of the hay cut at the donor site are all factors to be considered prior to the commencement of restoration management.  相似文献   

13.
The conservation of biological diversity requires an integrated approach covering the ecological demands of a multitude of species. Integration may be achieved by focusing on a careful selection of target species, which is rare in practice. Calcareous grasslands offer a case in point. Although they harbour a high diversity of both plant and insect species, in management the emphasis is placed on the flora. This results in an underestimation of, notably, the importance of structural heterogeneity in the vegetation. As an apparent consequence, conservation management in the Netherlands has been much more successful for the flora than for butterflies. In contrast, Germany shows promising efforts to integrate both plant and animal species in conservation management and landscape planning. The main constraints for a successfully integrated conservation management presently consist of a limited availability and exchange of information, and an insufficient organisation of research and management at an international level.  相似文献   

14.
Although spatial scale is important for understanding ecological processes and guiding conservation planning, studies combining a range of scales are rare. Habitat suitability modelling has been used traditionally to study broad-scale patterns of species distribution but can also be applied to address conservation needs at finer scales. We studied the ability of presence-only species distribution modelling to predict patterns of habitat selection at broad and fine spatial scales for one of the rarest mammals in the UK, the grey long-eared bat (Plecotus austriacus). Models were constructed with Maxent using broad-scale distribution data from across the UK (excluding Northern Ireland) and fine-scale radio-tracking data from bats at one colony. Fine-scale model predictions were evaluated with radio-tracking locations from bats from a distant colony, and compared with results of traditional radio-tracking data analysis methods (compositional analysis of habitat selection). Broad-scale models indicated that winter temperature, summer precipitation and land cover were the most important variables limiting the distribution of the grey long-eared bat in the UK. Fine-scale models predicted that proximity to unimproved grasslands and distance to suburban areas determine foraging habitat suitability around maternity colonies, while compositional analysis also identified unimproved grasslands as the most preferred foraging habitat type. This strong association with unimproved lowland grasslands highlights the potential importance of changes in agricultural practices in the past century for wildlife conservation. Hence, multi-scale models offer an important tool for identifying conservation requirements at the fine landscape level that can guide national-level conservation management practices.  相似文献   

15.
Habitat selection by East Caucasian tur (Capra cylindricornis), a species of global conservation concern, was examined in relation to terrain, climate and degree of human disturbance using a Geographical Information System and logistic regression. The study area was in the part of the Greater Caucasus of Georgia, where the species protection was not enforced. Two models of tur habitat requirements were obtained: one model at a scale of 20 × 20 m plots, and the other one at a scale of different habitat fragments made up of 20 × 20 m plots identified by the first model at its optimal cut-off value. The second model refined the first one.The first model suggested that the probability of a 20 × 20 m plot being part of tur habitat was positively correlated with slope, distances to roads and livestock summer camps, and negatively correlated with human population density and annual rainfall. The probability had a bell-shaped correlation with elevation, reaching its maximum at 3008.4 m. The second model suggested that a fragment of a land made up of 20 × 20 m plots with optimal characteristics for tur occurrence was more likely to contain tur if the area of the fragment was larger and its distance to the nearest area where tur occurred was shorter.The results show that the occurrence of East Caucasian tur is affected by climate, terrain, human disturbance and habitat fragmentation, and can be predicted regardless of seasonality in the species movements. These models can be applied to the management of the species and its habitat in the areas of the Caucasus that lie at >1000 m asl and have an annual rainfall >600 mm, and where the species protection is not enforced.  相似文献   

16.
The provision of recreational opportunities is one of the important human goals of marine protected areas. However, as levels of recreational use increase, human disturbance is likely to cause significant detrimental effects upon wildlife. Here we evaluate the best managing options to mitigate the impact of sea-based tourism on the foraging activity of an endangered population of European shags, Phalacrocorax aristotelis, in a coastal marine protected area (Cíes islands, north-western Iberia). Boat disturbance elicited a characteristic avoidance behavior that resulted in a substantial reduction in foraging activity as levels of boat use increased. Moreover, boats excluded shags from the best feeding areas, resulting in higher densities of foragers in areas of little boat traffic. We used a behavioral model to explore the effects of managing strategies aimed at reducing the impact of boats on the foraging activity of shags. Our model suggested that in low boat disturbance scenarios limiting the number of boats using the reserve would be a better management option than habitat protection (i.e. the establishment of set-aside areas free of boat traffic). On the contrary, when boat disturbance levels are high the protection of habitat is recommendable, even if spatial variation in habitat quality is unknown or poorly assessed. Our study stresses the point that management strategies to minimize disturbance to foraging seabirds may depend on the spatial overlap between sea-based recreational activities and foraging seabirds and the spatial variation in marine habitat quality for seabirds.  相似文献   

17.
东祁连山不同干扰生境草地土壤养分时空变化特征   总被引：2，自引：1，他引：2  
为探究东祁连山不同干扰高寒草地土壤养分及其生态化学计量学特征时空变化特征规律。研究了2017年8月在东祁连山不同干扰高寒草地土壤养分含量及其生态计量学特征,以及与2003年和2011年在相同地点、相同研究方法对该地区的土壤理化性质进行比较。结果表明:(1)在同一干扰生境不同土层下,土壤pH、电导率、含水量、土壤全量和速效养分均随着土层深度增加而逐渐减小,变化各异;在同一土层不同干扰生境下土壤养分间变化差异各不相同。(2)不同干扰生境土壤C/N为7.64~18.21,与我国陆地土壤平均值(10~12)基本接近,土壤C/P和N/P分别为109.16~144.79和7.52~15.65,远高于中国陆地土壤的C/P平均值52.7和N/P平均值3.9。(3)主成分分析结果表明,土壤理化性质(pH、电导率、含水量、全磷、全钾、速效磷、有机质、有机碳和速效氮)和土壤生态化学计量比(C/N、C/P和N/P)可以敏感的反映出不同干扰对草地土壤质量影响状况。土壤质量综合评价可知,不同干扰生境下草地土壤质量水平差异较大,多年生草地土壤质量较高,围栏外草地质量最差,在0-20,20-40cm土层中,均表现为多年生围栏内一年生围栏外。(4)与2003年和2011年相比较,4种不同干扰生境高寒草地土壤pH和含水量,总体呈上升的趋势,土壤有机质、全氮和全磷含量则呈现出明显下降趋势。综上,放牧干扰、人为种植对东祁连山4种干扰生境草地土壤养分含量有很大影响,为此应对该地区草地的利用和保护给予足够的重视,合理利用和科学管理草地。  相似文献   

18.
The construction of habitat models is a repeatable technique for describing and mapping species distributions, the utility of which lies in enabling management to predict where a species is likely to occur within a landscape. Typically, habitat models have been used to establish habitat requirements for threatened species; however they have equal applicability for modelling local populations of common species. Often, few data exist on local populations of common species, and issues of abundance and habitat selection at varying scales are rarely addressed. We provide a habitat suitability model for the common wombat (Vombatus ursinus) in southern New South Wales. This species is currently perceived as abundant throughout its extensive range across temperate regions of eastern Australia, yet little factual survey data exist and populations appear under threat. We use wombat burrows to reflect habitat selection and as our basis for ecological modelling. We found that environmental variables representing proximity to cover, measures of vegetation and proximity to watercourses are important predictors of burrow presence. Extrapolation of habitat models identified an abundance of habitat suitable for burrows. However, burrows in many suitable areas were abandoned. Our estimate of the population size was similar to the total annual mortality associated with road-kill. Theoretically, given the availability of suitable habitat, common wombat populations in the region should be thriving. It seems likely that this area once supported a much higher number of wombats; however limiting factors such as road mortality and disease have reduced the populations. The persistence of wombats in the study region must be supported by migration from other populations. Our findings challenge the perception that wombats are currently common and not in need of monitoring, suggesting that perceptions of abundance are often clouded by socio-political motives rather than informed by biological and ecological factors.  相似文献   

19.
Earthworms are widespread soil organisms that contribute to a wide range of ecosystem services. As such, it is important to improve our knowledge, still scanty, of the factors that drive the assembly of earthworm communities. The aim of the present study was to conjointly evaluate the effects on the assembly of earthworm communities of i) soil properties (texture, organic matter content, and pH), ii) grassland management (grassland age, livestock unit, and type of fertilization), iii) landscape diversity (richness, diversity of surrounding habitats, and grassland plant diversity), and iv) presence of hedgerows. The study was conducted in temperate grasslands of Brittany, France. Earthworms were sampled in 24 grasslands and, in three of these grasslands, they were sampled near a hedgerow or near a ditch (control without a hedgerow). Soil properties explained the larger portion of the variation in the earthworm community parameters compared to grassland management or landscape diversity. The increase in soil organic matter content and pH were the most favorable factors for earthworm abundance and biomass, in particular for endogeic species. Regarding grassland management, the increase in the livestock unit was the most damaging factor for earthworm communities, in particular for the anecic earthworm biomass and endogeic species richness. Surprisingly, landscape diversity negatively affected the total earthworm abundance and epigeic earthworm biomass, but it was related to an increase in the epi-anecic species. At a finer scale, we also demonstrated that the presence of hedgerows surrounding grasslands enhanced earthworm species richness, especially within the epigeic and anecic ecological categories. This study highlights that the earthworm ecological categories respond specifically to environmental filters; further studies need to be conducted to elucidate the factors that drive the assembly of earthworm communities at this ecological category level. We recommend that policymakers should act on landscape management to favor earthworm diversity in order to improve the ecosystem services they drive.  相似文献   

20.
The consideration of economic costs can dramatically influence the cost-effective allocation of resources spent on the conservation of biodiversity. In the face of ecological and economic data scarcity, simple rules are needed for the design of cost-effective habitat networks. A conceptual semi-analytical ecological-economic model is developed that is based on metapopulation theory and takes into account that the costs of conservation may be spatially heterogeneous and changing in time. The model is used to study trade-offs between amount, spatial connectivity and temporal continuity of habitat. The cost-effective levels of habitat connectivity and continuity are determined as functions of species attributes and economic parameters like the distribution of conservation costs. The results are relevant for the design of cost-effective conservation instruments.  相似文献   

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