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1.
The attacks of bark‐ and wood‐boring Coleoptera on broken conifer stems after severe snow‐breakage in early 1988 were studied in autumn 1988 and 1989. The study included twelve stands in the county of Värmland in central Sweden differing in age and edaphic conditions. The tops on the ground as well as the remaining rooted stem stumps of 94 Picea abies and 61 Pinus sylvestris were inspected. The presence of living branches on the stumps strongly influenced the incidence of insect attack. All stumps without branches were attacked during the two‐year period, whereas only a few spruce stumps with more than ten branches and no pine stumps with more than five branches were attacked. On spruce stumps, the most frequently encountered Coleoptera were all scolytids, i.e. the species Pityogenes chalcographus, Hylurgops palliatus, and the genera Polygraphus, Dryocoetes, and Trypodendron. On pine stumps, Tomicus piniperda was the most common species. Most of the spruce tops and virtually all pine tops were attacked during the two‐year period, and most of these attacks occurred during the first summer. P. chalcographus, Dryocoetes, and weevils of the genus Pissodes were the most frequent Coleoptera on spruce tops. On pine tops, the dominant insects belonged to the genera Pissodes and Pityogenes.  相似文献   

2.
For the forests in the county of Västerbotten in northern Sweden, this study attempts to quantify the non‐timber value arisen from on‐site consumptive use (berry‐ and mushroom‐picking), on‐site non‐consumptive use (hiking, camping, etc.), and off‐site visual experience. The contingent valuation method (CVM) was applied in a mail survey for obtaining empirical data. A follow‐up survey was also conducted in order to get knowledge about the representativeness of the respondents to the CVM survey. The results indicated that the non‐timber value accounted for a considerable portion of the total forest value. On‐site consumptive use was more valuable to rural people than to urban, while on‐site non‐consumptive use was more valuable to urban people. Taken together, these on‐site use values—which to a large extent depend on the Right of Common Access—accounted for two‐thirds of the non‐timber value. The value of the off‐site visual experience thus accounted for one‐third of the non‐timber value.  相似文献   

3.
A transition matrix model of an uneven‐aged Norway spruce (Picea abies (L.) Karst.) stand is presented. The transition probabilities are based on material from 48 study plots in eastern Finland. The ingrowth of the transition matrix is based on material from 8 study plots and on previous studies. In the model runs the sustainable harvest proportion for 5 years varied between 7 and 20% of the stocking depending on the stand basal area. The shape of the stand diameter distribution had no notable effect on the sustainable harvest proportion. The extension of the thinning interval from 5 to 10 years slightly reduced the total removal during the simulation time.  相似文献   

4.
Leader shoots from juvenile succulent Norway spruce seedlings were used as cuttings 11 weeks after sowing. The seedlings were grown for 6 additional weeks and then apically dominant and lateral shoots were also used as cuttings. These three cutting categories, leaders, dominants and laterals, rooted 99.4, 99.0 and 97.5 % respectively and performed differently with respect to rooting development, number of roots per cutting and response to IBA and PIBA treatments (synthetic auxins). A large individual variation in number of roots per cutting, which auxin treatments did not reduce, was present within each cutting categoty. Laterals from the three different families included in the study showed a different variation than did leaders and dominants. Generally, the juvenile succulent cuttings were easy to root. Seedlings grown at a rather high light intensity (62 W/m2) had the potential to give good rooting even without hormonal treatments.  相似文献   

5.
Models for predicting mortality in even‐aged stands were developed. The models rely on data from the Norwegian National Forest Inventory, and were designed for use in large‐scale forestry scenario models. A two‐step modelling strategy was applied: (1) logistic regression models predicting the probability of complete survival occurring,” and (2) multiplicative regression models for stem number reduction and diameter calibration. A joint model for all species predicting the probability of survival occurring on a plot was developed. Separate models for forests dominated by spruce, pine and broadleaved trees were developed for stem number reduction, while no appropriate models for diameter calibration were found. The phenomenon mortality is a stochastic, rare and irregular event, and this was reflected as low R 2 in the models. However, the model performance appeared logical and the results of validations based on independent data were reasonably good, i.e. the presented models may be applied to large‐scale forestry scenario analyses. With new rotations of permanent sample plot measurements, the models should be evaluated and, if necessary, revised.  相似文献   

6.
Two models for determination of the number of stems per hectare in forest stands (N) from attributes derived by aerial photo‐interpretation were developed. The models relied on the assumption that N could be determined by dividing the total stand volume per hectare with the volume of the “average tree”; defined by stand mean height and the diameter corresponding to mean basal area of a stand. Input variables of the models were stand mean height, crown closure and site quality. Additionally, model II required input of average stand volume per hectare and average mean diameter derived from stratified field sample plot inventories. Material for 143 coniferous stands was used for the testing of the models. The stands were recorded by intensive field measurements. Aerial photographs at the approximate scale of 1:15 000 were used for photo‐interpretation. The N value was underestimated in model I by 5.4–47.0%. The standard deviation for the differences was 15.2–26.2% for mature stands and 41.4–44.2% for young thinning phase stands. For model II, the mean difference between the predicted and observed N value was in the range ‐16.1% to 12.2%.  相似文献   

7.

Context and aim

This study aimed to examine the effect of the tension wood G‐layer on the viscoelastic properties of wood.

Methods

Tension wood and opposite wood samples were obtained from six French Guianese tropical rainforest species (Sextonia rubra, Ocotea guyanensis, Inga alba, Tachigali melinoni, Iyranthera sagotiana and Virola michelii); the tension wood of the former three of these species had a G‐layer, whilst the tension wood from the latter three had no G‐layer. Tensile dynamic mechanical analysis (DMA) was performed on green never dried wood samples in the longitudinal direction with samples submerged in a water bath at a temperature (30°C) and frequency (1 Hz) representative of the conditions experienced by wood within a living tree. Then, DMA was repeated with samples conditioned to an air-dried state. Finally, samples were oven-dried to measure longitudinal shrinkage.

Results

Tension wood did not always have a higher longitudinal storage (elastic) modulus than opposite wood from the same tree regardless of the presence or absence of a G‐layer. For the species containing a G‐layer, tension wood had a higher damping coefficient and experienced a greater longitudinal shrinkage upon drying than opposite wood from the same species. No difference was found in damping coefficients between tension wood and opposite wood for the species that had no G‐layer.

Conclusion

It is proposed that the different molecular composition of the G-layer matrix has an influence on the viscoelasticity of wood, even if a biomechanical gain is not yet clear. This study shows that rheological properties and longitudinal shrinkage can be used to detect the presence of a G‐layer in tension wood.
  相似文献   

8.
Today sawmills have started to use automatic methods for log grading. The methods used are either optical or gamma‐ray scanners. However, the signals from these scanners are too coarse for accurate log grading and for good control of the sawing process at the single log level. The objective of the present study was to determine the grading accuracy of a log‐scanner with two industrial X‐ray sources. The grading accuracy was compared with the accuracy of manual grading. The results showed that the manual grading of logs and boards is difficult. The accuracy of manual grading was low and the automatic grading systems were more reliable than manual ones. Possibilities for improving the automatic grading systems are discussed.  相似文献   

9.
Self‐thinning models were developed to describe the relationship between the stem number and mean diameter of even‐aged and monospecific tree stands undergoing self‐thinning. The models were developed separately for Pinus sylvestris L., Picea abies (L.) Karst. and Betula pendula Roth. stands. Data from 41 unthinned permanent sample plots were used. According to Reineke's equation and the —3/2 power rule of self‐thinning, a log‐log plot of average tree size and stem density will give a straight, self‐thinning line of constant slope. According to this study, the slope of the line consistent with Reineke's equation varies for different tree species. Within tree species, the intercept of the self‐thinning line varies according to site index.  相似文献   

10.
The defence of Norway spruce against a combined attack of the bark beetle Ips typographus and its associated blue‐stain fungi is based upon a) constitutive resin stored in ducts of the bark and sapwood, and b) induced resinosis in reaction zones surrounding the point of infection. Empirically, beetle epidemics are associated with external stresses, drought being a particularly prominent factor. An attempt was made to mechanistically explain the apparent link between drought and infestations, through studying after‐effects of drought on the induced defence in stressed trees. In the field, 3–5.5 m tall trees were exposed to artificial drought over three growth seasons to investigate whether this treatment would predispose them to fungal infection in a fourth season when drought was absent. Pre‐dawn xylem water potentials down to ‐1.85 MPa and a considerable foliage depletion were recorded. In the fourth season, the trees were inoculated with Ophiostoma polonicum, a pathogenic associate of Ips typographus. No difference in susceptibility could be seen between drought stressed trees and unstressed controls. Carbohydrate concentrations of foliage and branch bark were slightly enhanced and mineral nutrient concentrations strongly reduced in stressed trees.  相似文献   

11.
Altogether 1 080 two‐year‐old silver birch (Betula pendula Roth) were planted in groups of 36 plants at aspacing of 20 x 20 Cm in spring 1983 on a nursery field at lat. 60° 15 N and long. 16°00 E. In spring 1984 all plants were cut with 10 cm high stumps and light screens were arranged on the stump groups. Relative ¡rradiance (RI) 5, 10, 25, 50, 75 and 100% of full sunlight were used. After five growing seasons, all stumps exposed to 5 or 10% light intensity were dead, and 64–94% of stumps exposed to 25–100% RI were alive. 1.3–1.4 sprouts per living stump were living five years after cutting. The mean height of sprouts was highest on stumps exposed to 100% light. Also dry weights of leaves and sprouts, number of leaves per sprout, and leaf area were highest on stumps exposed to full sunlight. Stumps producting only one sprout five years after cutting have higher height and dry weight values than stumps with more sprouts. The biomass production per hectare and year for silver birch sprouts at a spacing of 0.2 m and with 90% survival was calculated as 5.4 tonnes. The biomass of leaves was 8.1 tonnes/ha/year five years after cutting.  相似文献   

12.
A stage‐structured model for projecting stand growth and yield is coupled with a gradient‐based algorithm to determine optimal treatment regimes for Norway spruce (Picea abies Karst.) plantations. The growth model forecasts the movement of trees between 3 cm diameter classes using a growth matrix that includes nonlinear equations for diameter growth. Because harvest treatments may be specified by tree diameter class, the model allows the determination of the optimal thinning timing, thinning intensity, and thinning type simultaneously with planting density and clearfelling age. The optimization model is used to determine treatment regimes with and without thinning for a wide range of values for economic and biological parameters. Results show that the optimal economic planting density is highly sensitive to interest rate, logging costs, planting costs and site class. For a given set of economic parameters, the optimal planting density is not sensitive to the number of thinnings performed or the thinning intensity. Top thinning is the optimal method for every regime in which thinning is scheduled. Results indicate that managers have great flexibility in choosing thinning schedules: few thinnings and shorter rotations give almost the same economic results as more frequent thinnings and longer rotations. Major uncertainties in these results and areas for further research are discussed.  相似文献   

13.
The variance of general combining ability (GCA) and specific combining ability (SCA) for height and diameter growth were estimated from a 13 years old provenance hybridization study in Norway spruce. The GCA variance component for both height and diameter growth was significant, but there was no significant SCA variance. Hybrid progenies outperformed the progenies from stands, but this superiority was not interpreted as a provenance heterosis effect. Estimated family heritabilities for height and diameter growth were moderately high (0.40 for both traits). A consequence of the low SCA variance for Norway spruce may be that clonal testing should replace progeny testing for estimating GCA.  相似文献   

14.
Stand differentiation, heritabilities and genetic correlations in 16 growth, bud and branch traits in Pinus contorta subsp. latifolia were studied for two growth periods in the greenhouse. Data for analyses were based on 116. open‐pollinated families from 33 stands in one breeding region in central Alberta, Canada. Family effect was significant at the 1% level of probability for all traits. The narrow‐sense heritabilities (h2) from 0.43 + 0.09 to 0.71 + 0.12 for traits of absolute height and diameter were the highest reported for growth traits in this conifer. In contrast, h2 varied between 0.15±0.06 and 0.33 ± 0.08 for bud, branch and growth increment traits. Except for relative branch length, all traits showed convariation with the other traits. Levels of heritability and genetic correlation suggest that selection procedures that exploit additive genetic variance and covariance might be practised, provided there would be high juvenile‐mature correlation for this sample of P. contorta subsp. latifolia.  相似文献   

15.
The occurrence of fungi in the underground parts of 1‐ and 2‐year‐old conifer stumps (Pimis sylveslris and Picea abies) was studied in two clear‐felled areas of eastern Uppland in Sweden, and most fungal isolates were identified to species.

Decay fungi (Basidiomycetes) were more frequent in spruce stumps than in pine stumps, and their incidence was highest in 2‐year‐old stumps. The occurrence of blue‐stain fungi and hyaline mycelia did not differ significantly between stumps of different species or age. Within stumps, decay fungi were most frequent in large roots close to the cut surface, whereas blue‐stain fungi and hyaline mycelia were most abundant in small roots. The occurrence of yeasts, bacteria and fast‐growing moulds was also recorded.  相似文献   

16.
The long‐term effects of lime application on fine roots of Norway spruce, Picea abies (L.) Karst, and Scots pine, Pinus sylvestris (L.), have been studied in five experimental forest stands subjected to different lime applications 5 to 18 years before the present study was undertaken. The effects of liming does not seem to significantly influence fine‐root development in forest stands in the long term. The only response to liming in measured root variables was a tendency to increased specific root length (SRL = fine‐root length/fine‐root dry weight, m/g). A correlation between increased SRL, decreased root biomass and increased stem volume growth was indicated. Changes in water extractable amounts of mineral elements—P, K, Ca, Mg, Mn, S, Al and Fe‐in bulk soil and rhizosphere soil from the mineral soil layers were studied in a control area and an area treated with 3830 kg CaCO3 ha‐1. Few significant differences were found between treatments, and then mainly in the case of Ca.  相似文献   

17.
First‐year seedlings of five latitudinal populations of Acer platanoides were subjected to decreasing photoperiod treatment under three different temperature regimes. The depth of the induced dormancy was quantified as the number of days to bud burst (DBB) under defined conditions favourable to growth. The results suggested a close relationship between autumn temperature and the strength of the induced dormancy, with high temperatures combined with short days leading to a deeper stage of dormancy. Northern and continental populations generally had bud burst earlier than southern. The results are discussed in relation to hypotheses for dormancy induction and release.  相似文献   

18.
Defoliation of conifers occasionally precedes bark beetle attacks, suggesting that a severe loss of foliage and ensuing reductions in carbohydrate availability may enhance host tree susceptibility. To shed light on this question, different degrees of defoliation on young Picea abies were simulated by removing whole whorls of branches from below, the trees retaining 100, 50, or 25% of their original crown biomass. After one week or one year, the trees were inoculated with Ophiostoma polonkum, a tree‐killing fungus transmitted by Ips typographus.

Fungal proliferation and tree mortality increased with increasing levels of pruning. Pruning reduced stem diameter growth, but not carbohydrate reserves in foliage and bark. Foliar N, P, and Ca increased with increasing pruning. The results lend support to the hypothesis that a reduction in the photosynthesis capacity increases host tree susceptibility to a beetle‐fungus attack, and that induced defence against infection depends on efficient translocation of assimilates to the sites of infection.  相似文献   

19.
The effect of soil scarification on decomposition of green Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) needles and mineral nutrient release was investigated. The treatments studied were ploughing, disk trenching, mound on inverted humus and untreated control. The study was conducted on two clear‐felled sites with different ground moisture conditions, located in south Sweden. The needles decomposed and released their contents of N and Ca considerably faster on scarified areas than on untreated ones. For P, Mg and especially K, only minor differences between scarified and untreated areas were established. At the wet site, needle decomposition and nutrient mineralization were generally affected to the same extent by all three scarification methods studied. At the dry to fresh site, mounding and ploughing enhanced needle decomposition and nutrient mineralization more than disk trenching. The effect of different scarification methods on the long‐term production capacity of soils is discussed.  相似文献   

20.
Clonal material propagated as cuttings and control seedling lots of Norway spruce (P. abies [L.] Karst.) were compared in two series of clonal tests. Variance components, heritability and expected genetic gain were calculated for height based on measurements carried out in the tests at the age of seven and nine years. The mean heights of the clones were significantly higher than those of the controls in one of the test series, but there was no significant difference between these two groups in the other series. The variance due to clone sources was significant in both test series and accounted for 10 to 19% of the total variation. Broad sense heritabilities were 0.21 ± 0.05 and 0.23 ± 0.02 on the individual level and 0.75 ± 0.06 and 0.78 ± 0.02 on the basis of the clone mean for the respective series. Theoretical genetic gains were calculated based on the assumption that the tallest 10% of the clones would be selected and propagated for reforestation purposes. The expected gain in height was 13.1 and 19.1% in these test series. Utilization of this gain is, however, difficult due to the ageing of the clones.  相似文献   

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