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1.
The amount and composition of sugar units comprising polysaccharides in sapwood and heartwood, or stemwood, of 11 industrially important pulpwood species were analysed. The polysaccharide content was between 60 and 80% (w/w) for all species, with cellulose as the predominant polysaccharide type and glucuronoxylans as the main non-cellulosic polysaccharides. The second most abundant non-cellulosic polysaccharides were either pectins, i.e. polygalacturonic acids, or glucomannans. The amount of acidic sugar units were 15–23% of the total amount of non-cellulosic sugar units in all samples, with the Acacia species in the high end. The amount and composition of water-soluble carbohydrates from ground wood samples were also analysed, since these are important in mechanical and chemimechanical pulping, and as a possible source of bioactive polymers. Sapwood released more carbohydrates than heartwood for most species. It is to be noted that the relative amount of dissolved acidic sugar units was larger from the heartwood than from the sapwood for all species. Probably due to the mild treatment conditions, the main dissolved polysaccharides were xylans only for a few samples, while easily soluble galactans, arabinogalactans, or mannans dominated in most species. Pectins dominated in heartwood of Populus grandidentata. Generally, pectins and acidic xylans were the main acidic polysaccharides.  相似文献   

2.
Water soluble polysaccharides from Norway spruce, Scots pine, and Siberian larch were compared. For all species the total amount of polysaccharides isolated from the heartwood was higher than that from the sapwood. The heartwood polysaccharides had a high content of galactose and arabinose units, and some glucuronic acid units, suggesting the presence of acidic arabinogalactans. The total amounts of recovered water-soluble arabinogalactans were 1.9 mg/g for spruce heartwood, 5.3 mg/g for pine heartwood, and as much as 106 mg/g for larch heartwood. The other water-soluble polysaccharides were mainly glucomannans. The average ratio of Gal:Ara:GlcA in the water-soluble arabinogalactans of spruce heartwood was about 4.3:1:1, pine heartwood about 4.5:1:0.2, and larch heartwood about 6.7:1:0.1. The corresponding molar ratios then being about 3.6:1:0.8 for spruce, 3.8:1:0.2 for pine, and 5.6:1:0.08 for larch. Thus, the content of glucuronic acid units was especially high in the spruce heartwood arabinogalactans. The content of arabinose was slightly higher in spruce and pine than in larch heartwood arabinogalactans.  相似文献   

3.
Dissolution and dispersion of spruce wood components into hot water   总被引:1,自引:0,他引:1  
Summary The dissolution and dispersion of components from Norway spruce (Picea abies) wood were examined in laboratory experiments to determine the factors influencing variations in dissolved and colloidal substances in mechanical pulp suspensions. Finely ground, fresh spruce wood was suspended in water at 90 °C and was agitated intensively for up to 12 h, after which the concentrations of dissolved and dispersed lipophilic extractives, lignans, carbohydrates and lignins were determined. Sapwood and heartwood were studied separately. Effects of pH and added electrolytes on the dissolution and dispersion of wood components were also investigated.Higher amounts of lipophilic extractives, and especially of triglycerides, were dispersed from sapwood than from heartwood. The release of lipophilic extractives continued for up to 3 h, after which the concentrations in the suspensions leveled off. At this stage the composition of the dissolved and dispersed lipophilic extractives equaled that of the wood. The amount of lipophilic extractives in the suspensions increased with increased pH, in the range of 4.5–6.7, but was lower in the presence of electrolytes. The dissolution of carbohydrates continued even beyond 3 h of agitation. The high water temperature induced hydrolytic reactions, thereby releasing especially arabinose. The release of arabinose through the hydrolytic cleavage from polysaccharides was more extensive at pH 4.5 than at pH 5.5 and 6.7. More polysaccharides containing galacturonic acid units (pectins) were dissolved at a higher pH. Much more polysaccharides containing glucose, most probably starch, were present in the sapwood suspensions. The dissolution of lignins also continued throughout the 12 h experiment. The measured UV-absorption, after extraction of lignans, was roughly the same for sapwood and heartwood suspensions. Slightly less lignins were released in the presence of electrolytes. Lignans were released only from heartwood.  相似文献   

4.
The fungal pathogen Heterobasidion irregulare has caused substantial damage to pine species in much of the south‐eastern United States, but the potential for damage from Heterobasidion root disease to forest tree species of the Great Lakes Region has not been thoroughly investigated. In each of three trials in two infested stands, fresh stem discs of six native conifer species were exposed to natural inoculum and examined for the asexual Spiniger stage of H. irregulare after incubation in the laboratory. In the first trial, the fungus was infrequently observed on discs, and differences among species were not found. But in the other two trials, detection of the pathogen was much more frequent on discs of tamarack (Larix laricina), red pine (Pinus resinosa) and white spruce (Picea glauca) than those of northern white cedar (Thuja occidentalis), eastern hemlock (Tsuga canadensis) and balsam fir (Abies balsamea). In a second study, the ability of an isolate of H. irregulare to decay root sapwood of these same species was compared using a soil‐block decay test. After 16 weeks of incubation, weight loss due to decay was greatest for red pine (23.2%) and larch (19.1%) and least for balsam fir (1.3%). Differences in the ability of H. irregulare to infect and saprophytically utilize substrate of these important tree species should prompt additional investigation of the relative risk posed to forests of the Great Lakes Region.  相似文献   

5.
Five Japanese timbers, four timbers from the USA, and one Malaysian timber were evaluated for their resistance to the invasive dry-wood termite Incisitermes minor (Hagen) using laboratory choice and no-choice feeding tests with holed specimens. The highest survival rates of I. minor in both the heartwood and sapwood no-choice feeding tests were more than 70% after 3 months. When offered sapwood and heartwood choice feeding tests and the combined choice feeding tests, the highest survival rates of I. minor were more than 75% after 3 months. With regards to the percentage of wood mass losses in the no-choice and choice feeding tests, karamatsu (Larix leptolepsis), buna (Fagus crenata), and Douglas fir (Pseudotsuga menziesii) were classified as “resistant” species among the ten sapwood specimens. In the heartwood no-choice and choice feeding tests, the resistant species were buna, karamatsu, Douglas fir, sugi (Cryptomeria japonica), akamatsu (Pinus densiflora), and western red cedar (Thuja plicata). The ranking of the resistance of the ten commercial timbers against I. minor was buna > karamatsu > sugi > western red cedar > Douglas fir > rubber > western hemlock > hinoki > spruce.  相似文献   

6.
Summary Use of UV-laser excitation to produce fluorescence spectra for heartwood and sapwood from jack pine (Pinus banksiana), white spruce (Picea glauca) and balsam fir (Abies balsamea) was examined. Spectra were fairly broad without sharp spectral features and overlap of spectra between species was common. Sample to sample and in-sample variation of the recorded fluorescence spectra was observed. The fluorescence spectra obtained from heartwood samples of jack pine showed evidence of photochemical bleaching as a result of the multiple laser pulses needed to produce a complete spectrum. Bleaching may have obscured differences between species. For the mix of species examined no sapwood nor heartwood samples were distinguishable by this technique with the detector used. Use of an optical multichannel analyzer (OMA) could reduce the number of laser pulses needed to obtain an entire spectrum. Under these conditions it would be possible to determine whether the minor differences in spectral features observed for the different species are more pronounced in the first few laser pulses and if they are characteristic of species. Certain aspects of the data suggest that with improved analytical equipment UV-fluorescence might prove to be a useful technique for the identification of certain species.  相似文献   

7.
ABSTRACT

Effects of climatic factors and material properties on the development of surface mould growth on wooden claddings were investigated in a laboratory experiment. Specimens of aspen (Populus tremula), Siberian larch (Larix Sibirica), American white oak (Querqus alba), Scots pine (Pinus sylvestris), Norway spruce (Picea abies) and thermally modified pine were incubated in eight climatic chambers at specified wetting periods (2 or 4?h per day), relative humidity (58–86%) and temperature conditions (10–27°C). Surface mould growth was assessed weekly for 13 weeks, and the results were evaluated statistically using Generalized Estimating Equations logistic regression models. All tested climatic factors had significant effects on the mould growth, and there were significant differences between the materials. The ranking of the materials varied with temperature and over time. Aspen, pine sapwood and oak were overall most susceptible to mould growth, and thermally modified pine least susceptible. There were significant differences between sapwood and heartwood for pine and spruce. The effect of density was tested on the spruce heartwood material, but was not found to be significant. The results can be used to further develop prediction models for mould growth on wooden claddings.  相似文献   

8.
Abstract

Untreated Scots pine (Pinus sylvestris) and Norway spruce (Picea abies) samples were exposed above ground in a durability test for 6 years. The samples consisted of three pieces of wood, 22×95×500 mm, screwed together; two pieces lengthwise with a third piece overlapping. Weight was measured, to calculate moisture content (MC), and samples checked regularly for cracks and fungal growth. Parameters investigated were heartwood/sapwood (pine), annual ring orientation (spruce), stand site, annual ring width and density. Stand site, annual ring width and density had no influence on MC or fungal growth for either pine or spruce. Spruce samples with vertical annual rings had fewer cracks than samples with horizontal annual rings. Pine sapwood samples had a high MC and a large amount of rot fungi, while heartwood had a lower MC and no rot. Most spruce samples were similar to pine heartwood, except from a few samples that had high MC and fungal growth. Those were all sawn from the outer part of the log. Therefore, it can be stated that spruce sawn from the inner part has almost the same properties as pine heartwood, while spruce from the outer part of the log has similar properties to pine sapwood.  相似文献   

9.
 Isolated water-soluble acidic arabinogalactans from Norway spruce and Scots pine heartwood were analysed and compared to Siberian larch heartwood arabinogalactans. The carbohydrate monomer composition was determined by acid methanolysis and gas chromatography, while structural studies were performed by 13C NMR spectroscopy and methylation analysis. The main structural features were found to be the same in the three types of arabinogalactans. However, the structure of the arabinogalactans from spruce and pine were found slightly different from the structure of larch arabinogalactans. The amount of single unit side-chains, consisting of arabinose and glucuronic acid units, was higher in the spruce and pine arabinogalactans than in the larch arabinogalactans. The amount of glucuronic acid was higher in the spruce arabinogalactans than in the pine arabinogalactans. The pine arabinogalactans had a higher amount of side chains with more than two sugar units than the spruce arabinogalactans. Received 30 December 1999  相似文献   

10.
Summary The purpose of this study was to examine the influence of the moisture level on the cell-wall material in wood using pulsed proton nuclear magnetic resonance. The wood species used were western hemlock (Tsuga heterophylla (Raf.) Sarg.) and sitka spruce (Picea sitchensis (Bong.) Carr.), distinguishing between heartwood and sapwood regions. The moisture contents of the specimens were below the fibre saturation point and they were conditioned to equilibrium moisture contents based on initial desorption, adsorption and secondary desorption processes. From the FID experiments, the NMR-based moisture contents and the solid-wood lineshape second moments were determined. Average relative proton-spin densities, which were needed to calculate the NMR-based moisture contents, were determined from known moisture contents and they were: hemlock sapwood: 0.616; hemlock heartwood: 0.537; spruce sapwood: 0.679; and, spruce heartwood: 0.446. The average RSD value, considering both heartwood and sapwood, for western hemlock species was 0.577 and for sitka spruce was 0.563; these are close to published RSD values for other species. The condition as to how the equilibrium moisture content was attained did not influence the second moment for hemlock; however, for spruce sapwood, the second moments were sorption dependent. The hemlock M2 decreased from about 5.1 × 109 s-2 at low MNMR to 4.5 × 109 s-2 (heartwood) and 4.3 × 109 s-2 (sapwood) at higher mnmr. The adsorption and secondary desorption M2 for the spruce sapwood region decreased from about 5.0 × 109 s-2 at low mnmr to about 4.1 × 109 s-2 near the MF, whereas M2 for the spruce heartwood decreased from about 4.3 × 109 s-2 at low MNMR to about 3.5 × 109 s-2 near MF. Extractives may have a key role in obtaining the RSD and second moments.This project was financially supported by the Science Council of British Columbia, MacMillan Bloedel Research. The Natural Sciences and Engineering Research Council of Canada is acknowledged for their support of the NMR spectrometer measurements  相似文献   

11.
Summary This study compared the susceptibility of five UK‐grown conifer species to colonization by sapstain fungi in two trials carried out in consecutive years. The conifers consisted of Sitka spruce (Picea sitchensis), Japanese larch (Larix kaempferi), Norway spruce (Picea abies), Scots pine (Pinus sylvestris) and lodgepole pine (Pinus contorta). Freshly cut 1‐m logs were exposed to the available inoculum of sapstain fungi from April to August in a woodland environment in the south east of England. Logs of each species were removed after 1‐, 2‐ and 4‐month exposure and sampled destructively to assess the amount of sapstain. In the second trial, per cent moisture content and concentrations of nitrogen, carbohydrate and phenolic compounds in the sapwood were also measured at the start and end of the trial. After 2 months, only the sapwood of both pine species had significant levels of sapstain; mean values of 37% and 19% for lodgepole pine (year 1 and year 2 respectively) and 12% and 1% for Scots pine. After 4 months, the levels of sapstain in both pine species exceeded 60% in both years. By contrast, very little sapstain developed in the other conifer species with maximum mean values of 10% for Norway spruce, 3.5% for larch and less than 1% for Sitka spruce. Overall, the moisture content of the logs decreased progressively in all species over the length of the trial. However, pine logs tended to retain higher levels of moisture throughout the trial compared with spruce or larch. The relatively higher moisture content of pine sapwood may be closer to the optimal moisture content that sapstain fungi require for infection and colonization, thereby contributing to the increased susceptibility of pine compared with the other conifer species. The pine logs also suffered from some colonization by bark beetles (Ips sexdentatus), which increased the inoculum potential and the opportunity for colonization by sapstain fungi. In addition, particular phenolic compounds in conifer sapwood may play a role in determining the resistance of some species to sapstain. Notably the most resistant species, Sitka spruce, was the only softwood that still retained detectable levels of phenolics in the sapwood to the end of the trial.  相似文献   

12.
Norway spruce [Picea abies (L.) Karst.] heartwood and sapwood have differing wood properties, but are similar in appearance. An investigation was made to see whether near-infrared spectroscopy (NIRS) could be used with multivariate statistics for separation between heartwood and sapwood in dry state on tangential longitudinal surfaces. For classification of wood into sapwood and heartwood, partial least square (PLS) regression was used. Orthogonal signal correction (OSC) filtering was used on the spectra. This study shows that a separation of sapwood and heartwood of spruce is possible with NIR spectra measured in a laboratory environment. The visible-wavelength spectra have significant influence on the predictive power of separation models between sapwood and heartwood of spruce. All 44 specimens in the calibration set were correctly classified into heartwood and sapwood. Validation of the model was done with a prediction set of 16 specimens, of which one was classified incorrectly.  相似文献   

13.
Vertical distribution of leaf area largely governs both tree structure and function. Models of this important tree attribute have been constructed for several commercially important conifers. However, a limited number of studies have compared alternative modeling techniques and inherent species differences. This study used several existing datasets for the five primary conifer species in Maine, namely balsam fir [Abies balsamea (L.) Mill.], northern white-cedar [Thuja occidentalis (L.)], eastern hemlock [Tsuga canadensis (L.) Carr.], eastern white pine [Pinus strobus (L.)], and red spruce [Picea rubens (Sarg.)] to examine species variation in total and vertical distribution of projected leaf area at the individual branch- and tree-levels. In addition, multiple methods for modeling the vertical distribution of leaf area were examined across the species. For a given branch diameter and location within the crown, eastern hemlock branches held the greatest amount of leaf area, followed by balsam fir, northern white-cedar, white pine, and red spruce. At the tree-level, eastern white pine held the greatest amount of leaf area followed by eastern hemlock, balsam fir, red spruce, and northern white-cedar for a given tree size. Across species, the two-parameter, right-truncated Weibull distribution performed the best for predicting vertical distribution of leaf area when compared to the four-parameter beta and Johnson's SB distributions (reduction of root mean square error of 1.7–21.1%). Northern white-cedar had a relative distribution of leaf area distinctly different than other species in this study with a mode shifted towards the upper crown. In contrast to red spruce and white pine, the mode of the relative distribution of leaf area for balsam fir and eastern hemlock occurred lower in the crown. Results of this study suggest that differences in total and vertical distribution of leaf area exist between species, but significant amounts of their variation are largely accounted for by bole and crown size.  相似文献   

14.
Summary Opposite wood, normal side wood, and compression wood were isolated from leaning stems of Abies balsamea, Larix laricina, Picea mariana, Pinus resinosa, and Tsuga canadensis and were subjected to analyses for lignin and relative carbohydrate composition. There were no statistically significant differences between the data obtained for opposite wood and side wood. Contrary to some earlier reports, opposite wood has exactly the same content of lignin, cellulose, and hemicelluloses as has corresponding normal wood.This paper is dedicated to Dean Edwin C. Jahn in honor of his 70th birthday.  相似文献   

15.
Reconstruction in twenty single-cohort stands across five climax series on the eastern slope of the Washington Cascade Range found a variety of species combinations and development patterns. Western larch (Larix occidentalis Nutt.) and lodgepole pine (Pinus contorta Dougl. ex Loud) were found to be very competitive species that usually occupied a dominant position in stands in the Abies grandis, Abies lasiocarpa, and Tsuga mertensiana climax series. Interior Douglas-fir (Pseudotsuga menziesii var. glauca (Mirb.) Franco) was found in all five climax series and, although its height growth was less than that of western larch or the lodgepole pine, it was usually found in the upper stratum. These results suggest that site classification based on climax potential should be used cautiously when applied to young stand management decisions regarding seral species. For example, the difference between an Abies grandis, and a Tsuga mertensiana climax series is quite large in terms of potential productivity but in either case western larch, if present, will likely dominate these stands.  相似文献   

16.
Abstract

The heartwood and sapwood from Scots pine (PS), Norway spruce (PA), and Oriental spruce (PO) were tested for susceptibility to discoloring fungi and water uptake. In addition, annual ring width and density were measured. The methods used were Mycologg for testing growth of fungi and a modified version of EN 927-5 to investigate water uptake. For pine, the heartwood showed a lower water uptake and no discoloring fungi growing in the tests. The heartwood had a significantly higher density and smaller annual ring width than the sapwood. In PA the heartwood had significantly lower discoloration than sapwood. The total water uptake in g/m2 was significantly higher in sapwood, but not the calculated moisture content. As for wood properties, the density was significantly higher in sapwood compared to heartwood, although there were no differences in annual ring width. Regarding PO, differences in water uptake could be seen between sapwood and heartwood although the densities were similar. These results show that susceptibility to discoloring fungi and water uptake is hard to correlate to a single inherent property when looking at different wood species.  相似文献   

17.
  • ? Allometry of sapwood/heartwood is essential for understanding tree growth, water transport and carbon allocation, timber production and use, but such an allometry is lacking for Chinese temperate tree species.
  • ? We studied the allometry and development of heartwood and sapwood for seven Chinese temperate tree species: Korean pine (Pinus koraiensis Sieb. et Zucc), Dahurian larch (Larix gmelinii Rupr.), Japanese elm (Ulmus davidiana Planch var. japonica (Rehd.) Nakai), Manchurian ash (Fraxinus mandshurica Rupr.), Manchurian walnut (Juglans mandshurica Maxim.), Amur cork-tree (Phellodendron amurense Rupr.), and Mongolian oak (Quercus mongolica Fisch.).
  • ? All heartwood parameters investigated, including heartwood radius (HR), heartwood formation rate (HFR), heartwood ring number (HRN), heartwood initiation age (HIA), and heartwood volume ratio (HVR), were positively correlated with tree cambial age (CA). The HR, sapwood width (SW), sapwood area (SA), heartwood and sapwood volumes were significantly related to stem diameter at breast height (DBH) or xylem diameter. There was a polynomial relationship between the sapwood ring longevity (SRL) and sapwood ring number (SRN). However, most of the allometric relationships were species-dependent. The hardwood formation patterns were different between coniferous and broadleaved tree species. A power function was suitable to scale SA from DBH, but the exponent varied from 1.32 for the larch to 2.19 for the cork-tree.
  • ? Our allometry provided a practical means to assess wood development and related physiology for the temperate tree species.
  •   相似文献   

    18.
    Abstract

    This study assessed the decay resistance of Pinus leucodermis wood to the brown-rot fungus Coniophora puteana. Based upon the median weight losses of 30.65% for heartwood and of 34.68% for sapwood obtained in the biological tests, both the heartwood and sapwood material examined was classified as not durable (durability class 5) according to the CEN/TS 15083-1 classification. Total extractives were low, 3.93% in heartwood and 1.00% in sapwood, while lignin content was 22.60% and 25.41% in heartwood and sapwood, respectively. It is highly recommended to use protective treatments before using P. leucodermis wood in outdoor conditions.  相似文献   

    19.
    3种不同处理方法对木材渗透性影响的研究   总被引:2,自引:0,他引:2  
    本文通过对长白鱼鳞云杉和臭冷杉生材分别进行普通气干处理和酒精置换处理以及对其气干材进行水浸处理,研究了这3种不同处理方法对木材气渗透性的影响及其影响机理。研究结果表明,长白鱼鳞云杉边材、心材和臭冷杉心村的生材经普通气干处理后,其气体渗透性较低,分别约为0.114、0.045和0.111darcy;长白鱼鳞云杉边材、心材和臭冷杉心材的生材经酒精置换处理后,其气体渗透性分别约为11.713、0.074和0.144darcy,比普通气干处理对照组试样的平均渗透性分别增加约101.5倍、62%和30%,t检验表明,前者差异非常显著,但后两者差异不显著;已气干18个月的长白鱼鳞云杉边材、心材和臭冷杉心材经水浸处理后,其平均气体浸透性较处理前分别增加约85%、49%、65.5%,t检验表明差异均显著。长白鱼鳞云杉生材边材经  相似文献   

    20.
    SEABY  D. A.; SELBY  C. 《Forestry》1990,63(2):197-207
    Seedlings of eight conifer species, Pinus contorta, Pinus sylvestris,Pinus nigra, Larix kaempferi, Picea abies, Picea sitchensis,Pseudotsuga, nenziesii and Abies grandis, were treated withauxin, in pot and nursery experiments. Dilute solutions of naphthaleneacetic acid (NAA) were applied at two treatment times, justbefore and just after cotyledon expansion. Lateral root numberswere increased by up to 20 times on responsive species. Sensitivityvaried considerably, the three pine species showing a much greaterresponse than the two species of spruce. In most cases the meannumber of induced roots increased steadily with increasing auxinconcentration. Forest nursery results showed that NAA couldprovide an inexpensive method for stimulating lateral root formationnear the soil surface. Practical nursery aspects of treatmentare briefly discussed.  相似文献   

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