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1.
Liana-dominated forest patches constitute 15–20% of old-growth forests in the Eastern Amazon but are generally excluded from management for timber production. Here we ask if liana-dominated patches may be brought into production by clearing lianas and conducting enrichment planting (EP) of native timber species. We present growth results from 8 years of such EP trials. Rapid growth and low mortality of all species in this study suggest that EP in cleared liana patches can contribute to timber stocks in second and third harvests of managed forests. The most vigorous individuals of Parkiagigantocarpa and Schizolobium amazonicum in each enrichment site grew more than 1 cm diameter per year (rates were initially >2 cm yr−1), and attained dominant canopy positions and diameters equal to those of small canopy trees in the surrounding forest within 8 years of planting (mean dbh ∼18 cm and ∼20 cm, respectively, at year 8). Limited data on Ceiba pentandra plantings indicate a similar trajectory for this species (dbh ∼40 cm in 8 years). The most vigorous Swietenia macrophylla grew at least 1 cm per year in enrichment plots (mean dbh ∼10 cm in 8 years), but take longer to attain dominant positions. Tabebuia serratifolia may take much longer to reach the canopy than other species tested (rates <1 m yr−1). We attribute the excellent performance to light availability; planting in intact soil with minimal compaction and abundant organic material; and low competition rates maintained by periodic thinning of competing vegetation.  相似文献   

2.
To insure adequate regeneration and future timber yields of mahogany (Swietenia macrophylla King), many logged forests will have to be restocked through enrichment planting and managed using silvicultural techniques that maintain this species’ long-term survival and growth. This study compared the effects of planting method and two silvicultural treatments on the survival and growth of mahogany seedlings in logging gaps in Bolivia. We tested the hypotheses that survival and growth will be higher among transplanted seedlings than seedlings established from sown seeds and higher in silvicultural treatments that reduce competing vegetation and increase light. The first silvicultural treatment consisted of gaps logged 6 months prior to planting, gaps logged just prior to planting, and gaps treated with herbicide prior to planting. The second treatment, applied 12 months after planting, consisted of manual vegetation cleaning around mahogany seedlings in half of the gaps. The first hypothesis was supported in terms of initial seedling growth but not survival, which was similar between planting methods during the 12-92 months after planting. Transplanted seedlings grew significantly faster than those established from sown seeds during the first year, but this growth advantage disappeared by the second year. Although transplants were 84 cm taller than seed-sown seedlings by the end of the study, this height gain was probably not worth the cost of growing and transplanting seedlings. The second hypothesis was supported in terms of both survival and growth. A significantly greater proportion of seedlings survived in herbicide (62%) compared to 6-month-old (46%) and recent gaps (18%) and in cleaned (51%) versus control gaps (39%). Seedlings initially grew faster in herbicide and recent gaps than in 6-month-old gaps. These differences among silvicultural treatments were largely explained by canopy cover, which, throughout the study, was at least 14% lower in herbicide gaps and 9% lower in cleaned gaps relative to their respective alternatives. By 64 months growth diminished to near zero and no longer differed among gap treatments, despite lower canopy cover in herbicide gaps. By 92 months, saplings in herbicide gaps were only 145 and 77 cm taller than those in recent and 6-month-old gaps, respectively. To maximize survival and growth of mahogany seedlings in logging gaps while minimizing costs, silvicultural strategies should focus on direct seed sowing and appropriately timed interventions (i.e. manual cleaning) to control competing vegetation.  相似文献   

3.
Virgin beech Fagus orientalis forests in northern Iran provide a unique opportunity to study the disturbance regimes of forest ecosystems without human influence. The aim of this research was to describe characteristics of natural canopy gaps and gap area fraction as an environmental influence on the success of beech seedling establishment in mature beech stands. All canopy gaps and related forest parameters were measured within three 25 ha areas within the Gorazbon compartment of the University of Tehran’s Kheyrud Experimental Forest. An average of 3 gaps/ha occurred in the forest and gap sizes ranged from 19 to 1250 m2 in size. The most frequent (58%) canopy gaps were <200 m2. In total, canopy gaps covered 9.3% of the forest area. Gaps <400 m2 in size were irregular in shape, but larger gaps did not differ significantly in shape from a circle. Most gaps (41%) were formed by a single tree-fall event and beech made up 63% of gap makers and 93% of gap fillers. Frequency and diversity of tree seedlings were not significantly correlated with gap size. The minimum gap size that contained at least one beech gap-filling sapling (<1.3 m tall) was 23.7 m2. The median gap size containing at least one beech gap-filling sapling was 206 m2 and the maximum size was 1808 m2. The management implications from our study suggest that the creation of small and medium sized gaps in mixed beech forest should mimic natural disturbance regimes and provide suitable conditions for successful beech regeneration.  相似文献   

4.
Seeds of mahogany and Spanish cedar were planted in gaps created by harvesting railroad ties at two locations in the Yucatan Peninsula. Gaps averaged 40 m2 in area. Germination after 2 months was very low (2%) for Spanish cedar and moderately low (29%) for mahogany at the one location where it was measured, probably due to predation and fungal damage. Annual seedling mortality was 25–40%, and height growth was moderate to normal, with average height equal to 45 cm after 3 or 4 years, depending upon the location. We conclude that gaps from railroad tie harvesting are not adequate for the establishment of either species from seed. If, however, gaps created by harvesting could be concentrated into at least moderately open areas 400 m2 in area or larger, they may be useful as sites for regenerating both species.  相似文献   

5.
A high incidence of Diplodia shoot blight (site means ranging 85-100%) was observed on recently planted red pine (Pinus resinosa) seedlings where mature red pine stands previously had been clearcut. An investigation of the potential of harvest debris as a source of inoculum of Diplodia pathogens then was conducted. Cones, bark, needles, stems from shoots bearing needles, and stems from shoots not bearing needles (both suspended above the soil and in soil contact) were collected from harvest debris left at sites where clearcutting occurred. Conidia were quantified, and their germination rate was assessed, and Diplodia species were identified using PCR. Conidia of Diplodia species were found at all study sites and conidia counts increased from samples collected from 6 to 18 months after harvest. Germinable conidia were obtained from debris collected 6 months to 5 years after harvest. Fewer conidia were obtained from debris collected at intervals of up to 4-5 years after harvest and the percentage of germinable conidia was lower after longer intervals following harvest. More conidia were obtained and a greater percentage germinated from debris collected above the soil than from debris in soil contact. The host substrate also influenced the number of conidia and the percentage that germinated. Planting red pine seedlings next to debris infested with Diplodia pathogens could provide a persistent source of inoculum. Results should prompt further consideration by land managers and researchers of the potential forest health risks, in addition to benefits, that may be associated with harvest debris.  相似文献   

6.
Studies of regeneration in African rain forests suggest that without silvicultural treatments, natural succession in logging gaps may not result in the establishment of timber species. In this paper we present the results of an experimental enrichment planting with moabi (Baillonella toxisperma Pierre), a valuable and important timber species harvested in Central Africa. Although forest gaps are generally considered as favourable for the regeneration of this species, a survey conducted in a forest concession in south-eastern Cameroon provided an estimate of only 12.7 seedlings ha−1, suggesting that the species was, in fact, poorly represented in logging gaps within the study area. To further investigate the dynamics of the moabi in logging gaps, 795 seeds were sown in 15 logging gaps and 410 nursery-raised seedlings were planted in 15 other gaps. A biannual monitoring program over a 30-month period showed a lower survival rate for seedlings from sowing (75.9%) compared to that of nursery-raised seedlings (95.3%). Planted seedlings reached an average of 229.3 cm tall whereas seedlings from sowing were 167.5 cm tall, with the observed difference roughly corresponding to the average height of the nursery-raised seedlings at the time they were introduced to the logging gaps. After 30 months, the diameters of planted seedlings (16.8 mm) were also greater than those of the directly sown individuals (12.5 mm). Forest gap characteristics significantly influenced the growth of the plants. Factors accounting for the differences were total solar radiation, the soil content of coarse sand, the topographic position of the gap, the vegetation cover and the density of Macaranga spp. Whilst total solar radiation had a positive influence on growth, the remaining factors had impacted growth negatively. A streamlined technique was tested by planting 7 seedlings in 250 gaps. Without additional site maintenance, 29.3% of the moabi seedlings emerged naturally from the competing vegetation after 24 months. With biannual maintenance some 89.4% of seedlings became successfully established. Clearance operations had no significant influence on the height of plants whilst plant diameter was greater in cleared gaps. The total cost of the enrichment technique was 5.5 EUR per gap without maintenance and 7.5 EUR per gap with a single maintenance measure. Whilst long-term monitoring is needed, this study suggests a high survival rate of moabi introduced in logging gaps, and a growth rate 10 times higher than previously reported under canopy cover. These findings, combined with the low costs of the enrichment technique, support the use of silvicultural measures in logging gaps to restore the forest.  相似文献   

7.
雪灾干扰下林窗对木荷幼苗更新的影响   总被引:1,自引:1,他引:1       下载免费PDF全文
林窗干扰是维持森林生态系统的重要驱动力之一,对种子萌发、幼苗等自然更新过程、森林物种组成和动态、森林生物多样性的维持具有重要作用。本研究以2008年雪灾干扰后的浙江江郎山木荷林为研究对象,对木荷林窗大小结构、幼苗更新、生长等进行调查研究,结果表明:扩展林窗以50 100 m2的林窗个数最多(占总数的45.45%),各等级林窗中以50 100 m2的林窗占总面积比例最大(占总面积的30.31%)。林窗中木荷幼苗的平均高度和地径较对照林分分别高1.44 cm和0.61 mm,幼树在林窗中的平均高度和地径则比对照林分中分别高45.37 cm和5.00 mm且差异显著;林窗大小对木荷幼苗、幼树的高度和地径生长影响显著,中林窗中幼苗的高度和地径均高于小林窗和大林窗中的幼苗且差异均显著(F=4.893,P=0.007;F=5.203,P=0.004;n=357);幼树的地径在不同大小林窗中差异显著(F=3.569,P=0.037;n=43)。林窗幼苗的更新密度随着林窗面积的增大而增大,在林窗面积达到76 m2时,更新密度达到最大值,而后随着林窗面积的增大下降;中林窗和小林窗中更新苗木以低矮植株(1级、2级)为主,面积100 m2大林窗中,木荷幼苗生长较快。与他人研究的森林天然林窗相比,雪灾干扰后改变了林窗的大小分布结构和面积,50 100 m2的林窗比例较大,一定程度上更利于幼苗更新,具有相对较大的林窗幼苗更新密度;不论林窗大小,林窗内的更新幼苗都比林内多,郁闭度较大的林内或大面积的空地上都不利于更新幼苗的生长。因此,从受灾木荷林窗大小结构、幼苗更新、生长等来看,中林窗是幼苗适宜更新的面积,为木荷灾后恢复与重建提供了科学依据。  相似文献   

8.
Some Eucalyptus species are widely used as a plantation crop in tropical and subtropical regions. One reason for this is the diversity of end uses, but the main reason is the high level of wood production obtained from commercial plantings. With the advancement of biotechnology it will be possible to expand the geographical area in which eucalypts can be used as commercial plantation crops, especially in regions with current climatic restrictions. Despite the popularity of eucalypts and their increasing range, questions still exist, in both traditional planting areas and in the new regions: Can eucalypts invade areas of native vegetation, causing damage to natural ecosystems biodiversity?The objective of this study it was to assess whether eucalypts can invade native vegetation fragments in proximity to commercial stands, and what factors promote this invasive growth. Thus, three experiments were established in forest fragments located in three different regions of Brazil. Each experiment was composed of 40 plots (1 m2 each one), 20 plots located at the border between the forest fragment and eucalypts plantation, and 20 plots in the interior of the forest fragments. In each experimental site, the plots were paired by two soil exposure conditions, 10 plots in natural conditions and 10 plots with soil exposure (no plant and no litter). During the rainy season, 2 g of eucalypts seeds were sown in each plot, including Eucalyptus grandis or a hybrid of E. urophylla × E. grandis, the most common commercial eucalypt species planted in the three region. At 15, 30, 45, 90, 180, 270 and 360 days after sowing, we assessed the number of seedlings of eucalypts and the number of seedlings of native species resulting from natural regeneration. Fifteen days after sowing, the greatest number of eucalypts seedlings (37 m−2) was observed in the plots with lower luminosity and exposed soil. Also, for native species, it was observed that exposed soil improved natural germination reaching the highest number of 163 seedlings per square meter. Site and soil exposure were the factors that have the greatest influence on seed germination of both eucalypt and native species. However, 270 days after sowing, eucalypt seedlings were not observed at any of the three experimental sites. The result shows the inability of eucalypts to adapt to condition outside of their natural range. However, native species demonstrated their strong capacity for natural regeneration in forest fragments under the same conditions where eucalypts were seeded.  相似文献   

9.
South African coastal dune forests are young, highly disturbed subtropical communities where conventional models of forest dynamics may be challenged. We tested predictions from the gap-phase regeneration model by comparing seedlings of three common species representing contrasting regeneration strategies: Acacia karroo as a ruderal, Celtis africana as a coloniser of forest gaps, and Diospyros natalensis as a late-successional species. We grew seedlings under contrasting light and nitrogen levels in a greenhouse and in the field for 1 year to compare their growth and survival rates, allocation and photosynthetic traits. Species’ growth rates generally followed the expected order: Acacia > Celtis > Diospyros, but Acacia responded strongly to light and Celtis responded strongly to nitrogen, leading to cross-overs in growth rates. The plasticity of allocation and photosynthesis did not clearly differentiate the strategies, although it was greater in the light-demanding species. Acacia and Celtis tended to survive better in Acacia stands than in forest plots. Leaf-level light compensation points (LCPs) were similar for the three species in most conditions, but auxiliary data suggest Diospyros has a lower whole-plant LCP than Acacia. Growth rates and LCPs were lower than most of those reported for primary-forest species in the literature, suggesting an unusual degree of shade-tolerance in this habitat. We discuss reasons why variation in shade-tolerance may be less important here than in the prevailing model for forest regeneration and suggest other biotic factors that may help differentiate regeneration niches.  相似文献   

10.
11.
Wild pistachio (Pistacia atlantica Desf.) is the most economically important tree species in many rural areas in the west of Iran. The species produces resin used for a wide variety of traditional uses. Because the resin can be harvested non-destructively, the trees are maintained until mortality occurs from natural causes. The result is that natural, managed stands include a variety of age classes. In recent years, a lack of smaller size classes has been observed in the Qalajeh forest, which is located in the Zagros Mountain region of western Iran. We established a series of plots in an area typical of Qalajeh forest to characterize the diameter distribution of the wild pistachio component. We confirmed a deficit of stems <30 cm dbh, based in the expectation that the landscape-level diameter distribution should be characterized by a negative exponential curve. For trees ≥30 cm dbh, de Liocourt's equation closely fit the diameter distribution (r2 = 0.93), translating to a q-factor of 1.34. We used this curve to estimate the deficit number of stems in diameter classes <30 cm. We estimate that this forest should have 19–24 wild pistachio trees/ha in the 5–25 cm classes, as compared to about 5 trees/ha found currently. Based on local conditions, we recommend that at least 30 seedlings/ha should be planted to allow 6–8 trees to reach to the 5 cm class.  相似文献   

12.
Silvicultural canopy gaps are emerging as an alternative management tool to accelerate development of complex forest structure in young, even-aged forests of the Pacific Northwest. The effect of gap creation on available nitrogen (N) is of concern to managers because N is often a limiting nutrient in Pacific Northwest forests. We investigated patterns of N availability in the forest floor and upper mineral soil (0–10 cm) across 6–8-year-old silvicultural canopy gaps in three 50–70-year-old Douglas-fir forests spanning a wide range of soil N capital in the Coast Range and Cascade Mountains of western Oregon. We used extractable ammonium (NH4+) and nitrate (NO3) pools, net N mineralization and nitrification rates, and NH4+ and NO3 ion exchange resin (IER) concentrations to quantify N availability along north-south transects run through the centers of 0.4 and 0.1 ha gaps. In addition, we measured several factors known to influence N availability, including litterfall, moisture, temperature, and decomposition rates. In general, gap-forest differences in N availability were more pronounced in the mineral soil than in the forest floor. Mineral soil extractable NH4+ and NO3 pools, net N mineralization and nitrification rates, and NH4+ and NO3 IER concentrations were all significantly elevated in gaps relative to adjacent forest, and in several cases exhibited significantly greater spatial variability in gaps than forest. Nitrogen availability along the edges of gaps more often resembled levels in the adjacent forest than in gap centers. For the majority of response variables, there were no significant differences between northern and southern transect positions, nor between 0.4 and 0.1 ha gaps. Forest floor and mineral soil gravimetric percent moisture and temperature showed few differences along transects, while litterfall carbon (C) inputs and litterfall C:N ratios in gaps were significantly lower than in the adjacent forest. Reciprocal transfer incubations of mineral soil samples between gap and forest positions revealed that soil originating from gaps had greater net nitrification rates than forest samples, regardless of incubation environment. Overall, our results suggest that increased N availability in 6–8-year-old silvicultural gaps in young western Oregon forests may be due more to the quality and quantity of litterfall inputs resulting from early-seral species colonizing gaps than by changes in temperature and moisture conditions caused by gap creation.  相似文献   

13.
Nitrogen fertilizer inputs increased sharply over the last decade in Brazilian eucalypt plantations. Due to the economic and potential environmental cost of fertilizers, mixed plantations with N-fixing species might be an attractive option to improve the long-term soil N status. A randomized block design was set up in southern Brazil, including a replacement series and an additive series design, as well as a nitrogen fertilization treatment. The development of mono-specific stands of Eucalyptus grandis (0A:100E) and Acacia mangium (100A:0E) was compared with mixed plantations in proportions of 1:1 (50A:50E), and other stands with different densities of acacia for the same density of eucalypts. The objective was to assess the effect of inter-specific interactions on the early development of the two species. Aboveground biomass was measured 6, 12, 18 and 30 months after planting, sampling 6–10 trees of each species per treatment at each age, and allometric equations were established in 0A:100E, 100A:0E, 50A:50E and 50A:100E. The height and basal area of E. grandis seedlings were enhanced by 12% and 30%, respectively by N fertilization at age 1 year. Inter-specific competition led to a stratified canopy, with suppression in acacia growth earlier for basal area than for height. The mean number of stems per acacia tree at 36 months after planting was significantly higher in pure stands (3.7), than in 50A:50E (2.7) and in the additive series (between 1.6 and 1.8). H/D ratios were highly sensitive to inter-tree competition for the two species. The suppressed acacia understorey in mixed-species stands did not influence biomass production and partitioning within eucalypts. This pattern led to biomass accumulation combining the two species in 50A:100E that was about 10% higher than in 0A:100E, from age 12 months onwards. Aboveground net primary production (ANPP) amounted to 25 Mg ha−1 and 37 Mg ha−1 from age 18 to 30 months in 100A:0E and 0A:100E, respectively. Acacia ANPP in 50A:100E amounted to 2 Mg ha−1 over the same period, as a result of substantial inter-specific competition. An increment in biomass production in these very fast-growing eucalypt plantations was achieved introducing acacia as an understorey and not in the 50A:50E design, as observed in other studies.  相似文献   

14.
The sustainability of current harvest practices for high-value Meliaceae can be assessed by quantifying logging intensity and projecting growth and survival by post-logging populations over anticipated intervals between harvests. From 100%-area inventories of big-leaf mahogany (Swietenia macrophylla) covering 204 ha or more at eight logged and unlogged forest sites across southern Brazilian Amazonia, we report generally higher landscape-scale densities and smaller population-level mean diameters in eastern forests compared to western forests, where most commercial stocks survive. Density of trees ≥20 cm diameter varied by two orders of magnitude and peaked at 1.17 ha−1. Size class frequency distributions appeared unimodal at two high-density sites, but were essentially amodal or flat elsewhere; diameter increment patterns indicate that populations were multi- or all-aged. At two high-density sites, conventional logging removed 93–95% of commercial trees (≥45 cm diameter at the time of logging), illegally eliminated 31–47% of sub-merchantable trees, and targeted trees as small as 20 cm diameter. Projected recovery by commercial stems during 30 years after conventional logging represented 9.9–37.5% of initial densities and was highly dependent on initial logging intensity and size class frequency distributions of commercial trees. We simulated post-logging recovery over the same period at all sites according to the 2003 regulatory framework for mahogany in Brazil, which raised the minimum diameter cutting limit to 60 cm and requires retention during the first harvest of 20% of commercial-sized trees. Recovery during 30 years ranged from approximately 0 to 31% over 20% retention densities at seven of eight sites. At only one site where sub-merchantable trees dominated the population did the simulated density of harvestable stems after 30 years exceed initial commercial densities. These results indicate that 80% harvest intensity will not be sustainable over multiple cutting cycles for most populations without silvicultural interventions ensuring establishment and long-term growth of artificial regeneration to augment depleted natural stocks, including repeated tending of outplanted seedlings. Without improved harvest protocols for mahogany in Brazil as explored in this paper, future commercial supplies of this species as well as other high-value tropical timbers are endangered. Rapid changes in the timber industry and land-use in the Amazon are also significant challenges to sustainable management of mahogany.  相似文献   

15.
Tropical montane cloud forest has been undergoing a drastic reduction because of its widespread conversion to pastures. Once these forests have been cleared exotic grasses are deliberately introduced for forage production. Exotic grass species commonly form monodominant stands and produce more biomass than native grass species, resulting in the inhibition of secondary succession and tree regeneration. The purpose of this study was to assess the effect of native vs. exotic grass species on the early establishment of two native tree seedlings (Mexican alder, Alnus acuminata and Jalapa oak, Quercus xalapensis) on an abandoned farm in central Veracruz, Mexico. Seedling survival and growth were monitored (over 46 weeks) in relation to grass cover and height, and available photosynthetic active radiation (PAR). More seedlings survived in the presence of the native grass Panicum glutinosum than those growing with the exotic grass Cynodon plectostachyus (92% vs. 48%). The causes of seedling mortality varied between species; Q. xalapensis was affected by herbivory by voles but mainly in the exotic grass-dominated stands, whereas A. acuminata seedlings died due to competition with the exotic grass. A. acuminata seedlings increased more in height in the exotic grass-dominated stands (102 ± 7.8 cm) compared to native grass-dominated stands (51 ± 4.7 cm). Grass layer height, cover and available PAR were correlated (Pearson; p < 0.05). In the exotic grass dominated plots, grass layer height was correlated with the relative height growth rates of Q. xalapensis (Pearson; p < 0.05). These results indicate that the exotic grass may be affecting tree regeneration directly (grass competition) and indirectly (higher herbivory). Passive restoration may occur once P. glutinosum dominated pastures are abandoned. However, when C. plectostachyus dominates, introduction of early and mid successional tree seedlings protected against vole damage is needed.  相似文献   

16.
Tropical forests are characterized by diverse assemblages of plant and animal species compared to temperate forests. Corollary to this general rule is that most tree species, whether valued for timber or not, occur at low densities (<1 adult tree ha−1) or may be locally rare. In the Brazilian Amazon, many of the most highly valued timber species occur at extremely low densities yet are intensively harvested with little regard for impacts on population structures and dynamics. These include big-leaf mahogany (Swietenia macrophylla), ipê (Tabebuia serratifolia and Tabebuia impetiginosa), jatobá (Hymenaea courbaril), and freijó cinza (Cordia goeldiana). Brazilian forest regulations prohibit harvests of species that meet the legal definition of rare – fewer than three trees per 100 ha – but treat all species populations exceeding this density threshold equally. In this paper we simulate logging impacts on a group of timber species occurring at low densities that are widely distributed across eastern and southern Amazonia, based on field data collected at four research sites since 1997, asking: under current Brazilian forest legislation, what are the prospects for second harvests on 30-year cutting cycles given observed population structures, growth, and mortality rates? Ecologically ‘rare’ species constitute majorities in commercial species assemblages in all but one of the seven large-scale inventories we analyzed from sites spanning the Amazon (range 49–100% of total commercial species). Although densities of only six of 37 study species populations met the Brazilian legal definition of a rare species, timber stocks of five of the six timber species declined substantially at all sites between first and second harvests in simulations based on legally allowable harvest intensities. Reducing species-level harvest intensity by increasing minimum felling diameters or increasing seed tree retention levels improved prospects for second harvests of those populations with a relatively high proportion of submerchantable stems, but did not dramatically improve projections for populations with relatively flat diameter distributions. We argue that restrictions on logging very low-density timber tree populations, such as the current Brazilian standard, provide inadequate minimum protection for vulnerable species. Population declines, even if reduced-impact logging (RIL) is eventually adopted uniformly, can be anticipated for a large pool of high-value timber species unless harvest intensities are adapted to timber species population ecology, and silvicultural treatments are adopted to remedy poor natural stocking in logged stands.  相似文献   

17.
We examined the relative susceptibility of four mahogany species, Khaya ivorensis, Khaya anthotheca, Entandrophragma angolense, and E. utile, to Hypsipyla robusta attack. Seeds were obtained from one to three parent trees for each species. The research was conducted in the moist semideciduous forest zone in Ghana and used a randomized complete block design. Tree height and diameter and height to first branch were measured until 24 months after out-planting in the field. H. robusta damage was assessed by counting the numbers of shoots attacked, branches, and dead shoots. Khaya spp. grew better but experienced more attack than Entandrophragma spp. The relative susceptibility to H. robusta attack, from most to least, of the four species was: K. anthotheca > K. ivorensis > E. angolense > E. utile. At 24 months, the mean number of shoots attacked per tree ranged from 1.0 for an E. utile seed source to 3.6 on for a K. anthotheca seed source. At 15 months, K. anthotheca and K. ivorensis started branching at about 1.5 m, but height of clear trunk increased over time due to self-pruning. As K. anthotheca grew taller, the number of H. robusta attacks per tree declined. This suggested that selection of genotypes and species that are tolerant of H. robusta attack based on infestation of young plants may not be appropriate. Genetic factors more completely reflecting the response of different species and genotypes to H. robusta attack may manifest themselves at later growth stages.  相似文献   

18.
Berries and mushrooms are increasingly appreciated products of Finnish forests. Therefore, there is a need to integrate them in silvicultural planning. Bilberry (Vaccinium myrtillus L.) is an economically important wild berry that is widely collected for household consumption and sale in North Karelia, Finland. In this study, bilberry yield models developed recently were included in a stand growth simulator and the joint production of timber and bilberry was optimized by maximizing soil expectation value (SEV) with 3% discounting rate, assuming that 75% of the bilberry yield is harvested. The effect of bilberry production on the optimal stand management increased with increasing bilberry price. With high bilberry prices (4–8 € kg−1) it was optimal to manage the mixed stand of Scots pine, Norway spruce and birch, and the pure stand of Norway spruce so as to promote bilberry production. In the Scots pine stand, where bilberry yields are higher, bilberry production affected optimal stand management already with a price of 2 € kg−1. Compared to timber production, joint production led to longer rotation lengths, higher thinning intensities, more frequent thinnings, and higher share of Scots pine in the mixed stand. The contribution of bilberries to the total SEV increased with increasing bilberry price and discounting rate. In the mixed stand and pine stand the SEV of bilberry production, calculated with 3% discounting rate, exceeded the SEV of timber production when bilberry price was 4 € kg−1.With 4% discounting rate this happened already with bilberry price of 2 € kg−1. It was concluded that forest management which promotes bilberry yields is the most profitable in pine stands where the potential bilberry yields are high.  相似文献   

19.
Few long-term studies have been conducted on changes in soil nutrients after afforestation in Iceland, a country with a young history of forest management. While fertilization was shown to improve survival of seedlings in the first years after planting on the nutrient limited soils, knowledge about the nutrients status of the soils that develop under maturing forest stands is still scarce. In a chronosequence study, the development of base cations and Olsen-phosphorus (Olsen-P) in the mineral soil was followed in six forest stands of two different tree species of increasing age (14–97 years): native birch (Betula pubescens) and introduced Siberian larch (Larix sibirica). A treeless heathland was included to present soil conditions prior to forest establishment. The sites are part of the largest forest area in Iceland, located in the east of the country. Results revealed an effect of stand age on all soil nutrients investigated except for potassium (K). Olsen-P increased in 0–10 cm depth of the mineral soil, indicating a better availability and thus improved P supply in maturing forest stands. Calcium (Ca) and magnesium (Mg) concentrations decreased with stand age in 0–10 and 10–20 cm soil depth, while sodium (Na) decreased only in the upper soil layer. Only Olsen-P and K concentrations were higher in the upper soil layer as compared to 10–20 cm depth. This indicates a higher biotic control as opposed to the geochemical control of the other base cations.  相似文献   

20.
Cedrela odorata (Spanish cedar) is a neotropical broadleaf tree species that is in high demand for furniture and interior fittings. In 1998, seed collections were made from Spanish cedar in the Yucatan Peninsula, Mexico, for genetic conservation and tree improvement projects. Progeny from these collections were established in genetic trials at Bacalar, Noh Bec, and Zoh Laguna in the Yucatan. Survival at 2 years was 73.5% at Bacalar, 72% at Noh Bec, but only 20% after 6 months for several replicates at Zoh Laguna. Mean 2-year height, number of years of Hypsipyla attack on apical shoots (over 2 or 3 years) and total branching over 2 years, were 1.3 m, 0.04 years, and 2.5 branches at Bacalar, and 1.9 m, 0.35 years, and 1.6 branches at Noh Bec. Selection of the three best provenances (Escárcega, Bacalar, and Calakmul) would result in an estimated cross-site gain of 8.6% in 2-year height. Within-provenance, cross-site heritabilities for 2-year height, shoot borer attack, and branching were 0.1 ± 0.02, 0.0 ± 0.02, and 0.0 ± 0.02, respectively. Single-site heritabilities for height at 2 years, shoot borer attack, and branching were 0.09 ± 0.05, 0.02 ± 0.04, and 0.09 ± 0.05 at Bacalar and 0.16 ± 0.07, 0.0 ± 0.06, and 0.0 ± 0.07 at Noh Bec. Given the known advantage of rapid growth to avoid Hypsipyla damage over the long term and the stable performance of provenances across sites for growth traits, we recommend provenance selection over multiple trials to reduce the risk of serious damage by the insect. Good provenances and other suitable seed sources, as well as proper site selection and silvicultural practices, are important tools for increasing yields from plantations of C. odorata.  相似文献   

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