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1.
Midcanopy layers are essential structures in “old-growth” forests on the Olympic Peninsula. Little is known about which stand and tree factors influence the ability of midcanopy trees in young-growth forests to respond to release; however, this information is important to managers interested in accelerating development of late-successional structural characteristics. We examined basal area growth response of midcanopy trees following variable-density thinning in an effort to determine the effect of thinning and local environment on the release of western hemlock (Tsuga heterophylla (Raf.) Sarg.) and western redcedar (Thuja plicata ex. D. Don) on the Olympic Peninsula in western Washington. Release was measured as the difference between average annual basal area growth over the 5-year prior to thinning and the 3-to-6 year period following thinning. Results indicate that while growth rates were similar prior to thinning (5.4 cm2 year−1in both thinned and unthinned patches) midcanopy trees retained in a uniformly thinned matrix grew significantly more (8.0 cm2 year−1) than those in unthinned patches (5.4 cm2 year−1) for western hemlock and for western redcedar. Crown fullness and crown crowding affected the release of western hemlock in the thinned matrix. Initial tree size, relative age, local crowding and measures of crown size and vigor affected the release of western redcedar in the thinned matrix. Our results indicate that midcanopy western hemlock and western redcedar retain the ability to respond rapidly with increased growth when overstory competition is reduced and the magnitude of response is related to neighborhood variables (intracohort competition, overstory competition, and tree vigor), thus suggest that variable-density thinning can be an effective tool to create variability in the growth of midcanopy trees in young-growth stands. We expect that this rapid response will produce even greater variability over time.  相似文献   

2.
林分断面积生长模拟理论与技术研究   总被引:2,自引:0,他引:2  
林分断面积是众多测树因子中的核心因子。文中对林分断面积生长规律及其模拟技术研究进展进行了综述, 分析了自然生长林分与间伐林分断面积生长的异同, 并重点从生长方程的选取、断面积模型的分类研制、研制途径及间伐指标、林分自变量及密度指标以及单木竞争指标等5个方面探讨了林分断面积模拟研究进展, 指出林分断面积生长模拟理论与技术领域存在的问题及未来研究重点, 以期为相关研究提供借鉴与参考。  相似文献   

3.
We studied the effects of line thinning on stand structure, microclimate and understory species diversity of two Cryptomeria japonica D. Don plantations in south-central Japan. In each of two study sites we compared stand structure between the thinned stand and an adjacent unthinned stand and found that line thinning increased the growth rate of residual trees such that stand basal area may recover within 10 years after thinning. In the thinned stand, more open canopy conditions resulted in higher maximum temperatures on the forest floor during the early growing season than in the unthinned stand. The thinned stand had greater understory plant species richness and biomass than the unthinned stand. This study suggested that line thinning could potentially enhance biodiversity while simultaneously increasing tree-growth rates in overstocked Cryptomeria japonica plantations.  相似文献   

4.
This study considered the effects of thinning on the development of compression wood in stems of 35-year-old stand of Corsican pine (Pinus nigra L.). Part of the stand had been thinned at 5-yearly intervals and part left unthinned. Twenty trees each from the thinned and unthinned stands were randomly selected and felled. Measurements were made on tree height, stem diameter, stem slenderness and canopy depth. Wood samples were removed from the central part of the main log and cross-sectional measurements made on ring width, basic density and compression wood content. Cross-sectional area of compression wood was found to be three time higher in stems from the unthinned trees in comparison with those from the thinned trees. No significant differences in mean radial ring width or basic density were found between treatments. Correlations indicated that, with increasing in stem diameter, compression wood content increased in the unthinned trees, while a decline in compression was observed in the thinned trees. Tree height was also positively correlated with compression wood content in unthinned trees, while no equivalent relationship was observed in thinned trees. Observations from this study, while not conclusive, suggest that phototropic stimulus may be producing stem inclinations in the unthinned stand as trees compete for space in the canopy, whereas crown competition has been largely eliminated in the thinned stand; and that this is responsible for compression wood levels recorded in this study.  相似文献   

5.
Medhurst JL  Beadle CL 《Tree physiology》2001,21(12-13):989-999
The crown structure of Eucalyptus nitens (Deane & Maiden) Maiden 6 years after thinning, and the development of stand leaf area index both immediately and 6 years after thinning, were investigated. Thinning did not alter branch angle, branching density or the relationship between branch size and branch leaf area. However, larger branches were found in the lower crown of thinned trees and the increase in leaf area as a result of thinning occurred on the northern aspect of the crown. The vertical distribution of leaf area in unthinned trees was skewed toward the top of the crown and correlated with live crown ratio. The vertical distribution of leaf area in thinned trees tended to be less skewed and was unrelated to tree size or dominance. Leaf area index, as estimated from light interception measurements, increased at a constant rate soon after thinning regardless of residual stocking. In the longer term, residual stocking had a strong influence on leaf area increase per tree and was correlated with changes in crown length.  相似文献   

6.
Thinning treatments in second-growth forest may be a practical means of accelerating the development of certain old-growth structural features in regions where old stands are presently uncommon. We used CANOPY, an individual-tree model calibrated with data from thinned and unthinned stands, to simulate effects of thinning on growth rates and development of old-growth structural features in second-growth northern hardwoods. Three simulated, moderately heavy thinnings over a period of 45 years nearly doubled the predicted mean radial increment of canopy trees, percent of stand basal area in large trees, and area of canopy gaps. Compared to untreated stands, thinned stands had fewer dead trees per ha, but the dead trees were larger in size and the overall volume of snags and logs was little affected. In a 77-year old even-aged stand, moderately heavy thinning was predicted to reduce the time needed to attain the minimum structural features of an old-growth forest from 79 to 36 years. Simulated treatments in an older, uneven-aged stand gave mixed results; the moderately heavy treatment stimulated individual tree growth, but the removal of some medium-sized canopy trees in conjunction with natural mortality delayed the development of old-growth structure. Total volume of dead wood may still be deficient under the thinning regimes investigated in this study, but predicted live-tree structure 45 years after moderately heavy thinning was typical of stands in the advanced transition and steady-state stages of old-growth development. Results suggest that thinning can substantially accelerate the development of old-growth structure in pole and mature northern hardwoods, but response in older, uneven-aged stands is more modest, and treatments in these stands may need to be more conservative to achieve restoration goals.  相似文献   

7.
Development in height, diameter and basal area was studied for beeted and for originally planted Norway spruce (Picea abies (L.) Karst.) seedlings. The study was carried out as a survey in three unthinned stands in southern Sweden. Beeted seedlings were separated from those originally planted by differences observed in the number of annual rings at stump height. In all stands height and diameter were smaller and growth rates in height and basal area were lower for beeted than for originally planted seedlings. The lower growth rates for beeted seedlings may be explained as effects of inter‐tree competition. The oldest stand was thinned at the time of inventory. 40% of the thinned trees were beeted representing 18% of the thinned volume.  相似文献   

8.
Silvicultural practices that provide a wide variety of vegetative composition and structure (habitats) in young stands should help manage for biological diversity across forested landscapes. This study was designed to test the hypotheses that: (i) abundance and diversity of stand structure attributes (species diversity and structural diversity of herb, shrub and tree layers) and forest floor small mammal communities, and (ii) relative habitat use by large herbivores, will increase from unthinned to conventionally thinned to chemically thinned stands of young lodgepole pine (Pinus contorta) forest. Replicate study areas were located near Summerland, Kelowna and Williams Lake in south-central British Columbia, Canada. Each study area had three treatments: a conventionally thinned, a chemically thinned and an unthinned stand. Pre-commercial thinning was conducted in 1993. Coniferous stand structure and understory vegetation were measured prior to thinning in 1993 and 5 years later in 1998. Small mammal populations were sampled intensively from 1993 to 1998. Relative habitat use by large herbivores was sampled in 1998.

Our results indicate that chemical thinning of young lodgepole pine stands produced an aggregated pattern of crop trees compared with stands subjected to conventional thinning. Diameter growth of crop trees in the chemically thinned stands was similar to that in the conventionally thinned, but also to that in unthinned stands. Although horizontal stratification (aggregates of trees) was enhanced, vertical stratification (structural diversity of vegetation) was less in the chemically than conventionally thinned stands. Abundance and diversity of understory vegetation and small mammal communities were generally unaffected by stand thinning in these particular installations. Relative habitat use by mule deer (Odocoileus hemionus) occurred in a gradient from highest in the conventionally thinned stand to lowest in the unthinned stand. Habitat use by snowshoe hares (Lepus americanus) tended to have the opposite trend. Moose (Alces alces) exhibited no difference in habitat use among stands. Thus, although there were few differences among treatment stands, chemical thinning could be used to develop an aggregated pattern of crop trees in pre-commercially thinned stands to maintain habitat for herbivores such as snowshoe hares and mule deer. Understory plant and forest floor small mammal communities would be maintained in these stands as well.  相似文献   


9.
We studied the combined effects of thinning on stand structure, growth, and fire risk for a Scots pine thinning trial in northern Spain 4 years following treatment. The thinning treatments were: no thinning, heavy thinning (32–46% of basal area removed) and very heavy thinning (51–57% of basal area removed). Thinning was achieved via a combination of systematic and selective methods by removing every seventh row of trees and then by cutting suppressed and subdominant trees in the remaining rows (i.e., thinning from below). Four years after thinning, mean values and probability density distributions of stand structural indices showed that the heavier the thinning, the stronger the tendency towards random tree spatial positions. Height and diameter differentiation were initially low for these plantations and decreased after the 4-year period in both control and thinned plots. Mark variograms indicated low spatial autocorrelation in tree diameters at short distances. Diameter increment was significantly correlated with the inter-tree competition indices, and also with the mean directional stand structural index. Two mixed models were proposed for estimating diameter increment using a spatial index based on basal area of larger trees (BALMOD) in one model versus spatial competition index by Bella in the other model. As well, a model to estimate canopy bulk density (CBD) was developed, as this variable is important for fire risk assessment. Both heavy and very heavy thinning resulted in a decrease of crown fire risk over no thinning, because of the reduction in CBD. However, thinning had no effect on the height to crown base and thus on the flame length for torching. Overall, although thinning did not increase size differentiation between trees in the short term, the increase in diameter increment following thinning and the reduction of crown fire risks support the use of thinning. Also, thinning is a necessary first step towards converting Scots pine plantations to more natural mixed broadleaved woodlands. In particular, the very heavy thinning treatment could be considered a first step towards conversion of overstocked stands.  相似文献   

10.
杉木林分密度效应研究   总被引:20,自引:3,他引:20       下载免费PDF全文
该项试验共分2个部分.(1)造林密度试验,小区面积为600 m2,5个处理,即1 667株.hm-2(A)、3 333株.hm-2(B)、4~983株.hm-2(C)、6 633株.hm-2(D)、9967株.hm-2(E);(2)造林密度调控试验,造林设计方法与造林密度试验相同,但在林分生长过程中,按密度管理图的密管线0.5为标准进行间伐,间伐后保留密度要与临近的下一个初植密度较稀植的林分密度基本相同,两者进行比较.试验结果(1)造林密度试验,优势高、平均高、平均胸径均随年龄的增加而递增,随密度的增加而递减,优势高9a、平均高6 a、平均胸径5 a,密度间差异已达显著性水平;林分蓄积量则随年龄增加而递增,5~7 a,密度间差异显著,8~18a,只有A密度与E、D密度差异显著;枝下高随密度、年龄的增加而递增,12~a后,C、D、E密度间的差异很小;冠幅随密度的增加而递减,随年龄的增加而递增,9~10a后,各密度则随年龄增加而缓慢递减.(2)造林密度调控试验,同一指数级,间伐后的林分与其密度基本相同未间伐的林分比较,其优势高、平均高差异不明显;立木蓄积前者小于后者;总蓄积(立木蓄积+间伐蓄积)前者大于后者;同一指数级,初植密度不同的林分,间伐后与密度基本相同未间伐林分的蓄积百分比,初植密度大的大于初植密度小的;指数级不同,而初植密度相同的林分,间伐后与密度基本相同未间伐的林分总蓄积百分比,高指数级的比值大于低指数级的比值.  相似文献   

11.
Ponderosa pine (Pinus ponderosa Dougl. ex P. Laws) forest stand density has increased significantly over the last century (Covington et al. 1997). To understand the effect of increased intraspecific competition, tree size (height and diameter at breast height (DBH)) and leaf area to sapwood area ratio (A(L):A(S)) on water relations, we compared hydraulic conductance from soil to leaf (kl) and transpiration per unit leaf area (Q(L)) of ponderosa pine trees in an unthinned plot to trees in a thinned plot in the first and second years after thinning in a dense Arizona forest. We calculated kl and Q(L) based on whole- tree sap flux measured with heat dissipation sensors. Thinning increased tree predawn water potential within two weeks of treatment. Effects of thinning on kl and Q(L) depended on DBH, A(L):A(S) and drought severity. During severe drought in the first growing season after thinning, kl and Q(L) of trees with low A(L):A(S) (160-250 mm DBH; 9-11 m height) were lower in the thinned plot than the unthinned plot, suggesting a reduction in stomatal conductance (g(s)) or reduced sapwood specific conductivity (K(S)), or both, in response to thinning. In contrast kl and Q(L) were similar in the thinned plot and unthinned plot for trees with high A(L):A(S) (260-360 mm DBH; 13-16 m height). During non-drought periods, kl and Q(L) were greater in the thinned plot than in the unthinned plot for all but the largest trees. Contrary to previous studies of ponderosa pine, A(L):A(S) was positively correlated with tree height and DBH. Furthermore, kl and Q(L) showed a weak negative correlation with tree height and a strong negative correlation with A(S) and thus A(L):A(S) in both the thinned and unthinned plots, suggesting that trees with high A(L):A(S) had lower g(s). Our results highlight the important influence of stand competitive environment on tree-size-related variation in A(L):A(S) and the roles of A(L):A(S) and drought on whole-tree water relations in response to thinning.  相似文献   

12.
We studied the effects of two types of selective thinning on beech stands formed by a shelterwood cut in 1910 — with lower number of crop trees and higher thinning intensity (T1) and higher number of crop trees with lower thinning intensity (T2). The stands were thinned in 1980, 1991 and 2001. Despite a lower stand density after thinning, the annual basal area increments of thinned stands in both thinning periods (1980–1991 and 1991–2002) were around 20% higher compared to those of the control (unthinned) stands. The mean annual basal area increment of dominant trees was 30–56% larger in the thinned plots compared to the control plots. Of 176 initial crop trees in the T1, 72% were chosen again during the last thinning. In the T2, 258 crop trees were chosen in the first thinning, and only 62% of these trees were chosen again during the last thinning. Only crown suppression and diameter classes of crop trees significantly influenced their basal area increment when diameter classes, crown size, crown suppression, and social status were tested. In the thinned stands, the dominant trees are more uniformly distributed if compared to the dominant trees in the control plots. Finally, the herbaceous cover and the species diversity were higher in the thinned plots.  相似文献   

13.
Diameter, height and volume increment of Scots pine (Pinus sylvestrisL.) and Norway spruce (Picea abies (L.) Karst.) stands wereinvestigated in long-term thinning experiments in southern Finland.The measurement period was on average 19 years, and thinningsconsisted of five different treatments: (1) unthinned; (2) selectivethinning from below where thinning intensity was determinedaccording to the number of stems; (3) selective thinning frombelow where thinning intensity was determined according to standbasal area; (4) systematic thinning where corridors were openedat regular intervals in the stand; and (5) half-systematic thinningwhere corridors were opened at regular intervals and the remainingstrips thinned from below, i.e. a combination of selective andsystematic thinning. The selective (basal area) thinning ofScots pine resulted in a volume growth reduction of about 12per cent, but in Norway spruce it resulted in no marked reductioncompared with the unthinned plots. In both tree species, volumeincrement in the half-systematic and selective (basal area)thinning differed only slightly from each other. The systematicthinning resulted in the lowest volume increment. In the systematicthinning, the total yield (m3 ha–1) somewhat decreasedwith increasing corridor width. In half-systematic thinning,on the other hand, the total yield was not clearly related tocorridor width. A corridor accelerated the diameter incrementof the edge trees. The edge effect progressively diminishedwhen moving deeper into the strip and ended at a distance ofabout 3–4 m from the corridor edge. After the thinning,the height increment first decreased, but later on it recovered.The unthinned plots had more dead trees than the other treatments,but there were no significant differences between the othertreatments. It can be concluded that the prevailing thinningmethod in the Nordic countries, i.e. half-systematic thinning,most probably results in no significant growth and yield losses.  相似文献   

14.
We assessed the effects of thinning (0, 20 and 30 % extraction of basal area) and canopy type (pine–beech vs. pine plots, beech accounting for 12 % of total basal area) on radial growth of dominant and codominant Scots pine at inter-annual scale and on microclimatic conditions, radial growth and xylogenesis 9 years after thinning at intra-annual scale. Thinning weakly affected pine growth, which was enhanced 3 years after harvesting. Over time, a gradual reduction in pine growth in mixed canopy relative to pure canopy occurred only in unthinned plots apparently due to beech expansion. Indeed, 9 years after thinning, a higher seasonal radial increment and a greater number of tracheids were produced under pine canopy in the unthinned plots, whereas no differences between canopy types were observed in the thinned plots. Radial increment and tracheid production were mainly affected by tree water status (air and soil humidity, throughfall). The differences of tree water status caused by treatments, and plausibly disparities in tree size and tree-to-tree competition, were the main drivers explaining the patterns observed for radial increment and xylogenesis. Our results suggest that the negative effects of beech competition on Scots pine growth in similar mixed forest may be controlled to some extent by thinning.  相似文献   

15.
Forest thinning and prescribed fire practices are widely used, either separately or in combination, to address tree stocking, species composition, and wildland fire concerns in western US mixed conifer forests. We examined the effects of these fuel treatments alone and combined on dwarf mistletoe infection severity immediately after treatment and for the following 100 years. Thinning, burning, thin + burn, and control treatments were applied to 10 ha units; each treatment was replicated three times. Dwarf mistletoe was found in ponderosa pine and/or Douglas-fir in all units prior to treatment. Stand infection severity was low to moderate, and severely infected trees were the largest in the overstory. Thinning produced the greatest reductions in tree stocking and mistletoe severity. Burning reduced stocking somewhat less because spring burns were relatively cool with spotty fuel consumption and mortality. Burning effects on vegetation were enhanced when combined with thinning; thin + burn treatments also reduced mistletoe severity in all size classes. Stand growth simulations using the Forest Vegetation Simulator (FVS) showed a trend of reduced mistletoe spread and intensification over time for all active treatments. When thinned and unthinned treatments were compared, thinning reduced infected basal area and treatment effects were obvious, beginning in the second decade. The same was true with burned and unburned treatments. Treatment effects on infected tree density were similar to infected basal area; however, treatment effects diminished after 20 years, suggesting a re-treatment interval for dwarf mistletoe.  相似文献   

16.
Thinning of Korean pine (Pinus koraiensis Sieb. et Zucc.) is used to facilitate timber and cone production. The present study in Northeast China investigated the effects of thinning intensity on individual tree growth, temporal variation in cone yield, and seed quality in Korean pine plantation. In 2005, five thinning intensity levels (none, extreme, heavy, moderate and light) were set in 15 permanent plots in a 32-year-old Korean pine plantation at Mengjiagang Forest Farm, Jiamusi City, Heilongjiang Province. We recorded tree growth and seed cone production from 2013 to 2016, i.e., from 8 to 11 years after thinning. Except for height growth, thinning increased tree growth (diameter at breast height and crown size) and improved cone yield. The extreme thinning treatment (to 300 trees per hectare) resulted in the largest tree diameter, tree volume, crown size and 4-year cone production per tree. The highest cone yield per tree in the mast year (2014) was observed when stands were thinned to 500 trees per hectare (heavy thinning). Although the best cone and seed quality and the largest cone and seed mass per tree were recorded in the heavily thinned stand, no significant differences were found between heavy and moderate thinning stands (750 trees per hectare). At the stand level, the moderately thinned stand had the highest basal area, stock volume and seed cone production per stand. Our results suggest that thinning to 750 trees per hectare will improve timber and cone productivity in 40-year-old P. koraiensis stands.  相似文献   

17.
长白落叶松人工林间伐林分的生长模拟   总被引:8,自引:1,他引:7       下载免费PDF全文
利用黑龙江省孟家岗林场长白落叶松人工林间伐标准地定位观测数据,其于间伐林分断面积与相同年龄、相同立地、相同保留木株数的未间伐林分面积,随时间趋于一致的假设,根据未间伐人工林林分断面积生长的一般预测模型导出了间伐林分的断面积生长预测模型,同时,结合抚育间伐技术批示的确定,进行和林分的动态模拟。  相似文献   

18.
Commercial thinning enables forest managers to meet timber production objectives. Thinning reduces tree density to alleviate competition for resources and favour growth of selected tree species. However, in doing so, thinning can homogenize the composition of mixed-species forests and raise biodiversity issues. There is increasing evidence that species richness can lead to higher productivity through a complementarity effect. Hence, thinning that would maintain species diversity of mixed-species forests could enhance stand productivity and help forest managers to reconcile timber production objectives and biodiversity issues. The objective of this study was to compare post-thinning stand production, experimentally over 10 years, in mixed and monospecific stands of black spruce (Picea mariana [Mill.] B.S.P.) and jack pine (Pinus banksiana Lamb.). The post-thinning stand production curve of the mixed stand converged toward that of the unthinned mixed stand while the production curves of the thinned and unthinned monospecific stands remained parallel. The convergent productivity of the mixed stand could be explained by a positive interaction between effects of thinning and niche complementarity. We propose that thinning that maintains species diversity of mixed stands could help forest managers who are implementing ecosystem management to reconcile timber production objectives with biodiversity issues.  相似文献   

19.
Pruning allows knot-free timber to be obtained, thereby increasing the value of the highest-value wood products. However, the effect of pruning on growth is under discussion, and knowledge about the tree response to the simultaneous development of thinning and pruning is scarce. The objective of this study was to analyze the effect of the interaction of thinning and pruning on tree and stand level and the annual radial growth of two pine species native to Mediterranean mountains. We used long-term data of three trials installed in pine stands where several combinations of pruning and thinning were developed. Five inventories were carried out for each trial, and the mean dasometric features of the different treatments were compared using linear mixed models including a competition index. In addition, we collected cores from ten trees per plot in order to evaluate the annual response of trees to the thinning and pruning. We analyzed the annual radial growth using a semiparametric approach through a smooth penalized spline including rainfall and temperature covariates. Pruning did not show any effect on growth. However, larger diameter and increased annual radial growth were found in thinned plots, both with and without pruning, as compared to unthinned plots. Also, we found significant effects of climate on annual radial growth. We recommend the application of thinning and pruning in stands of Mediterranean mountains in order to get knot-free timber since growth reduction was not found in thinned stands.  相似文献   

20.
日本落叶松人工林密度调控技术研究   总被引:1,自引:0,他引:1  
以日本落叶松中幼林抚育间伐样地监测数据,利用树冠竞争因子(CCF)、林分密度指数、单位断面积生长过程等指标,分析了抚育间伐对断面积和蓄积两方面的影响,结果表明:间伐林分的断面积和蓄积有相同的生长过程,即林分在一定间伐强度范围内(强度小于50%)间伐强度越大,年均生长率越大;用树冠竞争因子描述日本落叶松人工林断面积的生长过程,未间伐林分的立木度接近稳定状态,而间伐林分降低了林分的CCF值,随着林龄的增加,CCF值不断增加,逐渐向未间伐林分靠近。总的来说间伐林分在间伐初期要小于未间伐林分,但随着林龄的增大及林木竞争的加剧,间伐林分逐渐向未间伐林分靠近。  相似文献   

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