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1.
Dry Douglas-fir (Pseudotsuga menziesii) forests offer a wide range of timber and non-timber values, which may benefit from a balanced timber harvest by variable retention systems with conservation of biodiversity. A major component of biodiversity are forest floor small mammal communities whose abundance and diversity serve as ecological indicators of significant change in forest structure and function from harvesting activities. This study was designed to test the hypotheses that abundance, reproduction, and survival of (i) the southern red-backed vole (Myodes gapperi, formerly Clethrionomys gapperi), will decline; (ii) the deer mouse (Peromyscus maniculatus), will be similar; and (iii) the meadow vole (Microtus pennsylvanicus) and northwestern chipmunk (Tamias amoenus), will increase, with decreasing levels of tree retention. Small mammal populations were live-trapped from 1994 to 1997 in replicated sites of uncut forest, 20% and 50% volume removal by single tree selection, 20%, 35%, and 50% patch cuts based on openings of 0.1–1.6 ha, and small 1.6 ha clearcuts in Douglas-fir forest near Kamloops, British Columbia, Canada. M. gapperi dominated the small mammal community, starting with an abundance of 74–98 animals/ha with mean values ranging from 33 to 51 animals/ha. In the two post-harvest years, abundance, reproduction, and survival of M. gapperi populations were consistently similar among uncut forest and the various levels of tree retention. Thereafter, M. gapperi was seldom found on the small clearcuts. M. pennsylvanicus, T. amoenus, and P. maniculatus occurred predominantly in clearcut sites. As with other types of forest disturbance, responses to our treatments were species-specific. The most striking result was the high abundance and productivity of M. gapperi populations in a dry forest ecosystem, a novel result for this bio-indicator species of closed-canopy forest conditions. At least with respect to small mammals, the retention systems studied seem to enable timber extraction and maintenance of mature forest habitat in these dry fir ecosystems.  相似文献   

2.
The sub-Antarctic biome of South America is the world's southernmost forested ecosystem and one of the last remaining wilderness areas on the planet. Nonetheless, the region confronts various anthropogenic environmental impacts, such as the invasive North American beaver (Castor canadensis) and timber harvesting, particularly in stands of Nothofagus pumilio. Both of these disturbances can affect terrestrial and aquatic systems. To understand the influence and relative importance of these disturbances on sub-Antarctic watersheds, we characterized in-stream and riparian habitat conditions (pH, dissolved oxygen, conductivity, temperature, stream size, distance to riparian forest, bank slope, substrate heterogeneity, benthic organic matter) and benthic macroinvertebrate community structure (density, richness, diversity, evenness) and function (biomass, functional feeding group percent) in 19 streams on Tierra del Fuego Island. To explain the effects of beaver invasion and timber harvesting, we compared these physical and biotic variables among four habitat types: (a) beaver meadows, (b) shelterwood cut harvested areas without forested riparian zones, (c) variable retention harvested areas with riparian buffers, and (d) unmanaged old-growth primary forests. Most habitat variables were similar at all sites, except for dissolved oxygen (significantly higher in streams from old-growth primary forests). Benthic communities in beaver meadows had significantly lower diversity, compared to streams of unmanaged old-growth primary forests, and managed sites presented intermediate values between the two. Functionally, the benthic community in beaver meadows displayed a reduction of all functional feeding groups except collector-gatherers; again variable retention harvested areas with riparian buffers were similar to unmanaged old-growth primary forest streams, while shelterwood cut harvested areas occupied an intermediate position. These results indicated that current forestry practices that include both variable retention and legally mandated riparian forested buffers may be effective in mitigating impacts on stream benthic communities. Finally, these data demonstrated that C. canadensis invasion was a relatively larger impact on these streams than well-managed forestry practices.  相似文献   

3.
Large herbivores have potential to affect invertebrate community structure through numerous processes, but little work has been done to evaluate the relative importance of direct and indirect factors. In this study, we measured arthropod community assemblages on Ceanothus fendleri A. Gray (Fendler's ceanothus) plants that were growing inside and outside of 4-m2 exclosures. We used univariate analyses and structural equation modeling (SEM) to evaluate relationships within this herbivore–plant–arthropod system in ponderosa pine (Pinus ponderosa Laws.) forests of northern Arizona, USA. Results showed that individual arthropod abundance, family diversity, family richness, and functional group richness were significantly greater on plots where C. fendleri plants were protected from large ungulate herbivores (e.g., mule deer (Odocoileus hemionus) and Rocky Mountain elk (Cervus elaphus nelsoni)) than on unprotected plots in each of the three study years. Results also indicated the following: (1) arthropod abundance was significantly greater on protected plants than unprotected plants; (2) rarefaction curves suggested arthropod family richness was similar between protected and unprotected plants in two of the three years when scaled by number of individuals but the estimated total richness was consistently higher on protected plants; (3) arthropod abundance was directly affected by protection from herbivores, plant stem length, and number of flowers; (4) arthropod family richness was related to the number of individuals collected and affected by stem length. Results from this study illustrate that arthropod communities are directly affected by foraging vertebrate herbivores as well as indirectly affected through complex plant-mediated factors in this model system. Protection of preferred forage plants such as C. fendleri from ungulate herbivores can potentially increase diversity of arthropod assemblages in these forests, help conserve biological diversity, and enhance ecosystem restoration efforts.  相似文献   

4.
We used a 5-decade chronosequence of harvest openings to characterize population and community-level responses of small mammals to forest management targeting oak regeneration in southern Indiana. Live-trapping at 42 different sites allowed modeling of occupancy and relative abundance using environmental covariates while incorporating imperfect detection. Species richness was higher in smaller openings on southwest-facing aspects. Similarity between species richness of different age classes decreased with increasing site age. Eastern chipmunk (Tamias striatus) relative abundance was greater in early seral stages, i.e., at young sites with low basal areas. Relative abundance of white-footed mice (Peromyscus leucopus) exhibited different responses to coarse woody debris on sites versus microsites. Pine voles (Microtus pinetorum) and short-tailed shrews (Blarina brevicauda) were more likely to occupy older sites. We observed a greater relative abundance of short-tailed shrews at sites with steep and northeast-facing slopes. Northeast-facing slopes also resulted in higher short-tailed shrew occupancy rates. Incorporating detection probability enabled us to derive more accurate estimates of relative abundance and, when coupled with a Bayesian framework, permitted the estimation of occupancy for uncommon species. Our estimated responses can be used by forest managers to determine the potential impacts of even-aged and uneven-aged oak management on small mammals, and the statistical methodology we used can be applied even more broadly to improve understanding of wildlife responses to forest management.  相似文献   

5.
The establishment of terrestrial buffer zones around vernal pools has been recommended to provide upland habitat for pool-breeding amphibians in areas where forestry practices occur adjacent to breeding sites. However, few studies have empirically tested the effectiveness of buffers. We assessed post-breeding emigration behavior (net emigration distance, rate of movement, proportionate use of available habitats) of radio-tagged adult wood frogs (Lithobates sylvaticus) at nine vernal pools with experimental forest buffer treatments in central Maine, USA. Buffer treatments were either 30-m (N = 3) or 100-m (N = 3) forest buffers surrounded by a 100-m wide clearcut; pools surrounded by uncut forest served as reference sites (N = 3). We tracked 33 individuals in 2004 and 2005, for an average of 41 days, as they emigrated from breeding pools. Recently clearcut habitat was permeable to emigrating adult wood frogs, particularly females. A higher proportion of frogs at 30-m buffer sites than at 100-m buffer sites traveled through the clearcuts to reach intact forest beyond, suggesting that 30-m buffers may not provide sufficient upland habitat to support adult wood frog populations. There was high variability in emigration behavior among frogs, regardless of buffer treatment, and males and females tended to exhibit differential responses to different buffer sizes and to clearcut habitat. Although wood frogs in this study utilized both 30-m and 100-m forest buffers, variability between sexes and density-dependent effects could render small buffers (e.g., 30 m or less) inadequate to support these populations.  相似文献   

6.
Wildfires and timber harvest are two of the most prevalent disturbances in North American forests. To evaluate and compare their impact on small mammals, I conducted meta-analyses on (1) the effect of stand-replacement wildfires and several types of forest harvest (clearcutting followed by burning, clearcutting, and uniform partial harvest) on the abundance of deer mice (Peromyscus maniculatus) and red-backed voles (Myodes gapperi), (2) the impact of clearcutting and partial harvest on a broader array of small mammal species, and (3) the responses of small mammals to recent and older clearcuts (i.e. less than 10 years vs. 10–20 years after harvest). In coniferous and mixed forest, all disturbances except for partial harvest triggered significant increases in the abundance of deer mice and declines in red-backed voles. The increase in deer mice after wildfire was stronger than after clearcutting and marginally stronger than after clearcutting and burning. The abundance of red-backed voles was greatest in undisturbed or partially harvested stands, intermediate after clearcutting, and lowest after wildfire or clearcutting and burning. While the positive effect of clearcutting on deer mice did not persist beyond 10 years after disturbance, the negative effect on red-backed voles was similar between recent and older clearcuts. In deciduous forest, clearcutting did not result in a consistent change in abundance of deer mice and red-backed voles. For other small mammals, recent clearcutting tended to increase the abundance of yellow-pine chipmunks (Tamias amoenus), and meadow and long-tailed voles (Microtus pennsylvanicus and Microtus longicaudus). Woodland jumping mouse (Neozapus insignis), masked shrew (Sorex cinereus), and short-tailed shrew (Blarina brevicauda) did not show consistent response to timber harvest. Overall, the impact of different disturbances on the abundance of small mammals (i.e. positive or negative) appears to be species-specific, but disturbance type may influence the magnitude of this effect. Disturbance types can be ranked from severe to mild in terms of small mammal responses. The effects of forest harvest on small mammals are not equivalent to those of wildfire.  相似文献   

7.
Voles of the genera Microtus and Myodes feed on tree seedlings planted on cutover forest land in temperate and boreal forests of North America and Eurasia. This damage may have serious economic implications as well as limit regeneration of appropriate tree species in certain forest ecosystems. Prediction of vole population outbreaks and feeding damage to forest plantations, across even a limited geographic range, has yet to be achieved in North America. Thus, a major objective was a detailed analysis of changes in population dynamics of long-tailed voles (Microtus longicaudus), and to test three hypotheses (H) that vole populations would: (H1) rise and fall in accordance with the abundance of herbaceous plants (grasses and forbs) during early vegetative succession after forest harvesting, (H2) be positively associated with grass-seeded sites; and (H3) incidence of feeding damage to seedlings would be positively associated with vole abundance. Voles were live-trapped for 6 years (2004–2009) from the time of harvesting on intensive sites, as well as surveyed over a range of extensive sites. Population numbers were related to habitat characteristics and tree damage in young forest plantations near Golden, British Columbia, Canada.  相似文献   

8.
We examined the effects of white-tailed deer (Odocoileus virginianus) herbivory and microsite limitation on Tsuga canadensis regeneration in 39 randomly selected remnant T. canadensis stands in Michigan's Upper Peninsula. Deer of the region migrate to and congregate in T. canadensis stands in winter resulting in strong seasonal habitat use patterns. In each study stand, we quantified vegetation, microsite availability, and deer use (via pellet counts). While some stands contained high densities of T. canadensis regeneration (stems < 4.0 cm dbh), we found complete T. canadensis regeneration failures in 6 out of 39 stands. Additionally, 17 and 22 stands respectively, had complete failures in the small and large sapling categories. General linear models suggested that deer use was the primary limiting factor in the small sapling size class, even at relatively low levels of deer use. T. canadensis seedling density was positively associated with the availability of high-decay coarse woody debris and negatively associated with basal area of Acer saccharum in the overstory. This latter association may be due, at least in part, to negative effects of broadleaf litter on T. canadensis establishment and a general trend toward increasing Acer abundance in the regeneration layer. Our results suggest that differential tolerance to browsing (Tsuga vs. Acer) in conjunction with reduced germination substrate availability may set up a scenario where successful T. canadensis establishments is more limited by legacy and indirect than direct effects given contemporary levels of deer use.  相似文献   

9.
Predation risk is an important factor influencing the distribution of ungulates and their impact on forest structure. However, simultaneous predation risk by wolves and humans is rarely considered in the analyses of habitat selection by ungulates. We counted ungulate pellets on transects to analyse the influence of wolves and humans on ungulate density distribution in the Białowieża Forest, Poland. We assessed whether (1) forest exploitation influenced ungulate habitat selection, (2) ungulate density was higher in areas without human hunting, (3) ungulates avoided the surroundings of a major road, (4) prey density was higher in the strife zone between home ranges of wolf packs both in the presence and absence of human hunting, (5) ungulates avoided areas selected by wolves, and (6) wolf kill sites were in high prey density areas. Red deer (Cervus elaphus) and wild boar (Sus scrofa) selected unexploited over exploited forests and areas without hunting, whereas roe deer (Capreolus capreolus) preferred exploited forests. Wild boar and European bison (Bison bonasus) avoided the area within 300 m of a major road, whereas we could not detect any avoidance by red or roe deer. Prey density was not higher in wolf strife zones, regardless of human hunting. Ungulates did not avoid areas selected by wolves. Wolves killed red deer in areas with prey density of about 4 red deer/km2, regardless of whether the average red deer density in those areas was higher or lower. We conclude that habitat alteration by forest exploitation and hunting by humans influenced the density distribution of ungulates more than predation risk by wolves.  相似文献   

10.
Voles and shrews are key species in northern forest ecosystems. Thus, it is important to quantify to what extent new forestry practices such as planting of non-native tree species impact these small mammals. In northern Norway stands of coastal subarctic birch forests have increasingly been converted to non-native spruce stands during the last century. This leads to changes in the forest floor vegetation and soil conditions that can be expected to negatively impact the community of ground-dwelling small mammals. In this 10-year trapping study we contrasted seasonal small mammal population abundances in spruce plantations with four birch forest varieties. Six different small mammal species were trapped (in descending order of abundance; common shrew Sorex araneus, red vole Myodes rutilus, field vole Microtus agrestis, grey-sided vole M. rufocanus, pygmy shrew S. minutus and water shrew Neomys fodiens). None of the voles appeared to exhibit temporal dynamics resembling population cycles. The three most numerous species were clearly less abundant in the spruce plantations compared to the other forest types. Autumn abundances were most impacted by spruce plantations, indicating that growth rates in the reproductive season were more influenced than winter declines. Species associated with productive forest habitats (i.e. field vole and common shrew) were most impacted by tree species conversion. Still young spruce plantations inter-mixed with birch trees and the ecotone habitat, sustained small mammal abundances comparable to the native birch forests. This implies that managing spruce plantations to maintain a mix of different tree species and high spatial heterogeneity (i.e. more ecotones), will reduce the negative impacts on the small mammal community. On the contrary, if young spruce plantations, as they age become spruce monocultures covering larger parts of the landscapes than they do presently, the negative effects on small mammal communities may be larger than observed in the present study.  相似文献   

11.
The fisher (Martes pennanti) is a forest mustelid endemic to North America that has experienced range reductions in Pacific states that have led to their listing under the Endangered Species Act as warranted but precluded by higher priorities. The viability of the southern Sierra Nevada fisher population is of particular concern due to its reduced historical range, isolated nature, and low genetic variability. We located resting structures of radio-collared fishers in the southern Sierra Nevada and compared resting and available habitat to examine selection for specific features of resting sites. Resting structures provide protection from predators and unfavorable weather and are believed to be the most limiting habitat element across fisher home ranges. Resting structures were found primarily in live trees (76%) and snags (15%). Trees used by fishers for resting were among the largest available and frequently had mistletoe infestations. Ponderosa pines (Pinus ponderosa) were used more often than expected and incense cedars (Calocedrus decurrens) less than expected. Snags were also large and in fairly advanced stages of decay. Habitat at fisher resting sites had higher canopy cover, greater basal area of snags and hardwoods, and smaller and more variable tree sizes compared to random sites. Resting sites were also found on steeper slopes and closer to streams. Canopy cover was consistently the most important variable distinguishing rest and random sites. In western North America, fishers are generally associated with late-successional forests, but changes in these forests due to logging and fire suppression have resulted in a transition to forest stands characterized by fewer large trees and more small stems. These conditions are consistent with our finding that the large rest structures were surrounded by smaller than average trees. Management practices that support the growth and retention of greater numbers of large trees and snags, while maintaining a minimum of 61% (based on moosehorn) or 56% (generated via Forest Vegetation Simulator) canopy cover and a complex horizontal and vertical forest structure, can improve and provide for future fisher habitat.  相似文献   

12.
Large wildfire events in coniferous forests of the western United States are often followed by postfire timber harvest. The long-term impacts of postfire timber harvest on fire-associated cavity-nesting bird species are not well documented. We studied nest-site selection by cavity-nesting birds over a 10-year period (1994–2003), representing 1–11 years after fire, on two burns created by mixed severity wildfires in western Idaho, USA. One burn was partially salvaged logged (the Foothills burn), the other was primarily unlogged (the Star Gulch burn). We monitored 1367 nests of six species (Lewis’s Woodpecker Melanerpes lewis, Hairy Woodpecker Picoides villosus, Black-backed Woodpecker P. arcticus, Northern Flicker Colaptes auratus, Western Bluebird Sialia mexicana, and Mountain Bluebird S. currucoides). Habitat data at nest and non-nest random locations were characterized at fine (field collected) and coarse (remotely sensed) spatial scales. Nest-site selection for most species was consistently associated with higher snag densities and larger snag diameters, whereas wildfire location (Foothills versus Star Gulch) was secondarily important. All woodpecker species used nest sites with larger diameter snags that were surrounded by higher densities of snags than at non-nest locations. Nests of Hairy Woodpecker and Mountain Bluebird were primarily associated with the unlogged wildfire, whereas nests of Lewis’s Woodpecker and Western Bluebird were associated with the partially logged burn in the early years after fire. Nests of wood-probing species (Hairy and Black-backed Woodpeckers) were also located in larger forest patch areas than patches measured at non-nest locations. Our results confirm previous findings that maintaining clumps of large snags in postfire landscapes is necessary for maintaining breeding habitat of cavity-nesting birds. Additionally, appropriately managed salvage logging can create habitat for some species of cavity-nesting birds that prefer more open environments. Our findings can be used by land mangers to develop design criteria for postfire salvage logging that will reserve breeding habitat for cavity-nesting birds.  相似文献   

13.
We examined the short-term effects of group-selection harvesting with seed-tree retention on ground-dwelling and bark-dwelling arthropod communities in a northern hardwood forest in the Upper Peninsula of Michigan. Arthropods were sampled in 16 group-selection openings and 8 closed canopy reference plots. Two opening sizes were examined—radii of 0.5 (320 ± 27 m2, n = 8) and 1.0 (1217 ± 62 m2, n = 8) times the mean canopy height (22 m). Each opening and reference plot was centered on a single Betula alleghaniensis Britt. (yellow birch). Ground-dwelling arthropods were sampled using pitfall traps that were opened for two 1-week periods (rounds 1 and 2), and bark-dwelling arthropods were sampled with sticky traps attached to the centrally located B. alleghaniensis trees. Family-level diversity of ground-dwelling arthropods was lower in reference plots than in the openings, but the only significant difference occurred during round 2, between the matrix and large openings (P < 0.01). During both sampling periods, the ground-dwelling community exhibited a distinct clustering of family-level composition along environmental gradients such as opening size. Families such as Staphylinidae (rove beetles) and Trombidiidae (red velvet mites) were not favored by higher canopy openness while families such as Acrididae (grasshoppers) and Lycosidae (wolf spiders) were captured more in openings than in the forest matrix. Landing rates of wood-boring insects such as Buprestidae (metallic wood-boring beetles) and Xiphydriidae (wood wasps) were significantly higher on seed trees in group-selection openings than in reference plots (P < 0.05). Our results suggest that integrating group-selection openings within northern hardwood forests can lead to an increase in the family-level diversity of ground-dwelling arthropods, at least in the short term. Furthermore, seed trees left in such openings may be more attractive to bark- and wood-boring insects, which warrants further investigation into the susceptibility of these seed trees to damage by certain insect pests.  相似文献   

14.
Ruffed grouse (Bonasa umbellus; hereafter grouse) populations in the central and southern Appalachians (CSA) are declining due to widespread maturation of forest cover. Effective management of this species requires a sex- and age-specific understanding of habitat preferences at multiple temporal and spatial scales. We used multivariate logistic regression models to compare habitat within 1440 grouse home ranges and 1400 equally sized buffered random points across 7 CSA study areas. On most sites, grouse home ranges were positively associated with roads and young forest (<20 years old). Sex and age status affected habitat preference. In general, males used younger forest than females, likely because of differences in habitat use during reproductive periods. Juveniles had fewer vegetation types preferred by adult grouse and more of the avoided vegetation types within their home ranges, indicative of competitive exclusion. Adult females had the greatest specificity and selectivity of habitat conditions within their home ranges. Habitat selection varied among seasons and years on most sites. Winter habitat use reflected behavior that maximized energy conservation, with open vegetation types avoided in the winter on the northernmost study areas, and topography important on all areas. Summer habitat selection reflected vegetation types associated with reproductive activities. Scale influenced habitat preference as well. Although roads and forest age predominantly influenced grouse home range location within the landscape, mesic forest types were most important in determining core area use within the home range. This was likely a result of increased food availability and favorable microclimate. Habitat management efforts should attempt to maintain ∼3–4% of the landscape in young forest cover (<20 years old), evenly distributed across management areas. Roads into these areas should be seeded as appropriate to enhance brood habitat and provide travel corridors connecting suitable forest stands.  相似文献   

15.
Red-shouldered hawks (Buteo lineatus) are threatened in Wisconsin and when nest sites are found during the cruising or marking stage of timber harvesting, the harvest is altered to accommodate the hawks. If nest site locations are known before initiation of timber harvest, foresters can employ a proactive approach to manage red-shouldered hawks while maintaining timber production. We searched for red-shouldered hawks nest sites on Marinette County Forest (MCF) which encompasses 94,000 ha in northeastern Wisconsin and is the second largest county forest in the state. We used a comparative modeling approach to evaluate distribution and habitat relationships of red-shouldered hawk nest sites in relation to a suite of environmental variables in MCF. Models were used to develop forest management recommendations for red-shouldered hawks in Wisconsin. During the spring of 2006 and 2007, we broadcasted conspecific calls to survey 1121 calling stations along forest roads and trails. We located 20 and 25 active nesting territories in 2006 and 2007, respectively (11 of which were active in both years). To understand nest site selection, we measured 22 habitat variables within 0.04-ha plots at active nest sites (n = 34) and at stratified random sites (n = 61). Logistic regression with information-theoretic model selection identified a model including greater tree species richness and closeness to forested wetland as the best-approximating model. Variable selection with Discriminant Function Analysis (DFA) indicated that nest selection was best explained by greater number of tree species, closer distance to forested wetlands, greater volume of downed woody debris, fewer small sawlogs, and increased proximity to streams. Univariate comparisons identified four of the five aforementioned variables in the DFA model as significant. Red-shouldered hawks are likely more common in Wisconsin than their state status suggests. Forest management for red-shouldered hawk nest sites should focus on increasing tree species richness, increasing down woody debris volume, and protecting forested wetlands. These recommendations may assist property managers to locate and plan for continued persistence of this species on MCF.  相似文献   

16.
This study was designed to answer questions about the patterns of understory diversity in managed forests of southern New England, and the factors that appear associated with those patterns. At the landscape-level, we used plot data to answer questions regarding the spatial distribution of forest understory plant species. Data from a combination of fixed area (understory vegetation) and variable radius (overstory trees) plot methods are combined with site variables for the analysis. Univariate and multivariate statistical methods are used to test for understory diversity relationships with overstory cover types and topography separately, and in combination. Analyses also test for relationships between specific understory species and cover types. In general the understory flora is dominated by four common clonal species that occur across the range of forest cover types: wild sarsaparilla (Aralia nudicaulis L.), Canada mayflower (Maianthemum candense Desf.), star flower (Trientalis borealis Raf.), and partridgeberry (Mitchella repens L.). Results also show that over story composition and structure can be used to assess understory species richness. Species richness follows a general trend among cover types of: hardwood ≥ regenerating forest, hardwood–pine, and pine ≥ mixed ≥ hardwood–hemlock > hemlock. Eastern hemlock (Tsuga canadensis L. Carriere) and mountain laurel (Kalmia latifolia L.) (which decreased in dominance from ridge to valley) both showed negative trends with understory species richness. Topographic position also appears associated with understory floristic patterns (particularly for the hardwood cover type), both in terms of species richness and compositional diversity which both increased from ridge, to midslope, to valley. However, overstory composition (covertype) appears to have a higher order influence on vegetation and mediates the role of topography. The results from this study provide foresters with a better understanding for maintaining floristic diversity and composition of the understory in managed forests.  相似文献   

17.
Old growth stands of boreonemoral spruce (Picea abies) forests frequently have a shrub layer dominated by hazel (Corylus avellana) – a species which is generally excluded in intensively managed forests due to clearcutting activities. We sampled understory species composition, richness and biomass, as well as environmental variables beneath these two species and also within forest ‘gaps’ in order to determine the effect of overstory species on understory vegetation. Species richness and biomass of herbaceous plants was significantly greater under Corylus compared with plots under Picea and in forest gaps. Indicator species analysis found that many species were significantly associated with Corylus. We found 45% of the total species found under woody plants occurred exclusively under Corylus. Light availability in spring and summer was higher in gaps than under forest cover but no difference was found between plots under Corylus and Picea. Hence, reductions in light availability cannot explain the differences in species composition. However, Ellenberg indicator values showed that more light demanding species were found under Corylus compared to Picea, but most light demanding species were found in gaps. The litter layer under Picea was three times thicker than under Corylus and this may be an important mechanism determining differences in understory composition and richness between the woody species. The presence of Corylus is an important factor enhancing local diversity and small-scale species variation within coniferous stands. Hence, management should maintain areas of Corylus shrubs to maintain understory species diversity in boreal forests.  相似文献   

18.
A high incidence of Diplodia shoot blight (site means ranging 85-100%) was observed on recently planted red pine (Pinus resinosa) seedlings where mature red pine stands previously had been clearcut. An investigation of the potential of harvest debris as a source of inoculum of Diplodia pathogens then was conducted. Cones, bark, needles, stems from shoots bearing needles, and stems from shoots not bearing needles (both suspended above the soil and in soil contact) were collected from harvest debris left at sites where clearcutting occurred. Conidia were quantified, and their germination rate was assessed, and Diplodia species were identified using PCR. Conidia of Diplodia species were found at all study sites and conidia counts increased from samples collected from 6 to 18 months after harvest. Germinable conidia were obtained from debris collected 6 months to 5 years after harvest. Fewer conidia were obtained from debris collected at intervals of up to 4-5 years after harvest and the percentage of germinable conidia was lower after longer intervals following harvest. More conidia were obtained and a greater percentage germinated from debris collected above the soil than from debris in soil contact. The host substrate also influenced the number of conidia and the percentage that germinated. Planting red pine seedlings next to debris infested with Diplodia pathogens could provide a persistent source of inoculum. Results should prompt further consideration by land managers and researchers of the potential forest health risks, in addition to benefits, that may be associated with harvest debris.  相似文献   

19.
云南松天然林及人工林群落结构和物种多样性比较   总被引：11，自引：0，他引：11       下载免费PDF全文
通过对云南省滇中地区云南松天然林和人工林的群落结构调查及物种多样性研究,结果表明:干扰较少的云南松天然林具有较为稳定、合理的群落结构,其乔木层优势种群具有较强的自然更新能力和长远的演替潜力,并且物种多样性和生境多样性。干扰较少的云南松天然林的这些特性均优于干扰较大的云南松天然林,干扰较大的云南松天然林又优于人工林。随着森林起源过程中干扰(人为干扰和自然干扰)程度的逐渐增加,云南松群落的稳定性、生物多样性、协调平衡能力及自然修复能力相应下降;而入侵种紫茎泽兰的分布范围和数量则相应上升,提示其或可作为森林群落生物多样性、干扰程度和森林健康评价的指示生物。  相似文献   

20.
China's subtropical forests have experienced severe deforestation and most native forests have been fragmented into an archipelago-like landscape. The genetic effect of habitat fragmentation depends largely on the level of gene flow within and among population fragments. In the present study, Eurycorymbus cavaleriei, a canopy tree found throughout subtropical China, was selected as a representative insect-pollinated plant species to assess genetic consequence of forest fragmentation. Contemporary pollen dispersal and mating patterns were estimated in two physically isolated stands of E. cavaleriei within fragmented forests using six highly polymorphic microsatellite loci. We found high genetic diversity (HE = 0.670–0.754) in both adults and offspring in the fragmented agricultural landscape, suggesting that habitat fragmentation did not necessarily erode genetic diversity of E. cavaleriei. Although substantial pollen travelled less than 100 m, paternity analysis revealed that a large amount of long-distance pollination events occurred, with the average pollen dispersal distance being 1107 m and 325 m for the two stands, respectively. Extensive pollen immigration (39.3–42.6%) indicated that there was effective genetic connectivity among E. cavaleriei stands in the fragmented forests. twogener analysis revealed that the exponential power model was the best-fitting dispersal curve with a fat-tailed (b < 1) dispersal feature. The results from a multilocus mating system analysis suggested that a small amount of biparental inbreeding and some correlated mating events occurred in the fragmented forests, which were similar to our parallel findings in the continuous forests of E. cavaleriei. Estimates of pollen pool structure (ΦFT = 0.128–0.174) indicated large genetic differences between pollen clouds accepted by maternal trees. The number of effective pollen donors (Nep) in E. cavaleriei, estimated using both mltr (Nep = 4.2–5.3) and twogener (Nep = 2.9–3.9) models, was equivalent to the number of effective pollen donors detected in continuous forests of E. cavaleriei. The pollen dispersal and mating patterns detected here indicated that habitat fragmentation did not have a negative impact on pollen movement in E. cavaleriei, possibly due to its generalist pollination system and the resilient foraging behavior of its pollinators in response to changes in landscape structure. The long distances of pollen-mediated gene flow between patches highlight the conservation value of remnant forest fragments in maintaining genetic connectivity at the landscape scale in subtropical China.  相似文献   

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