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1.
Lodgepole pine is native to western North America, but it is also planted as a fast-growing alternative to Scots pine in Sweden. The production of these two species, when grown as native and as exotic species, was compared in a transcontinental two-species provenance experiment. The tests were planted in 1986 on five sites in northwestern Canada and two sites in Sweden, and included full-sib families, half-sib families, seed orchard collections and natural stand seed collections of both species. After 25 years, lodgepole pine produced 48% more volume (m3ha?1) and had 27% higher survival than Scots pine at one Swedish site, and had similar volume production and survival at a second. In the five Canadian sites, Scots pine produced on average 22% more volume than lodgepole pine. The variation between sites was, however, large. This higher volume of Scots pine in Canada could be due to higher survival (+28%) and less frequent damage; but higher top height for lodgepole pine in Canada indicated higher potential productivity. The results indicate that an exotic species may produce more than the native species, possibly thanks to higher survival, but it is also possible to increase production with successful population selection of the native species.  相似文献   

2.
Between April and September 1994, six plots within Aberfoyle Forest in Scotland, three of Scots (Pinus sylvestris) and three of lodgepole pine (Pinus contorta), were surveyed for spiders and harvestmen, as these are predators of several forest invertebrate pests.A total of 64 species of spider were trapped in Scots and lodgepole pine; 56 in Scots pine and 32 in Lodgepole pine. The most abundant spiders, both in Scots and Lodgepole pine, were Linyphiidae.Scots pine plots were floristically more diverse and structurally more complex than lodgepole pine plots. More species of spiders and harvestmen (midsummer only) were found at ground level in Scots pine than lodgepole pine plots. More species and a greater numbers of spiders were found in the canopy of Scots pine than in that of lodgepole pine. However, no differences in the abundance of harvestmen were found between the canopies of Scots and lodgepole pine. Spider species diversity was greatest in Scots pine.  相似文献   

3.
Seedlings of different provenances of Scots pine (Pinus sylvestris L.), lodgepole pine (Pinus contorta Dougl., var. latifolia Engelm.) and Norway spruce (Picea abies (L.) Karst.) were planted in three Scots pine shelterwoods (125, 65 and 43 stems ha−1) and a clear-cut, all in northern Sweden. The sites were mounded and planting took place during 2 consecutive years (1988 and 1989). The solar radiation experienced by the individual seedlings was determined using a simulation model. Height development of the seedlings was examined during their first 6 years after planting. During the final 3 years of the study, height growth of Norway spruce was relatively poor, both in the shelterwoods and the clear-cut area. Height growth of lodgepole pine was significantly greater than that of Scots pine, both in the shelterwoods and the clear-cut. In contrast to Norway spruce, Scots pine and lodgepole pine displayed significantly greater height growth in the clear-cut than in the shelterwoods. For all three species in the shelterwoods, regression analyses showed that height growth was more strongly correlated with the distance to the nearest tree than with the amount of radiation reaching the ground, i.e. growth was reduced in the vicinity of shelter trees. Therefore, we conclude that the significant reduction in height growth of seedlings of Scots pine and lodgepole pine in Scots pine shelterwoods was partially caused by factors associated with the distance to the nearest shelter tree. Because the substrate was a nitrogen-poor sandy soil, we suggest that root competition for mineral nutrients, especially nitrogen, accounts for the reduction in height growth.  相似文献   

4.
Needle samples of six provenances each of lodgepole pine (Pinus contorta Dougl. var. latifolia) and Scots pine (Pinus sylvestris L.), originating from latitudes 55 to 68 degrees N in western Canada and northern Sweden, were collected during the autumn and subjected to freezing temperatures in the range of -8 to -29 degrees C on three occasions in September and October. Needle injury was assessed by two different methods: visual assessment and chlorophyll a fluorescence. Chlorophyll a fluorescence data showed a highly significant correlation with the visual assessments of injury, indicating that the technique can be used as a simple, non-destructive and objective measure for rapid detection of freezing injury and for ranking of needle materials with respect to development of cold acclimation. The analyses showed that, during the autumn, lodgepole pine needles were more hardy and acclimated to low temperatures earlier than Scots pine needles.  相似文献   

5.
One-year-old Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and lodgepole pine (Pinus contorta Dougl.) seedlings were grown for 17 weeks in 100-cm deep, 7.8-liter containers. Two Douglas-fir provenances, one from a wet and one from a dry site in coastal British Columbia, and two lodgepole pine provenances, one from a wet and one from a dry site in interior British Columbia, were grown in wet (522% water content) or dry (318% water content) peat/vermiculite soil in a factorial design. Each container was sealed so that water loss occurred only through the seedling. Five harvests were made at three to five week intervals and water use, dry matter increment, root length and root weight were determined at each harvest. Stomatal conductance and shoot water potentials were measured during the last 12 weeks of the experiment. Lodgepole pine seedlings had greater dry matter production, water use, stomatal conductance and new root length than Douglas-fir seedlings. New root weight of lodgepole pine seedlings exceeded that of Douglas-fir seedlings during the last five weeks of the experiment, and specific root length (root length per unit root weight) of new roots was higher for lodgepole pine seedlings throughout the experiment. Douglas-fir seedlings showed higher water use efficiency (WUE) than lodgepole pine seedlings, and both species showed higher WUE in the dry soil treatment. Douglas-fir seedlings had lower water potentials and higher water uptake rates per unit of new root length than lodgepole pine seedlings, although water uptake rates per unit of root dry weight showed little difference between species. Soil water treatment influenced specific root length of new roots, water uptake per unit of new root length, and WUE in Douglas-fir seedlings more than in lodgepole pine seedlings.  相似文献   

6.

The effect of site fertility, spacing and mode of regeneration on the survival and stand development of lodgepole pine, Scots pine and Norway spruce was studied in a series of experiments comprising 22 study areas in Finland. After 13-14 yrs, lodgepole pine had a mean survival of 68% in planted and 61% in seeded plots, while Norway spruce had the highest (92%) and Scots pine the second highest survival (82%). The survival of planted lodgepole pine was better the wider the spacing. Best survival was achieved on subdry and dry sites, both with planting and with direct seeding. Dominant height was not affected by spacing, but both basal area and volume at the age of 13-14 yrs were significantly higher the denser the spacing. The average difference in the value of the estimated site index H 50 was slightly under 3 m for the superiority of lodgepole compared with Scots pine. Seeding resulted in site indices almost as high as those of lodgepole planting.  相似文献   

7.
The patterns of current‐year shoot, needle and terminal bud elongation in seedlings of three Scots pine (Pinus sylvestris L.) and three lodgepole pine (Pinus contorta Dougl. var. latifolia Engelm.) provenances were compared during the third and fourth growing seasons after planting. Lodgepole pine produced longer shoots and buds than did Scots pine, mainly because lodgepole pine formed more stem units and elongated at a faster rate. Stem unit length and the duration of shoot and bud elongation differed relatively little between species and provenances. Lammas or polycyclic growth occurred in some lodgepole pine provenances, but not in any Scots pine provenance, and was associated with enhanced shoot elongation. Needle elongation commenced earlier, proceeded at a faster rate, and was greater in lodgepole pine than in Scots pine, but ceased about the same time in all species and provenances. The heat sum required to attain 50% of final length was lower for shoots and needles in lodgepole pine than in Scots pine, and for shoots in northern provenances than in southern ones. Mitotic activity in the apical meristem of the terminal bud, which occurred less than one week after the seedlings were free from snow, started and ceased about the same time in each species, but was higher in lodgepole pine than in Scots pine early in the shoot elongation period.  相似文献   

8.
Abstract

The objective of this study was to compare volume growth and external timber quality properties of lodgepole pine (Pinus contorta Dougl. var. latifolia) stands planted in different spacings (1.41×1.41, 2.00×2.00, 2.83×2.83, 4.00×4.00) in southern Sweden. In northern Sweden lodgepole pine has been grown for a long time and shows superior volume production compared to Scots pine (Pinus Sylvestris L.). The Swedish Forestry Act restricts establishment of lodgepole pine plantations in southern Sweden. However, it is important to increase the knowledge about lodgepole pine and its potential use also in this part of the country. Two experiments in southern Sweden were studied. Each trial was designed as a random block experiment with two blocks. Both investigated stands had an age of 23 years at the time of assessment. The mean diameter and the volume production differed significantly between the spacings. The highest volume production was found in the narrowest spacing, 313% compared to the widest spacing. Significant differences between spacings were also found concerning external quality traits. The frequency of dominant and co-dominant trees without defects was 24% in the most open spacing compared to 46% in the narrowest one. Under current circumstances approximately 2500 seedlings/ha in the initial stand seems to be a reasonable compromise between growth, diameter development and timber quality.  相似文献   

9.
Abstract

Pine weevil (Hylobius abietis L.) damage to seedlings after overstorey removal was investigated in a survey study in six shelterwoods in the south–central part of Sweden. The shelterwoods predominantly consisted of Scots pine, except at one site where the shelter trees mainly consisted of Norway spruce. Before final cutting, 10 plots were laid out at each site and measurements of shelter trees and marked seedlings were taken. The seedlings were examined during the 2 years after final cutting. The study showed that removal of shelter trees increases the risk of severe damage by pine weevil and the variable that was most strongly correlated with the risk was the seedling root collar diameter. Both Scots pine and Norway spruce seedlings were severely damaged by pine weevil, and most of the feeding occurred during the first year after cutting. The amount of debarked area was significantly larger for Scots pine than for Norway spruce seedlings. Vitality (growth of the leading shoot before final cutting) of the seedlings also affected the probability of damage. Seedlings with high vitality were less damaged by pine weevil than seedlings with low vitality. For Scots pine the shelterwood density before final cutting was correlated to the intensity of pine weevil feeding after cutting. In conclusion, after the final cutting of a pine or spruce shelterwood, pine weevils will probably invade the area. To avoid serious damage, Norway spruce and Scots pine seedlings should have reached a diameter of at least 10–12 mm.  相似文献   

10.
Abstract

Tree vitality and height of lodgepole pine (Pinus contorta var. latifolia) originating from the six Swedish seed orchards and native Scots pine (Pinus sylvestris L.) were estimated in three 8-yr-old test plantations established in the Komi Republic (north-west Russia). A randomized row-plot design with 9–13 replicates of each entry was used. In general, the Scots pine tree vitality was better than that of lodgepole pine but these differences were significant only for seed sources of southern origins Larslund, Rumhult, and Österby. On the contrary, the lodgepole pine height growth was faster (6–21%) except for seed sources of southern origins Rumhult and Österby. In comparison with the native Scots pine, lower mean heights (4–10%) were recorded for these seed sources. Among the lodgepole pine seed sources the northern origins had better vitality, while the effect of latitude on the tree height was insignificant.  相似文献   

11.
Scots pine and different provenances of lodgepole pine were compared in regard to root characteristics and the distribution of biomass within individual trees. Tap root formation was more frequent and the root/shoot ratio was higher on Scots pine than on lodgepole pine but significant variation of these traits was found between provenances of planted lodgepole pine. A non‐destructive method for estimation of root biomass was developed.  相似文献   

12.
The mountain pine beetle Dendroctonus ponderosae Hopkins is endemic to lodgepole pine, Pinus contorta var. latifolia Engelmann, forests in western Canada. However, the current beetle epidemic in this area highlights the challenges faced by forest managers tasked with prioritizing stands for mitigation activities such as salvage harvesting and direct control methods. In western Canada, the operational risk rating system for mountain pine beetle is based on biological knowledge gained from a rich legacy of stand-scale field studies. Due to the large spatial (millions of hectares affected) and temporal (over 10 years) extents of the current epidemic, new research into large-area mountain pine beetle processes has revealed further insights into the landscape-scale characteristics of beetle infested forests. In this paper, we evaluated the potential for this new knowledge to augment an established system for rating the short-term risk of tree mortality in a stand due to mountain pine beetle. New variables explored for utility in risk rating include direct shortwave radiation, site index, diameter at breast height, the temporal trends in local beetle populations, Biogeoclimatic Ecosystem Classification and beetle–host interaction variables. Proportional odds ordinal regression was used to develop a model for the Vanderhoof Forest District in west-central British Columbia. Prediction on independent data was assessed with the area under the receiver operator curve (AUC), indicating good discriminatory power (AUC = 0.84) for predicting levels of mountain pine beetle-caused pine mortality.  相似文献   

13.
In boreal forests of western Canada, lodgepole pine (Pinus contorta Dougl. ex. Loud.) and white spruce (Picea glauca (Moench) Voss) often grow together with numerous tall shrubs such as green alder (Alnus crispa (Ait.) Pursh) and little-tree willow (Salix spp.). In an area south of Grande Prairie, Alberta, Canada, we examined the effects of shrubs, herbs and other trees on nutrient and light availability and growth of white spruce and lodgepole pine. For white spruce the best competition measure (tested against volume increments of the past 3 years) was visually estimated % ground cover times the height of the competitor (VCHT) with light (DIFN) ranking in third place. For lodgepole pine, DIFN was the best competition measure for predicting volume increment and the best competition index was again VCHT. Taller conifers had a stronger competitive effect than tall shrubs, with their effect on white spruce being larger than that on lodgepole pine.  相似文献   

14.
At Florence, Italy, several species of pine were experimentally infected with blister rust. Inoculations were carried out on 3 and 15 months-old seedlings. After antificial and natural inoculation, Brutia, Aleppo, Austrian, Swiss mountain, Maritime and Italian stone pine showed pyenia and aecia. Spotted seedlings of Ponderosa pine showed only mycelium of C. flaccidum in needle and stem tissues. The exotic species seemed to have a very high degree of resistance to blister rust.  相似文献   

15.
Predictions of damage risk from snow and wind at sites using tree characteristics of Scots pine (Pinus sylvestris L.), were made using a subset of data from permanent sample plots within the Swedish National Forest Inventory (NFI). The plots were sampled twice at five-year intervals between 1983 and 1992. A logistic risk assessment model was developed using data originating from 286 plots, dominated by Scots pine (> 65% of basal area), within one county situated in the boreal zone in northern Sweden (Västerbotten). The model was evaluated with NFI-data from two other counties, one adjacent to Västerbotten (Västermorrland, 99 plots), which is also in the boreal zone, and one (Kalmar, 138 plots) in the hemi-boreal zone in southern Sweden. In each plot, measurements at first inventory of tree characteristics for the largest undamaged sample tree, and measurements at second inventory of damage from snow and wind on all sample trees were used to develop a logistic model that predicts the damage probability for each site. The best predictors were upper diameter (ud, diameter at 3 or 5 m) and the ratio of height/diameter at breast height (rhd). According to the model calculations, the overall damage probability never exceeded 0.26 for any of the sample plots used for model development. At a given ud the probability of damage is higher for a site with trees of low rhd. The fit of the model was better for the adjacent Västernorrland county than for the southern county, Kalmar. This inferior predictability was explained by differences in tree characteristics between Kalmar and the other counties. The results show that it is possible to predict damage from snow and wind at a site by using only single tree characteristics.  相似文献   

16.
The aim of this study was to investigate the differences in infections caused by Agrobacterium tumefaciens in a conifer, Scots pine (Pinus sylvestris), and in a non-host deciduous species, silver birch (Betula pendula). All the Agrobacterium tumefaciens strains tested caused crown-gall formation in both tree species, but the infection rates varied remarkably. In Scots pine, the development of galls was rare, and slower than in silver birch. Inoculation into the base of the stem were the most successful in gall induction. Silver-birch galls were large, often surrounding the whole stem, in contrast to Scots pine galls, which were characterized by their small size and neck-like connection with the host plant. In silver birch, no other morphological changes could be seen. In Scots pine, abnormal phenotypes with proliferating short shoots above the galls were observed during the second and third growing season. The results indicate that, of the two non-host tree species, the deciduous one, silver birch, is more susceptible to an A. tumefaciens infection than the conifer, Scots pine. The matrix for A. tumefaciens infection in silver birch differs from that in Scots pine, since the terpene compounds of Scots pine seem either to kill the agrobacteria or to suppress their growth. The differences between the species could be partly caused by their difference in sensitivity to phytohormones. These features reflect evolutionary incompatibility between A. tumefaciens and a gymnosperm.  相似文献   

17.
An outbreak of the mountain pine beetle (Dendroctonus ponderosae Hopkins; MPB), currently affecting over 10.1 million hectares of lodgepole pine forests (Pinus contorta Dougl.) in British Columbia, Canada, is the largest in recorded history. We examined the dynamics of even-aged lodgepole pine forests in southern British Columbia, which were undergoing this MPB outbreak. Using dendroecology and forest measurements we reconstructed the stand processes of stand initiation, stand disturbances, tree mortality, and regeneration, and explained the current stand structure and the potential MPB impacts in selected stands. Our results indicate that stand-replacing fires initiated even-aged seral lodgepole pine stands in this region. In the absence of fire in the 20th century, multiple MPB disturbances, which each resulted in partial canopy removal, modified the simple one-layer structure of the fire-origin stands by the initiation of post-MPB disturbance regeneration layers, transforming the stands into complex, multi-aged stands. Despite high overstory mortality due to the current MPB outbreak, regeneration layers, which are likely to survive the current outbreak, will provide important ecological legacies and will contribute to mid-term timber supply.  相似文献   

18.
Tree species response to climate change-induced shifts in the hydrological cycle depends on many physiological traits, particularly variation in water relations characteristics. We evaluated differences in shoot water potential, vulnerability of branches to reductions in hydraulic conductivity, and water source use between Pinus contorta Dougl. ex Loud. var. latifolia Engelm. (lodgepole pine) and Pseudotsuga menziesii (Mirb.) Franco (interior Douglas-fir), and determined the consequences for seasonal changes in photosynthetic capacity. The Douglas-fir site had soil with greater depth, finer texture and higher organic matter content than soil at the lodgepole pine site, all factors that increased the storage of soil moisture. While the measured xylem vulnerability curves were quite similar for the two species, Douglas-fir had lower average midday shoot water potentials than did lodgepole pine. This implied that lodgepole pine exhibited stronger stomatal control of transpiration than Douglas-fir, which helped to reduce the magnitude of the water potential gradient required to access water from drying soil. Stable hydrogen isotope measurements indicated that Douglas-fir increased the use of groundwater during mid-summer when precipitation inputs were low, while lodgepole pine did not. There was a greater reduction of photosynthetic carbon gain in lodgepole pine compared with Douglas-fir when the two tree species were exposed to seasonal declines in soil water content. The contrasting patterns of seasonal variation in photosynthetic capacity observed for the two species were a combined result of differences in soil characteristics at the separate sites and the inherent physiological differences between the species.  相似文献   

19.
In the southern Rocky Mountains, current mountain pine beetle (Dendroctonus ponderosae Hopkins) outbreaks and associated harvesting have set millions of hectares of lodgepole pine (Pinus contorta var. latifolia Engelm. ex Wats.) forest onto new stand development trajectories. Information about immediate, post-disturbance tree regeneration will provide insight on dynamics of future stand composition and structure. We compared tree regeneration in eight paired harvested and untreated lodgepole pine stands in the Fraser Experimental Forest that experienced more than 70% overstory mortality due to beetles. New seedlings colonized both harvested and untreated stands in the first years after the beetle outbreak. In harvested areas the density of new seedlings, predominantly lodgepole pine and aspen, was four times higher than in untreated stands. Annual height growth of pine and fir advance regeneration (e.g., trees established prior to the onset of the outbreak) has doubled following overstory mortality in untreated stands. Growth simulations based on our regeneration data suggest that stand basal area and stem density will return to pre-beetle levels in untreated and harvested stands within 80-105 years. Furthermore, lodgepole pine will remain the dominant species in harvested stands over the next century, but subalpine fir will become the most abundant species in untreated areas. Owing to terrain, economic and administrative limitations, active management will treat a small fraction (<15%) of the forests killed by pine beetle. Our findings suggest that the long-term consequences of the outbreak will be most dramatic in untreated forests where the shift in tree species composition will influence timber and water production, wildfire behavior, wildlife habitat and other forest attributes.  相似文献   

20.
Glyphosate is the herbicide most extensively used for site preparation and conifer release. It is a broadspectrum herbicide and therefore crop safety is a critical issue. This study assessed the early effects of 14 different treatments, including no weed control, manual weed control, and 12 foliar-applied herbicide treatments at low,intermediate, high, and highest application rates and application timing on glyphosate phytotoxicity of containerized seedlings of Austrian pine(Pinus nigra J.F.Arnold.), Scots pine(P. sylvestris L.) and maritime pine(P.pinaster Aiton), conifer species widely used for afforestation and supplementary plantings in Turkish forestry. In general, Scots pine seedlings were tolerant to glyphosate compared to the other species. Glyphosate phytotoxicity varied significantly according to the time and rate of application. Seedlings were relatively tolerant to glyphosate in April whereas they were intolerant in May. The highest herbicide rate(1.2% v:v) was consistently phytotoxic to all species. Moreover, the effect of herbicide rate on seedling survival and growth varied significantly according to application date(i.e., application rate 9 date interaction). Seedlings appeared tolerant to glyphosate at low and intermediate rates(0.2, 0.4% v:v) between midspring and mid-summer, whereas they demonstrated significant sensitivity to the highest rate across all time periods. Glyphosate at the high rate(0.8% v:v) was particularly more phytotoxic when applied in May. Application of glyphosate at rates up to 0.8% could be recommended for weed control without significant pine damage in midspring when the needles presumably have a dense leaf epicuticular wax layer limiting herbicide penetration.Applications of 0.8 and 1.2% v:v are not recommended during May–June.  相似文献   

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