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1.

? Context

Coarse woody debris (CWD, ≥10 cm in diameter) is an important structural and functional component of forests. There are few studies that have estimated the mass and carbon (C) and nitrogen (N) stocks of CWD in subtropical forests. Evergreen broad-leaved forests are distributed widely in subtropical zones in China.

? Aims

This study aimed to evaluate the pools of mass, C and N in CWD in five natural forests of Altingia gracilipes Hemsl., Tsoongiodendron odorum Chun, Castanopsis carlesii (Hemsl.) Hayata, Cinnamomum chekiangense Nakai and Castanopsis fabri Hance in southern China.

? Methods

The mass of CWD was determined using the fixed-area plot method. All types of CWD (logs, snags, stumps and large branches) within the plot were measured. The species, length, diameter and decay class of each piece of CWD were recorded. The C and N pools of CWD were calculated by multiplying the concentrations of C and N by the estimated mass in each forest and decay category.

? Results

Total mass of CWD varied from 16.75 Mg ha?1 in the C. fabri forest to 40.60 Mg ha?1 in the A. gracilipes forest; of this CWD, the log contribution ranged from 54.75 to 94.86 %. The largest CWD (≥60 cm diameter) was found only in the A. gracilipes forest. CWD in the 40–60 cm size class represented above 65 % of total mass, while most of CWD accumulations in the C. carlesii, C. chekiangense and C. fabri forests were composed of pieces with diameter less than 40 cm. The A. gracilipes, T. odorum, C. carlesii and C. chekiangense forests contained the full decay classes (from 1 to 5 classes) of CWD. In the C. fabri forest, the CWD in decay classes 2–3 accounted for about 90 % of the total CWD mass. Increasing N concentrations and decreasing densities, C concentrations, and C:N ratios were found with stage of decay. Linear regression showed a strong correlation between the density and C:N ratio (R 2?=?0.821). CWD C-stock ranged from 7.62 to 17.74 Mg ha?1, while the N stock varied from 85.05 to 204.49 kg ha?1. The highest overall pools of C and N in CWD were noted in the A. gracilipes forest.

? Conclusion

Differences among five forests can be attributed mainly to characteristics of the tree species. It is very important to preserve the current natural evergreen broad-leaved forest and maintain the structural and functional integrity of CWD.  相似文献   

2.
Narrowing the uncertainties in carbon (C) and nitrogen (N) dynamics during decomposition of coarse woody debris (CWD) can significantly improve our understanding of forest ecosystem functioning. We examined C, N and pH dynamics in the least studied CWD component—tree bark in a 66-year-long decomposition chronosequence. The relative C concentration decreased by ca. 32% in pine bark, increased by ca. 18% in birch bark and remained stable in spruce and aspen bark. Nitrogen increased in bark of all tree species. In conifer bark, it increased along with epixylic succession. Over 45 years, the relative C/N ratio in bark decreased by 63 and 45% for coniferous and deciduous species, respectively. Bark pH did not change. Due to bark fragmentation, the total C and N amounts in bark of individual logs of aspen, birch, pine and spruce decreased at average rates of 0.03, 0.02, 0.26 and 0.05 year?1, and 0.02, 0.02, 0.03 and 0.03 year?1, respectively. At the forest stand level, the total amounts of C and N in log bark were 853 and 21 kg ha?1 or 11.2 and 45.5% of the C and N amounts stored in downed logs and ca. 2.3–3.8 and 2.2–2.4%, respectively, of total C and N amounts stored in forest litter. In boreal forests, decomposing log bark may act as a long-term source of N for wood-inhabiting communities.  相似文献   

3.

Fineroots (≤ 2 mm diameter) are dynamic components of the forest ecosystems and play important role in water and nutrient acquisition in forests. These roots are sensitive to forest fertilization and therefore, the response of fineroots to nutrient application would provide better understanding of the forest carbon and nutrient dynamics that will be helpful in sustainable forest management plans. Two fertilization treatments, including (1) F400: 400 g P2O5 (16.5%)/tree and (2) F600: 600 g P2O5 /tree, and F0: control (without fertilization), were applied in an Acacia mangium plantation with a planting density of 1100 trees/ha (3 m?×?3 m). The evaluation of fineroot growth across seasons showed that fertilization significantly increased production and subsequent mortality and decomposition. The total decomposition associated with F600 application was 7.95 tons ha?1 year?1, equaling 121% of F400 and 160% of the control. Mortality in F600 was 8.75 tons ha?1 year?1, equaling 111% of F400 and 198% of F0, while production in F600 was 10.40 tons ha?1 year?1, equaling 127% of F400 and 143% of F0. Fineroot production, mortality, and decomposition are seasonally dependent, with higher values measured in the rainy season than in the dry season. Stand basal area increment was significantly correlated with fineroot production (R2?=?0.75), mortality (R2?=?0.44), and decomposition (R2?=?0.48). This study showed that fertilization could facilitate fineroot production, which can then lead to a higher turnover of carbon and nutrients through the decomposition of the greater mass of the fineroots.

  相似文献   

4.
Coarse woody debris (CWD) is involved in important forest ecosystem functions and processes, e.g., habitat provision, water retention, and organic matter decomposition. However, a quantitative, CWD-produced soil organic carbon (SOC) imprint has not yet been detected, possibly due to lack of free adsorption sites on soil minerals. To circumvent this potential constraint, we selected plots with and without CWD in a beech (Fagus sylvatica L.) primeval forest in the West Carpathian volcanic range (Slovakia). Local andic soil contains abundant allophane and amorphous Fe-compounds as important SOC binding agents. The C concentration in the fine earth of sampled soils was determined by the dry combustion method. We established that organic carbon concentration decreased with depth from 0.20 kg kg?1 (0.0–0.3 m) to 0.11 kg kg?1 (0.3–0.5 m) in soil with CWD and from 0.13 kg kg?1 (0.0–0.3 m) to 0.07 kg kg?1 (0.3–0.5 m) in soil without CWD. The respective average differences in soil organic carbon concentration (0.07 kg kg?1) and stock (15.84 kg m?2) between the two series of plots within the upper 0.3 m were significant according to the t test (P < 0.05 or P < 0.01, respectively). Also, corresponding differences within the 0.3–0.5 m layer (0.04 kg kg?1 and 5.51 kg m?2) were significant (P < 0.05, P < 0.001). Our results represent the first indication that CWD-produced SOC imprint may reach deeper than just a few centimeters in soils featuring high adsorption capacity, such as Andosols.  相似文献   

5.
The decomposition of harvest residues (brash) in managed forests has an important influence on the carbon (C) and nitrogen (N) stocks of these ecosystems. The brash input from thinning events in a 25-year-old Sitka spruce plantation was determined. A litter-bag method was used to determine the mass loss and decomposition rate of brash left on the forest floor. The changes in C and N concentrations and the C:N ratio of the needles and branches were also monitored as decomposition progressed for 2.5 years. Using the decomposition rate (k b) and estimated brash inputs, we then determined the total cumulative stock of C that the brash could supply to the deadwood pool over a 41-year rotation period. The three thinning events resulted in the addition of 37.99 t C ha?1 and 0.61 t N ha?1 to the forest floor. A significant mass loss of 44 % was recorded from brash decomposition bags after 2.5 years, with a rapid loss of 35 % in the first year, after which the rate of decomposition slowed. The k b-value and residence time (95 % decomposition) were 0.311 year?1 and 9.6 years, respectively. There was a 69 % increase in the N concentration of needles after 1.5 years, while an increase of 185 % in the N concentration of branches was recorded after 2.5 years. The C concentration (48.55 ± 0.20 %) did not differ significantly between the needles and branches over time. The accumulated C stock from decomposing brash at clearfell was estimated at 18.51 t C ha?1.  相似文献   

6.
Intensification of coffee (Coffea arabica) production is associated with increases in inorganic fertilizer application and decreases in species diversity. Both the use of organic fertilizers and the incorporation of trees on farms can, in theory, reduce nutrient loss in comparison with intensified practices. To test this, we measured nutrient concentrations in leachate at 15 and 100 cm depths on working farms. We examined (1) organically managed coffee agroforests (38 kg N ha?1 year?1; n = 4), (2) conventionally managed coffee agroforests (96 kg N ha?1 year?1; n = 4), and (3) one conventionally managed monoculture coffee farm in Costa Rica (300 kg N ha?1 year?1). Concentrations of nitrate (NO3 ?-N) and phosphate (PO4 3?-P) were higher in the monoculture compared to agroforests at both depths. Nitrate concentrations were higher in conventional than organic agroforests at 15 cm only. Soil solutions collected under nitrogen (N)-fixing Erythrina poeppigiana had elevated NO3 ?-N concentrations at 15 cm compared to Musa acuminata (banana) or Coffea. Total soil N and carbon (C) were also higher under Erythrina. This research shows that both fertilizer type and species affect concentrations of N and P in leachate in coffee agroecosystems.  相似文献   

7.
The effect of forest conservation on the organic carbon (C) stock of temperate forest soils is hardly investigated. Coarse woody debris (CWD) represents an important C reservoir in unmanaged forests and potential source of C input to soils. Here, we compared aboveground CWD and soil C stocks at the stand level of three unmanaged and three adjacent managed forests in different geological and climatic regions of Bavaria, Germany. CWD accumulated over 40–100 years and yielded C stocks of 11 Mg C ha?1 in the unmanaged spruce forest and 23 and 30 Mg C ha?1 in the two unmanaged beech–oak forests. C stocks of the organic layer were smaller in the beech–oak forests (8 and 19 Mg C ha?1) and greater in the spruce forest (36 Mg C ha?1) than the C stock of CWD. Elevated aboveground CWD stocks did not coincide with greater C stocks in the organic layers and the mineral soils of the unmanaged forests. However, radiocarbon signatures of the O e and O a horizons differed among unmanaged and managed beech–oak forests. We attributed these differences to partly faster turnover of organic C, stimulated by greater CWD input in the unmanaged forest. Alternatively, the slower turnover of organic C in the managed forests resulted from lower litter quality following thinning or different tree species composition. Radiocarbon signatures of water-extractable dissolved organic carbon (DOC) from the top mineral soils point to CWD as potent DOC source. Our results suggest that 40–100 years of forest protection is too short to generate significant changes in C stocks and radiocarbon signatures of forest soils at the stand level.  相似文献   

8.
We measured the rates of respiration from snags and logs (“coarse woody debris”, CWD) of Japanese red pine (Pinus densiflora Sieb. et Zucc.) to examine the rate of decomposition and CO2 efflux from these materials in a temperate secondary broad-leaved forest in Japan. At this site, a high quantity of CWD of P. densiflora had accumulated as a result of pine wilt disease during the 1970s. Respiration rates were measured using a dynamic closed chamber method combined with an infrared gas analyzer. We measured the respiration rate of 7 samples of snags and 10 samples of logs from August 2003 to January 2004. The responses of the respiration rates of snags (Rsnag) and logs (Rlog) to changing temperature were both exponential and the responses to water content were quadratic, and the same function could be used to estimate annual values of both Rsnag and Rlog. Intensive measurements of water contents of snags and logs showed a marked difference in water content. The mean water content of snags was 20% of log water content. This difference was likely responsible for the observed difference in annual Rsnag and Rlog. The decay rate constants estimated from the respiration rates measurement of snags and logs were 0.019 and 0.081 year−1, respectively. Despite being lower than Rlog, Rsnag was a significant compartment of the CWD carbon budget at this site.  相似文献   

9.
Some land-use systems in Saskatchewan, Canada include the nitrogen-fixing trees buffaloberry (Shepherdia argentea Nutt.), caragana (Caragana arborescens Lam.) and sea buckthorn (Hippophae rhamnoides L.). These species provide various ecological functions such as ameliorating soil moisture, light and temperature but little work has been done quantifying biological nitrogen fixation by these species. Greenhouse experiments were conducted to quantify N2-fixation using the 15N natural abundance and the 15N dilution methods. Buffaloberry failed to form nodules in all but one of the four replicates in the natural abundance experiment. Using the 15N dilution method, the percentage of N derived from atmosphere (%Ndfa) in the shoot of buffaloberry averaged 64 %. For caragana, the mean  %Ndfa was 59 and 65 % and seabuckthorn was 70 and 73 % measured using the natural abundance and dilution methods, respectively. Because of large variability in biomass production between plants grown in the natural abundance experiment and the dilution experiment, the amounts of N2 fixed also were very variable. Buffaloberry fixed an average of 0.89 g N m?2; the average for caragana ranged from 1.14 to 4.12 g N m?2 and seabuckthorn ranged from 0.85 to 3.77 g N m?2 in the natural abundance and dilution experiments, respectively. This corresponds to 16 kg N ha?1 year?1 for buffaloberry; an average of 15–73 kg N ha?1 year?1 in caragana and 11–67 kg N ha?1 year?1 in seabuckthorn. The substantial amounts of N2 fixed by these species indicate that they have the potential to contribute to the overall N balance in land-use systems in which they are included.  相似文献   

10.
Root biomass plays an essential role in carbon sequestration at both individual and ecosystem scales, yet few feasible methods for measuring root biomass of individual trees have been developed. We sampled 11 trees in a pure 20-year-old Pinus tabuliformis plantation to estimate total root, fine root and leaf biomass of individual trees using the nested regression method. The relationships between fine root biomass and surface area as well as leaf biomass and area were examined across tree sizes. Fine roots of P. tabuliformis were estimated to consume 40 % of the net primary production for turnover, and absorbed a total of 1364 g m?2 year?1 of major nutrients. A significant linear relationship was determined between roots and leaves in terms of both biomass and surface area. The findings added weight to the concept that the functional balance theory is applicable to even-aged mature trees of P. tabuliformis.  相似文献   

11.
The effects of different simulated rainfall regimes on water relations, growth, and biomass production and allocation of five provenances of Cedrela fissilis, C. saltensis and C. balansae were assessed in a pot-grown seedling experiment in greenhouse under uncontrolled conditions. Four simulated rainfall regimes were applied: 600 mm year?1 (severe water deficit), 800 mm year?1 (mild water deficit), 1,000 mm year?1 (mild wet), and 1,200 mm year?1 (well-watered) used as control. Provenances were compared in terms of water relations (midday leaf water potential, Ψ md and relative water content, RWC), growth (root, shoot and leaf length, neck diameter), growth rate (shoot- and neck growth rate, SGR and NGR), biomass production (root, shoot and leaf dry weight), and biomass allocation (root:shoot ratio, RSR). Multivariate analyses revealed that water relations differed significantly under all simulated rainfall treatments by means of several variables throughout time. Effects of species × treatment, and provenance nested within species × treatment interactions significantly were important. SGR and RSR also were important to explain differences in Ψ md and RWC between treatments and provenances. Broadly C. balansae and C. saltensis provenances seem to display a better performance than C. fissilis provenances under severe water deficit only. However, despite differences between provenances, it was not possible to clearly separate them according to their origin, based on water relations and growth performance. Even though the results of this study performed on pot-grown seedlings are based on relatively few observations and require additional confirmation, they allow an opportunity for speculation on the factors that are related to intraspecific variations that are likely to occur during the growth of seedlings from different geographical sites in field condition.  相似文献   

12.
The growth patterns of annually resolved tree rings are good indicators of local environmental changes, making dendrochronology a valuable tool in air pollution research. In the present study, tree-ring analysis was used to assess the effects of 16 years (1991–2007) of chronic nitrogen (N) deposition, and 10 years (1991–2001) of reduced nitrogen input, on the radial growth of Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) growing in the experimental area of Lake Gårdsjön, southwest Sweden. In addition to the ambient input of c. 15 kg N ha?1 year?1, dissolved NH4NO3 was experimentally added to a 0.52-ha watershed at a rate of c. 40 kg ha?1 year?1. Atmospheric N depositions were reduced by means of a below-canopy plastic roof, which covered a 0.63-ha catchment adjacent to the fertilized site. The paired design of the experiment allowed tree growth in the N-treated sites to be compared with the growth at a reference plot receiving ambient N deposition. Nitrogen fertilization had a negative impact on pine growth, while no changes were observed in spruce. Similarly, the reduction in N and other acidifying compounds resulted in a tendency towards improved radial growth of pine, but it did not significantly affect the spruce growth. These results suggest that spruce is less susceptible to changes in the acidification and N status of the forest ecosystem than pine, at least in the Gårdsjön area.  相似文献   

13.
Afforestation and ecological restoration have often been carried out with fast-growing exotic tree species because of their high apparent growth and yield. Moreover, fast-growing forest plantations have become an important component of mitigation measures to offset greenhouse gas emissions. However, information on the long-term performance of exotic and fast-growing species is often lacking especially with respect to their vulnerability to disturbance compared to native species. We compared carbon (C) storage and C accumulation rates in vegetation (above- and belowground) and soil in 21-year-old exotic slash pine (Pinus elliottii Engelm.) and native Masson pine (Pinus massoniana Lamb.) plantations, as well as their responses to a severe ice storm in 2008. Our results showed that mean C storage was 116.77 ± 7.49 t C ha?1 in slash pine plantation and 117.89 ± 8.27 t C ha?1 in Masson pine plantation. The aboveground C increased at a rate of 2.18 t C ha?1 year?1 in Masson pine and 2.23 t C ha?1 year?1 in slash pine plantation, and there was no significant difference in C storage accumulation between the two plantation types. However, we observed significant differences in ice storm damage with nearly 7.5 % of aboveground biomass loss in slash pine plantation compared with only 0.3 % loss in Masson pine plantation. Our findings indicated that the native pine species was more resistant to ice storm because of their adaptive biological traits (tree shape, crown structure, and leaf surface area). Overall, the native pine species might be a safer choice for both afforestation and ecological restoration in our study region.  相似文献   

14.
The effect of different planting densities (100,000 and 167,000 plants ha?1) and levels of nitrogen fertilization (0, 261, 521, and 782 kg N ha?1 year?1) on biomass production and chemical composition of Moringa oleifera was studied in a split-plot design with four randomized complete blocks over 2 years with eight cuts year?1 at the National Agrarian University farm in Managua, Nicaragua (12°09′30.65″N, 86°10′06.32″W, altitude 50 m above sea level). Density 167,000 plants ha?1 produced significantly higher total dry matter yield (TDMY) and fine fraction yield (FFDM), 21.2 and 19.2 ton ha?1 respectively, compared with 11.6 and 11 ton ha?1 for 100,000 plants ha?1. Growth rate in 167,000 plants ha?1 was higher than in 100,000 plants ha?1 (0.06 compared with 0.03 ton ha?1 day?1). Average plant height was 119 cm irrespective of planting density. Fertilization at the 521 and 782 kg N ha?1 year?1 levels produced the highest TDMY and FFDM in both years of the study and along all cuts. The interaction between cut and year was significant, with the highest TDMY and FFDM during the rainy season in the second year. Chemical composition of fractions showed no significant differences between planting densities. Significantly higher crude protein content was found in the coarse fraction at fertilizer levels 521 and 782 kg N ha?1 year?1 (87.9 and 93.7 g kg?1 DM) compared with lower levels. The results indicate that Moringa can maintain up to 27 ton ha?1 dry matter yield under dry tropical forest conditions over time at a planting density of 167,000 plants ha?1 if the soil is regularly supplied with N at a level of approximately 521 kg ha year?1 in conditions where phosphorus and potassium are not limiting.  相似文献   

15.
Carbon (C) sequestration potential was quantified for five tree species, commonly used in tree-based intercropping (TBI) and for conventional agricultural systems in southern Ontario, Canada. In the 25-year-old TBI system, hybrid poplar (Populus deltoides × Populus nigra clone DN-177), Norway spruce (Picae abies), red oak (Quercus rubra), black walnut (Juglans nigra), and white cedar (Thuja occidentalis) were intercropped with soybean (Glycine max). In the conventional agricultural system, soybean was grown as a sole crop. Above- and belowground tree C Content, soil organic C, soil respiration, litterfall and litter decomposition were quantified for each tree species in each system. Total C pools for hybrid poplar, white cedar, red oak, black walnut, Norway spruce and a soybean sole-cropping system were 113.4, 99.4, 99.2, 91.5, 91.3, and 71.1 t C ha?1, respectively at a tree density of 111 trees ha?1, including mean tree C content and soil organic C stocks. Net C flux for hybrid poplar, white cedar, red oak, black walnut, Norway spruce and soybean sole-crop were 2.1, 1.4, 0.8, 1.8, 1.6 and ?1.2 t C ha?1 year?1, respectively. Results presented suggest greater atmospheric CO2 sequestration potential for all five tree species when compared to a conventional agricultural system.  相似文献   

16.
Although agrosystems are recognized for their socio-economic value, few works have been conducted to assign its sequestration potential and ecological services. Accordingly, this study aimed to evaluate the ecological services of the eucalyptus stands in order to permit to small producers the access in carbon credit market. Three stands were selected according to age. Data were compared to that of a savannah (control). In total, 12,817 individuals belonging to 30 families, 53 genera and 70 species were identified in the plantations against 7107 individuals belonging to 24 families, 36 genera and 42 species in the savannah. Gmelina, Annona, Hymenocardia, Allophyllus, Daniellia, Terminalia and Piliostigma were the most represented genera. There was no significant difference between Savannah and plantations in terms of diversity (p > 0.05). The largest stock of carbon was found in oldest stands (108.51 ± 26.46 t C/ha) against 13.62 ± 3.03 t C/ha in Savannah. Eucalyptus saligna stored 39.66 t C/ha (4 t C ha?1year?1) in young stands; 57.28 t C/ha (6 t C ha?1year?1) in medium stands and 85.46 t C/ha (9 t C ha?1year?1) in old stands. The sequestration potential was higher in eucalyptus stands (398.25 t CO2eq/ha) than savannah (50.05 t CO2eq/ha). In total 956.82 t CO2eq/ha were sequestered for an economic value of $9568.45/ha against 50.05 t CO2eq/ha corresponding to $500.56/ha in Savannah. Eucalyptus stands are carbon sinks and could be an opportunity for financial benefits in the event of payment for environmental services in the context of the CDM process.  相似文献   

17.
Coarse woody debris (CWD) has become recognised as an important component of the carbon (C) pool in forest ecosystems. In Ireland, managed Sitka spruce (Picea sitchensis (Bong) Carr.) forests account for 52.3% of the total forest estate. To determine the stock and decay dynamics of above and belowground CWD, field surveys using fixed area sample plots, were conducted in six even-aged Sitka spruce stands, representing the young, intermediate and mature stages of a typical commercial rotation. The volume, mass, density loss and C:N ratio of all CWD types (logs, stumps, and coarse roots) were determined using a five-decay class (DC) system. The decay rates and half life of CWD was also determined. To estimate CWD coarse root mass; roots associated with stumps classified in different decay classes were excavated. The coarse roots were categorised into small (2-10 mm), medium (10-50 mm) and large (>50 mm) diameter classes.CWD C-mass ranged from 6.98 to 18.62 Mg ha−1 and was highest in an intermediate forest (D35), while the aboveground volume varied from 6.31 to 42.27 m3 ha−1. Coarse roots accounted for 21% to 85% of the total CWD C-pool in the surveyed stands. The total CWD C-mass was poorly correlated with the number of thinning events (R2 = 0.29), when data from D35 was excluded. The density loss was significant in logs (45%), stumps (58%), and small- (38%), medium- (50%) and large roots (38%) as decay progress from DC 0 to 4. There was a 46%, 41%, 51%, 72% and 57% decline in C:N ratio of logs, stumps, small-, medium- and large roots, respectively, as decay progressed from DC 0 to 4. The density decay rates were 0.059, 0.048 and 0.036 kg m−3 year−1 for logs, stumps and coarse roots, respectively. The size classification of roots did not significantly affect their decay rate. The half life (50% decomposition) of CWD was estimated has 12-, 14- and 19 years for logs, stumps and roots of Sitka spruce. Regression curves showed a strong correlation between the density and C:N ratio (R2 = 0.69, 0.74 and 0.93 for logs, stumps and coarse roots, respectively). The long term storage of C and its slow rate of decomposition make CWD a vital structural and functional component of the CWD C-pool and a major controller of forest ecosystem C-retention.  相似文献   

18.
In tropical areas of Mexico, Leucaena leucocephala is widely used in silvopastoral systems. However, little information exists on other native woody species of high forage potential, such as Guazuma ulmifolia. The aim of this study was to evaluate the components of biomass, forage yield and quality, and availability of N in fodder banks of L. leucocephala, G. ulmifolia, and a mixture of both species during dry and rainy seasons, under sub-humid tropical conditions. The experimental unit was a 5 × 10 m plot, containing three rows with 2 m between rows; each row had 20 plant positions with 0.50 m between plants. Within each plant position there was either a single plant, in the case of pure-crop, or two plants, in the case of mixed of both species. A complete randomized block design with three repetitions was used. In both seasons, there were a significantly greater proportion of leaves in the G. ulmifolia fodder banks (71 %) and in mixed fodder banks (69 %) than in L. leucocephala fodder banks (64 %). Consequently, these systems had leaf-to-stem ratios of 2.4, 2.2 and 1.9, respectively. The forage yield of fodder banks was not influenced by season. The mixed fodder bank had greater forage yield (5.1 t DM ha?1) than the L. leucocephala fodder bank (3.4 t DM ha?1) in each season. Additionally, the mixed fodder bank accumulated more forage yield during the experimental period (10.2 t DM ha?1 year?1) than G. ulmifolia (9.0 t DM ha?1 year?1) or L. leucocephala (6.9 t DM ha?1 year?1). The concentrations of CP, C and C:N were not influenced by season. Forage NDF and ADF concentrations were greater in the rainy season (476 g kg?1 DM) compared with the dry season (325 g kg?1 DM). Mixed fodder banks had the greatest N yield (185.9 kg ha?1) and consequently the greatest availability of N (371.8 kg N ha?1 year?1). We conclude that mixed fodder banks of L. leucocephala and G. ulmifolia are a better option for improving productivity and forage quality in comparison with pure fodder banks in Yucatan, Mexico.  相似文献   

19.
We monitored variation in seasonal and annual net ecosystem production (NEP), gross primary production (GPP), and ecosystem respiration (R E) based on 7-year eddy covariance measurements above a cool?Ctemperate deciduous broad-leaved forest (Japanese beech forest). The 7-year means (±SD) of annual NEP, GPP, and R E were 312?±?64, 1250?±?62, and 938?±?36?g?C?m?2?year?1, respectively. Variation in NEP was much larger than variation in GPP and R E. During the growing season, the main factor controlling carbon balance was air temperature; variation in seasonal integrated NEP was regulated by accumulated air temperature (degree-day) with a significant negative correlation, whereas the seasonal ratio of R E to GPP was correlated positively with accumulated air temperature. Because the deviation of seasonal NEP was also significantly correlated with seasonal R E/GPP, NEP was controlled by R E/GPP, depending on air temperature during the growing season. Seasonal R E in the defoliation and snow seasons was also important for evaluating the annual carbon balance, because the total number of days in the two seasons was quite large owing to a long snowy winter. In the defoliation and snow seasons, we found defoliation season length was a major factor determining seasonal integrated R E, illustrating the positive correlation between R E and defoliation season length. The major factors controlling interannual variations in forest carbon balance are discussed.  相似文献   

20.
In the Northern and Baltic countries, grey alder is a prospective tree species for short-rotation forestry. Hence, knowledge about the functioning of such forest ecosystems is critical in order to manage them in a sustainable and environmentally sound way. The 17-year-long continuous time series study is conducted in a grey alder plantation growing on abandoned agricultural land. The results of above- and below-ground biomass and production of the 17-year-old stand are compared to the earlier published respective data from the same stand at the ages of 5 and 10 years. The objectives of the current study were to assess (1) above-ground biomass (AGB) and production; (2) below-ground biomass: coarse root biomass (CRB), fine root biomass (FRB) and fine root production (FRP); (3) carbon (C) and nitrogen (N) accumulation dynamics in grey alder stand growing on former arable land. The main results of the 17-year-old stand were as follows: AGB 120.8 t ha?1; current annual increment of the stem mass 5.7 t ha year?1; calculated CRB 22.3 t ha?1; FRB 81 ± 10 g m?2; nodule biomass 31 ± 19 g m?2; fine root necromass 11 ± 2 g m?2; FRP 53 g DM m?2 year?1; fine root turnover rate 0.54 year?1; and fine root longevity 1.9 years. FRB was strongly correlated with the stand basal area and stem mass. Fine root efficiency was the highest at the age of 10 years; at the age of 17 years, it had slightly reduced. Grey alder stand significantly increased N and Corg content in topsoil. The role of fine roots for the sequestration of C is quite modest compared to leaf litter C flux.  相似文献   

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