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1.
The effects of initial tree spacing on wood density at breast height were examined for 22-year-old Japanese larch (Larix kaempferi). The experiment involved the use of three plots with different initial tree spacing densities (300, 500, and 1000 trees/ha).
For five trees selected from each plot, the total tree height, diameter at breast height, height to the base of the live crown,
and crown diameter were measured. Ring width and wood density for individual growth rings were determined by X-ray densitometry.
A mixed-effects model was applied for fitting the radial variation in wood density in relation to initial spacing. Models
having various mean and covariance structures were tested in devising an appropriate wood density model. The model, consisting
of the mean structure with quadratic age effects and heterogeneous first-order autoregressive covariance, was able to describe
the radial variation in wood density. Closer spacing of trees (1000 trees/ha) resulted in a faster increase in wood density
from the pith outward than for more widely spaced trees, indicating that initial tree spacing may influence the age of transition
from juvenile to mature wood. 相似文献
2.
The influence of rate of growth on the density and heartwood extractives content of eucalypt species
J. Wilkes 《Wood Science and Technology》1984,18(2):113-120
Summary Basic density and extractives content, of the sapwood and outer heartwood respectively, were compared for a fast-and a slow-grown tree in five, 40 year old, coppice clumps of each of six eucalypt species. Rate of growth did not appreciably influence extracted wood density, although outer heartwood extractives levels were significantly (P<0.05) higher in the larger stems. The juvenile core, as defined by radial variation in basic density, occupied a similar proportion of the tree diameter in the fast-and slow-grown trees, suggesting that sampling was effected in tissues of similar maturity in both tree groups.Helpful discussions with Dr. W. A. Heather, a co-worker, and Dr. W. E. Hillis, Visiting Fellow of this Department, are gratefully acknowledged 相似文献
3.
The radial pattern of both maximum ring density and ring area of 36 black spruce trees were used to determine the transition age from juvenile wood to mature wood. The data were obtained by X-ray densitometry and both segmented linear and polynomial regressions were used to point out the age of the juvenile wood boundary. Three stand densities (1,790, 2,700 and 3,400 stems/ha) and three sampling heights (2.4, 5.1 and 7.8 m) were studied. Although maximum ring density and ring area presented similar radial patterns, they gave two significantly different results of transition ages. The maximum ring density over-estimated the transition age (17.6 years) in contrast to ring area (14 years). The results show that the transition from juvenile wood to mature wood occurred after 12 years at 7.8 m (versus 13.1 years at a height of 5.1 m, and 17.6 years at 2.4 m). Although transition age occurred later in the high stand density group (21 years), the difference was not significant between the three stand density groups. Nevertheless, transition age remains difficult to determine since no standard definition exists. The transition occurs over years, and most probably a transition wood exists between juvenile wood and mature wood. Estimation of the juvenile wood proportion in volume shows that it remains constant along the stem and it increases with stand density. 相似文献
4.
We evaluated heartwood, sapwood and bark content in teak trees. A total of 27 sample plots were laid out in teak plantations raised by State Forest Department in Karnataka covering different age groups... 相似文献
5.
《Scandinavian Journal of Forest Research》2012,27(4):330-350
To study and model the variation of wood properties, sample trees were selected from 42 Norway spruce and 20 Scots pine stands covering a wide variation in climatic and site conditions, stand maturation and tree sizes. Plot and tree measurements were followed by sampling wood from different heights in each sample tree and laboratory measurements of wood properties. Mixed linear and non-linear prediction models were developed using diameters, number of annual rings and climatic indices as explanatory variables. The variation in spruce properties explained by these variables was: basic density 50%, latewood content 52%, juvenile wood diameter 85%, heartwood diameter 94% and bark thickness 76%. The corresponding values for pine were 59, 54, 79, 92 and 85%. Random among-tree variance was an important contributor to the remaining variation for density and latewood. In general, only a minor part of the random variation was related to variance between stands. Predictions derived from the models for density and juvenile wood in both species, and heartwood in pine showed good agreement when validated with data sets from two other studies. The resulting models may be used for predicting wood properties in forest planning and in bucking computers in harvesters, provided that the essential information is available. 相似文献
6.
The wood quality from 50- to 70-year-old Tectona grandis trees from an unmanaged forest in East Timor was assessed. The aim was to evaluate teak in mature stands that had undergone
uncontrolled disturbances, e.g., fire and local community usage. Heartwood represented 91% of the tree radius at a height
of 1.7 m, and sapwood contained on average nine rings. The mean ring width showed within-tree and between-tree variability.
The chemical compositions of heartwood and sapwood were similar. Within-tree chemical variation occurred only in terms of
extractives, which increased from the pith (8.3%) to the heartwood-sapwood transition (12.7%) and decreased in the sapwood
(9.2%). Overall, the wood properties of teak from a unmanaged forest in East Timor were comparable to those reported for plantation
teaks of other origin: 607 kg/m3 basic density, 3.5% and 5.2% radial and tangential shrinkage, 141 N/mm2 modulus of rupture, 10684 N/mm2 modulus of elasticity, and 50 N/mm2 maximum crushing strength in compression parallel to the grain. Disturbances on individual tree growth arising from the unmanaged
status of the stand were evidenced by higher within-tree variability of ring width. However, the longitudinal and radial variations
of wood density and mechanical properties were of low magnitude and in a degree that did not negatively impact on timber quality. 相似文献
7.
《Forest Ecology and Management》1998,102(1):75-88
To evaluate the relationship of overstory residual trees to the growth of unmanaged young-to-mature understory Douglas fir (Pseudotsuga menziesii (Mirb.) Franco) and western hemlock (Tsuga heterophylla (Raf.) Sarg.), the basal area and volume of 14 paired plots with and without residual trees were examined in the Willamette National Forest, Oregon. Residual trees were large survivors of the fires that initiated the understory between 55 and 121 yr ago. Understory stands were naturally regenerated and not managed in any way. High residual tree and understory densities were negatively associated with understory volume. The relation of density of residual trees to total understory and Douglas-fir basal areas and volumes was best described by a negative logarithmic function. The rate of decrease in total understory and Douglas-fir basal areas and volumes per individual residual tree became smaller with increasing residual-tree density. Predicted total understory volume reduction was 23% with five residual trees/ha and 47% with 50 residual trees/ha, averaging 4.6% and 0.9% per residual tree, respectively. After including the estimated volume growth of residual trees since initiation of the understory, stand volume was still 19% lower with five residual trees/ha and 41% lower with 50 residual trees/ha than in stands with no residual trees, averaging a reduction of 38% and 0.8% per residual tree, respectively. In mixed stands of Douglas fir and western hemlock, predicted Douglas-fir basal area and volume declined more rapidly than did total understory basal area and volume when residual-tree densities exceeded about 15 trees/ha. This difference was probably due to the relative shade-intolerance of Douglas fir. Predicted Douglas-fir volume reduction was 13% with five residual trees/ha and 75% with 50 residual trees/ha, averaging 2.6% and 1.5% per residual tree, respectively. The southern aspects had more than 150% the total understory basal area and volume and more than 200% the Douglas-fir volume and basal area of the northern aspects. Lower density and basal area of understory trees, particularly of dominant and codominant Douglas fir, were associated with increasing residual-tree densities. Given the same diameter at breast height (DBH), heights of Douglas fir were not related to residual trees. Regardless of understory age, understory volume was greatest in stands with the lowest understory densities. These results suggest that timber production in unthinned green-tree retention units may be reduced and may depend on the density of leave-trees. Thinning of understory trees is recommended to reduce growth loss from intraspecific competition. 相似文献
8.
We studied heartwood and sapwood variation in western redcedar (Thuja plicata) at three sites, including a 95-year-old naturally regenerated, unmanaged stand, a 35-year-old planted spacing trial, and a 30-year-old naturally regenerated stand to which thinning and fertilization treatments had been applied. In the 95-year-old stand, we studied within-tree variation in heartwood and sapwood. In the thinning/fertilization trial and the planted spacing trial, we studied effects of cultural practices and growth rate on heartwood and sapwood. In the trees that we studied, sapwood width was generally fairly narrow, rarely exceeding 3.5 cm. Heartwood formation in western redcedar appeared to begin at a relatively small stem diameter (7 cm) and at a young age, probably 10–15 years. The amount and proportion of heartwood increased with distance downward from the top of the tree, with the implication that older trees will contain a greater proportion of heartwood than younger trees. For any given age, it appears that cultural treatments that favor rapid growth will result in stems with greater amounts of both sapwood and heartwood, and a greater proportion of heartwood. 相似文献
9.
Stoneman GL Crombie DS Whitford K Hingston FJ Giles R Portlock CC Galbraith JH Dimmock GM 《Tree physiology》1997,17(4):267-274
We studied the effects of five thinning treatments (T1 = 5.5, T2 = 11, T3 = 16.5, T4 = 22.5 and T5 = 28.5 m(2) ha(-1) basal area under bark) x two fertilizer treatments (F0 = unfertilized and F1 = fertilized with 400 kg ha(-1) N plus 229 kg ha(-1) P) on growth and water relations of pole-sized Eucalyptus marginata J. Donn ex Sm. trees growing in southwestern Australia. Thinning reduced leaf area index (LAI) from 2.1 in the T4 and T5 treatments to 0.8 in the T1F0 treatment. Fertilizer had no effect on LAI in the T2, T4 or T5 treatments, but increased LAI by 45 and 20% in the T1 and T3 treatments, respectively. Thinning plus fertilizing increased diameter growth most in the fastest growing trees, from 0.4 cm year(-1) for trees in the T5F0 and T5F1 treatments to 0.7 and 1.2 cm year(-1) for trees in the T1F0 and T1F1 treatments, respectively. In both fertilizer treatments, stand basal area and volume growth increased with increasing stand density up to 15 m(2) ha(-1), and thereafter declined with increasing stand density, such that the growth rate of trees in the T5 treatment was only half of that at a stand density of 15 m(2) ha(-1). In response to fertilizer, growth rates of the slowest and fastest-growing trees increased from 0.35 and 3.5 m(2) ha(-1) year(-1) (F0) to 0.56 and 5.4 m(3) ha(-1) year(-1) (F1), respectively. Stand growth efficiency (growth per unit LAI) increased in response to thinning, and fertilizer increased stand growth efficiency at all stand densities. Throughout the dry season, T5 trees had lower predawn shoot water potentials (Psi(pd)) (minimum of -1.5 MPa) than T1 or T2 trees (minimum of -0.7 MPa). Fertilizer decreased Psi(pd) in T5 trees (by -0.9 and -1.5 MPa, respectively, in F0 and F1), but not in T1 or T2 trees. Stand growth rate was closely related to cumulative midday water stress (CMWS) over the dry season, and volume growth rate declined sharply from 6 m(3) ha(-1) year(-1) at a CMWS of 130 MPa days, to zero at a CMWS of 220 MPa days. Application of fertilizer to thinned stands increased LAI, stand growth efficiency and stand growth. In unthinned stands, fertilizer increased stand growth efficiency and stand growth; however, it also increased tree water stress, which limited the fertilizer-induced increases in LAI and growth. We attribute the increase in tree and stand growth in response to application of fertilizer to increased photosynthetic rates, increased allocation to stem wood, and in thinned stands also to higher LAIs. 相似文献
10.
Yoshio Kijidani Naoki Ohshiro Daisuke Iwata Masanori Nagamine Taiga Nishiyama Junji Matsumura Shinya Koga 《Journal of Wood Science》2014,60(3):177-185
Recently, breeding programs have attempted to produce high growth rates for shorter rotation cycles in plantation trees. In these trees, the ratio of juvenile wood increases; thus, the juvenile wood properties should be improved for structural use. To this end, it is important to understand the influences on juvenile wood properties precisely. In this study, we report on the indole acetic acid (IAA) amounts of juvenile sugi (Cryptomeria japonica) trees in September and compare the IAA amounts to those in mature trees. The IAA amounts at the lower trunks in juvenile trees were significantly larger than those in mature trees and the IAA amounts decreased with tree height. In each stand, except a mature tree stand, there is no significant effect of IAA amounts on latewood width and MFA. However, put together all samples, the latewood width and MFA increased with IAA amounts in samples with IAA <200 ng/cm2. The samples at lower trunk in juvenile trees had significantly larger IAA amounts, larger MFA and larger latewood width than the samples in mature trees (p < 0.01). The very large IAA amounts may have a certain relation with juvenile wood properties. 相似文献
11.
本文就中条山油松株内幼龄材与成熟材材性差异的比较研究,讨论对幼龄期划分的依据。根据木材解剖特征、物理力学性质的径向变异规律,确定其幼龄期为14年。随着树干高度的增加,油松木材幼龄期逐渐缩短、株内幼龄材范围及所占断面上的比例变小。株内幼龄材与成熟材材性差异显著。幼龄材管胞长度短、直径小,胞壁薄,微纤丝角度大,生长轮较宽,晚材率低,浸提物含量高,基本密度较大。幼龄材的力学强度远远小于成熟材。 相似文献
12.
Alexander Berrocal Johana Gaitan-Alvarez Roger Moya David Fernandez-Solis Edgar Ortiz-Malavassi 《林业研究》2020,31(2):667-676
To elucidate the development of heartwood,bark,sapwood,pith and specific gravity of wood in fastgrowing teak(Tectona grandis)plantations in Costa Rica,we sampled three trees in each of 55 plantations and modelled each variable with age,site and different tree heights.Age and stand density of plantations were significant correlated with stem diameter at breast height and total height of the tree.Formation of heartwood was initiated at the age of 4-year-old and increased in direct proportion with age.The age of plantation had a significant relationship with stem diameter at breast height,heartwood percentage,sapwood thickness,sapwood percentage,percentage of bark,pith diameter and percentage,and specify gravity.The model for these tree parameters was model with these parameters as dependent variable and in relation to age as independent variable. 相似文献
13.
This study considered the effects of thinning on the development of compression wood in stems of 35-year-old stand of Corsican
pine (Pinus nigra L.). Part of the stand had been thinned at 5-yearly intervals and part left unthinned. Twenty trees each from the thinned
and unthinned stands were randomly selected and felled. Measurements were made on tree height, stem diameter, stem slenderness
and canopy depth. Wood samples were removed from the central part of the main log and cross-sectional measurements made on
ring width, basic density and compression wood content. Cross-sectional area of compression wood was found to be three time
higher in stems from the unthinned trees in comparison with those from the thinned trees. No significant differences in mean
radial ring width or basic density were found between treatments. Correlations indicated that, with increasing in stem diameter,
compression wood content increased in the unthinned trees, while a decline in compression was observed in the thinned trees.
Tree height was also positively correlated with compression wood content in unthinned trees, while no equivalent relationship
was observed in thinned trees. Observations from this study, while not conclusive, suggest that phototropic stimulus may be
producing stem inclinations in the unthinned stand as trees compete for space in the canopy, whereas crown competition has
been largely eliminated in the thinned stand; and that this is responsible for compression wood levels recorded in this study. 相似文献
14.
Nishtman Hatami Peter Lohmander Mohammad Hadi Moayeri Soleiman Mohammadi Limaei 《林业研究》2020,31(1):99-106
The aim of this study was to develop and test a new basal area growth model in mixed species continuous cover forests in northern Iran.Weanalyzed 421 core samples from 6 main species in the forest area to develop our growth model.In each plot,we measured variables such as total tree height(m),diameter at breast height(DBH)(cm)and basal area of larger trees as cumulative basal areas of trees(GCUM)ofDBH[5 cm.The empirical data were analyzed using regression analysis.There was a statistically significant nonlinear function between the annual basal area increment,as the dependent variable,and the basal area of the individual trees and competition as explanatory variables.Reference area from the largest trees,was circular plot with area of 0.1 ha.GCUM was estimated for trees of DBH>5 cm.Furthermore,we investigated the dependencies of diameter growth of different species on stand density at different levels of competition,and diameter development of individual trees through time.The results indicate that competition caused by larger neighborhood trees has a negative effect on growth.In addition,the maximum diameter increment is affected by competition level.Therefore,the maximum diameter increment of species occurs when the trees are about 35–40 cm in dense-forest(40 to 0 m^2 per ha)and when the trees are about 60 to 70 cm in very dense forest(60 to 0 m^2 per ha)which is more likely to Caspian natural forests with high level density due to uneven-aged composition of stands. 相似文献
15.
Significant efforts have been made to improve teak; however, evaluation in juvenile step is limited. The objective of this study was to conduct an early assessment of the wood properties of 4-year-old Tectona grandis. Samples of 36 clones were collected to determine their morphological tree parameters [diameter a breast height (DBH), diameter of the second log, tree height, and log quality]. Presence of growth stress, heartwood percentage, shrinkage (radial, tangential, and volumetric), basic density, and ultrasound velocity (USV) were determined for standing trees, logs, green lumber, and dry lumber. The results indicate that DBH and USV in standing trees can be used to predict elastic module (ED), mainly ED of the standing tree and dry lumber, as well as the possible presence of growth stress. Additionally, growth stress can be predicted by UVS in standing tree. Tangential and volumetric shrinkage were not predicted by tree morphology, but radial shrinkage was predicted by diameter and UVS was not affected by any shrinkage. Basic density was predicted by DBH and UVS measured in log. 相似文献
16.
Chun-Sheng Wang Ji Zeng Sebastian Hein Zhi-Gang Zhao Jun-Jie Guo 《Scandinavian Journal of Forest Research》2017,32(8):679-687
Both crown and branch characteristics greatly affect tree growth and timber quality, and their development is closely related to planting density (PD). Here crown and branch attributes of Betula alnoides were investigated in a 14-year-old experimental plantation in southern China with five planting densities ranging from 500 to 3333 stems per hectare (sph). The results showed that high PD significantly reduced crown and branch sizes. The ratios of crown diameter to stem diameter at breast height at the stand level and for dominant and co-dominant trees were almost constant regardless of PD. PD also had no significant influence on the number and angle of branches, and only planting with the highest density significantly increased branch density and mortality. Insertion angle (IA) of dead branches was greater than that of live branches and these differences were significant at the two low planting densities (833 and 500?sph). Linear mixed models revealed that branch diameter (BD) was negatively correlated with its relative height in the crown and IA. The binary-logistic regression model indicated that branch status (live or dead) was codetermined by BD and PD. These findings can help farmers cultivate high-quality wood of B. alnoides and obtain a higher economic return. 相似文献
17.
Adzo Dzifa Kokutse Alexia Stokes Nomessi Kuma Kokutse Kouami Kokou 《Annals of Forest Science》2010,67(4):407-407
18.
Context
Loblolly pine is often grown in intensively managed plantations for wood production. In order to fully evaluate the effects of management practices on wood quality and ultimately value, it is necessary to relate mechanical properties to management practices.Aims
The aim of this study was to evaluate the effect of planting density on mechanical properties of lumber recovered from loblolly pine trees from a 27-year-old spacing trial and develop prediction equations for modulus of elasticity and modulus of rupture from stand, tree, and board characteristics.Methods
Regression methods were applied to sample trees from three planting densities (2,989, 1,682, and 746 trees ha?1) and used to relate mechanical properties of lumber extracted from the trees to stand, tree, and board characteristics.Results
Initial planting density was found to be correlated with modulus of elasticity and, to a lesser extent, with modulus of rupture. Including board characteristics and utilizing the visual grade and board position as regressors produced improved prediction equations.Conclusions
The mean modulus of elasticity declines with decreasing planting density while the variability increases, suggesting that planting density is a surrogate for frequency and size of knots. Thus, lower planting densities, while producing more lumber, may produce proportionally fewer boards of greater modulus of elasticity than higher planting densities. 相似文献19.
Secondary xylem of woody plants has a large volumetric proportion of gas occupying spaces that would otherwise be filled with water. We examined whether these gas-filled voids have a mechanical role by either decreasing the fresh mass the tree must support (by replacing some of the water with gas) or by providing inexpensive filler to increase stem diameter (thereby increasing the second moment of area at the expense of the modulus of elasticity and modulus of rupture). Calculations from published data show that temperate softwood species (n = 26) average 18 and 50% gas by volume for sapwood and heartwood, respectively; temperate hardwood species (n = 31) average 26% gas by volume in both the sapwood and heartwood; and tropical species (n = 52) with mixed sapwood and heartwood have 18% gas by volume. In this paper, we develop equations to show how gas affects the mechanical behavior of tree stems, and describe model results to show how gas affects mechanical stability, based on mass and stem diameters for six 34-year-old Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) trees. For the same applied load, modeled stems in which the gas space was filled with water differed in their surface stresses by < 2% from modeled stems in the native state (partially gas-filled), indicating no practical benefit from a reduction in stem mass due to gas. A second modeling scenario compared the native state to stems in which gas was removed and stem diameters decreased (and material properties adjusted to concur with the increased wood density) to conserve mass. Removal of the gas-filled voids resulted in up to 41% higher surface stress for the same applied load, caused by a decrease in the second moment of area greater than the increase in modulus of elasticity. Trees with gas removed had higher modulus of rupture, but could withstand up to 14% lower maximum wind forces than trees in their native state, suggesting a biomechanical role for the gas if the model assumptions are valid. The gas content may, however, have evolved in response to pressures unrelated to biomechanics. We discuss some of its potential effects on sapwood physiology. 相似文献
20.
《Scandinavian Journal of Forest Research》2012,27(3):247-257
Abstract This study assessed variation in modulus of elasticity of trees and logs of Scots pine (Pinus sylvestris) trees. The study used 192 sample trees (c. 90–150 years) selected from 24 clear-felling forests in central and southern Sweden. Modulus of elasticity (MOE) assessed with transit-time technology on standing tree stems at 0.5–2.0m on the southern and northern side of each tree varied from 8.6 to 17.6 GPa. No systematic MOE difference was found between the southern and northern side of tree stems. The sometimes large MOE variations seen in some individual trees are probably a result of wood variation and wood defects. MOE assessed with resonance-based technology varied between 7.4 and 14.1 GPa for logs cut at similar height (<6.0 m). Models of MOE variation were derived from factors related to growth conditions at stand and tree level, with an R 2 adj of c. 0.46–0.62. The models indicate that growth and tree attributes associated with and/or creating less stem taper would yield trees with higher MOE. 相似文献