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1.
A field study was conducted in the Western Ghats of peninsular India to test the following two hypotheses: (1) lower floristic diversity for the cardamom hill reserves (CHR —a traditional agroforestry system of the tropics) compared to undisturbed evergreen forests and (2) a truncated vegetation structure for shade trees in the cardamom areas. The experimental sites involved three CHRs and an evergreen forest site.The CHRs, regardless of their locations, were characterized by lower floristic diversity and density than the evergreen forest site. The undisturbed forest site at Ayyappancoil registered the highest floristic richness and diversity (Simpson's floristic diversity index, D=0.93), followed by the well-managed CHR site, suggesting that managerial interventions may have a strong bearing on the floristic diversity of CHRs. The current suite of species in the CHRs included both heliophilic as well as shade-tolerant components. However, dominant tree species, their density and relative abundance exhibited marked variations among the CHR sites, albeit about one-third of the species were common at all sites.Stand physiognomy was characterized by the dominance of a single layer of trees in the CHRs, while the wet evergreen forest exhibited a multilayered canopy structure. Some of the lower height classes were poorly represented in the cardamom areas, whereas the evergreen forests depicted an inverse J shaped height distribution pattern. Implicit in the truncated stand structure of the CHR is the poor regeneration status, due to systematic removal of the lower size classes. 相似文献
2.
Interactions between forest canopy characteristics and plants in the forest understory are important determinants of forest community structure and dynamics. In the highlands of southwestern, China the dwarf bamboo Bashania fangiana Yi is an understory dominant beneath a mixed canopy of the evergreen Abies faxoniana (Rheder & Wilson) and the deciduous Betula utilis (D. Don). The goal of this study was to better understand the role of bamboo dominance, canopy characteristics, and periodic bamboo dieback on forest development. To achieve this goal, we measured tree seedling, tree saplings, and trees, forest canopy characteristics, and bamboo cover in permanent forest (n = 4) and gap plots (n = 31) in a mixed A. faxoniana and B. utilis forest in Sichuan, China. Dwarf bamboos died off in 1983 in the gap plots, and in three of the four forest plots. Forest development was assessed for the period 1984–1996. The seedling bank in forest and gap plots increased after bamboo die-off. A. faxoniana seedlings increased more than B. utilis in forest plots; the opposite pattern characterized gap plots. The proportion of seedlings on raised micro-sites on the forest floor also changed and new seedling were more abundant on the forest floor. By 1996, bamboo seedling cover and biomass had recovered to ca. 45% or their pre-flowering values. Rates of bamboo seedling recovery were faster beneath canopy gaps and deciduous trees than beneath forest or evergreen trees. Tree mortality exceeded recruitment in plots with dense bamboo; the opposite pattern was found in the plot with little bamboo. The mortality rate for B. utilis trees (2.4% year−1) was higher than that for A. faxoniana (0.8% year−1) and forests with dense bamboos became more open over the census period. Tree mortality was size-dependent and intermediate sized trees had the lowest rates of mortality. Stand basal area increased mainly due to greater basal area gain than loss for A. faxoniana. Interactions between tree species life history, canopy type, and bamboo life-cycles create heterogeneous conditions that influence tree and bamboo regeneration and contribute to the coexistence of A. faxoniana and B. utilis in old-growth forests in southwestern China. 相似文献
3.
To better understand tree regeneration trajectories and the resultant coexistence of Abies with co-dominants, Picea jezoensis var. hondoensis, Tsuga diversifolia and Betula ermanii, in an old-growth subalpine forest, we investigated spatial mortality patterns during the regeneration of Abies mariesii and A. veitchii, which are abundant in the understory reflecting their shade tolerance. Regeneration of these Abies spp. from shaded understory to canopy status is affected by other canopy co-dominants. Snags of understory Abies spp. were common, suggesting that the primary mortality agent is suppression by the overstory. Although live, small Abies trees in the understory were positively associated with a Picea canopy, the long-term survival was reduced among Abies trees close to the canopy, suggesting that shading by large Picea in the overstory negatively affects understory Abies plants. The existence of shade-intolerant canopy co-dominants such as Picea and also Tsuga, which are larger and longer lived than the shade-tolerant Abies, may play an important role in preventing the Abies spp. from competitively displacing these other tree species, which are much rarer in the understory, though common in the canopy. Moreover, in spite of the fact that Betula canopies fostered recruitment and growth of Abies saplings, Abies showed no association with Betula canopy and their survival at later-stage was rather reduced near or beneath Betula canopies at the subsequent understory small tree stage. Based on spatially significant events related to tree death, this study detected such “habitat shifts” in the trajectory of tree regeneration. Accordingly, it can be concluded that careful consideration of the regeneration habitat is required for a fuller understanding of ecological processes in spatially complex old-growth forest systems. 相似文献
4.
Forests in the montane and lower subalpine ecoregions of the southern Canadian Rocky Mountains may have been more open and structurally diverse at the beginning of the 20th Century than today. Today, the area of mature Pinus contorta subsp. latifolia Dougl. forest that is conducive to mountain pine beetle (MPB, Dendroctonus ponderosae Hopkins) infestations, infrequent high severity fires, and herbivory appears to have increased in Banff and Kootenay National Parks. Based on a review of the literature, we hypothesised that MPB infestations increase forest stand structural diversity and tree species diversity. Stand structure parameters were investigated in mesic montane and lower subalpine stands approximately 15, 25, and 65 years after MPB outbreaks. Parameters measured were stand density (number of trees per ha), diameter at breast height, height class, species, age class distribution, and coarse woody debris mass. Influences of fire frequency, time since fire, and fire severity on these parameters were assessed to determine whether fire history had a confounding influence on stand structure. The Shannon–Wiener index indicated higher stand structural diversity 15 years but not 25 and 65 years after MPB infestations. MPB infestations led to general decreases within stands in the number of living trees, small diameter snags, and Pinus tree species and an increase within stands in the number of large diameter snags. Management that allows the occurrence of the natural fire regime of variable severity fires, in addition to some MPB infestations, would provide for more open and diverse stands. MPB infestations have some effects on stand structure that are similar to those of fire. Changes in stand structure resulting from recent declines in burning rates within Banff and Kootenay National Parks can be reversed to some extent by MPB infestations. 相似文献
5.
In this study three species of the genus Quercus section Erythrobalanus (Quercus coccinea, Quercus rubra, Quercus velutina) were investigated. All occur together as canopy trees in forests of southern New England. Acorns of each Quercus species were planted in plots located in five zones that represent a range of forest gap/canopy conditions that can occur within a southern New England forest. These five zones were demarcated adjacent to and across large openings of two physiographic sites—valley and ridgetop. Experiments were designed to monitor germination and initial growth of seedlings for the first three growing seasons. During the start of the first growing season germination was monitored. At the end of the first growing season measurements of height and number of flushes were taken and destructive samples of seedlings made for dry mass of root, stem and leaves. At the end of the third growing season height was recorded for surviving seedlings. Comparisons were made of germination and growth of seedlings located in the different gap/canopy conditions.
Results demonstrated clear differences in patterns of germination and early growth among species and among gap/canopy conditions of the sites. All species showed an increased lag in germination with reduced amounts of light. Highest growth and flushing rate were in the center conditions of the openings for all species during the first growing season. Quercus rubra had the greatest height growth the first growing season but a lower number of flushes than Q. velutina and Q. coccinea. In comparison with the other species, Q. rubra had the greatest total dry mass in most gap/canopy conditions after the first growing season. However, Q. velutina had the greatest total dry mass in the center of the ridgetop opening. In almost all gap/canopy conditions Q. velutina had greater proportions of dry mass allocated to roots compared with the other species.
After 3 years, greatest height growth in any of the gap/canopy conditions was recorded for all three species in the center of the valley site. Under this condition Q. rubra had significantly greater growth than Q. velutina and Q. coccinea. Quercus rubra also had significantly greater height growth and survival beneath the canopy conditions of the valley site than the other species. On the ridgetop site regeneration failed to establish beneath canopy conditions that provided low amounts of light. Quercus velutina showed greatest height growth after three years in the center and edge conditions of the ridgetop opening compared with the other species. Environmental influences that determine species germination and growth performance are suggested. 相似文献
6.
为探讨川西亚高山针叶林边界木树冠对林窗干扰的响应,对王朗自然保护区的这一植被类型进行了调查,分析了该类型森林中边界木的数量、树高、胸径和偏冠率的分布。结果表明：一些林窗的个别边界木个体较小,但大多数边界木个体与群落乔木层中主要树木的个体大小相一致,一个林窗中边界木的平均高度可以用来表示林窗的高度;约1／5的边界木未出现偏冠现象,绝大多数边界木的偏冠率在0．5～0．7之间(占61．72％),不同树种间的偏冠有显著差异;单个林窗的边界木最少为5株,最多为17株,平均每个林窗拥有9．51株边界木。 相似文献
7.
William F. Laurance Henrique E. M. Nascimento Susan G. Laurance Richard Condit Sammya D&#x;Angelo Ana Andrade 《Forest Ecology and Management》2004,190(2-3):131-143
We used data from a long-term (14–18 years) demographic study to infer the maximum longevity for populations of 93 relatively abundant tree species in central Amazonia. We also assessed the influence of several life-history features (wood density, growth form, mortality rate, recruitment rate, stem diameter, growth increment, population density) on tree longevity. Data on 3159 individual trees were collected in 24 permanent, 1 ha plots in undisturbed forest arrayed across a large (ca. 1000 km2) study area. For each species, three estimates of longevity were generated (by dividing the stem diameter of the largest tree by the median, upper quartile, and upper decile of observed diameter-growth rates), and the mean of these three values was used as a longevity estimate. Longevity values ranged from 48 years in the pioneer Pourouma bicolor (Cecropiaceae) to 981 years for the canopy tree Pouteria manaosensis (Sapotaceae), with an overall mean of 336±196 years. These growth-based estimates of maximum tree age were concordant with those derived from analyses of mean mortality rates. Tree longevity was positively correlated with wood density, maximum stem diameter, and population density, and negatively correlated with annual mortality, recruitment, and growth rates. On average, pioneer species had much lower longevity than did non-pioneers, whereas among old-growth trees, emergent species had greater longevity than did canopy species. Our results are consistent with radiocarbon-based studies that suggest that Amazonian trees can occasionally exceed 1000 years of age. 相似文献
8.
Yasuhiro Kubota 《Journal of Forest Research》2006,11(3):191-201
This article reports the regeneration dynamics of a temperate Abies–Tsuga forest in Kirishima Yaku National Park, southwestern Japan, and examines the influence of species coexistence mediated by
gap disturbances on biomass production. All trees taller than 2 m in a 1-ha plot were monitored over four growing seasons.
Three growth-form groups occupied different vertical layers. Evergreen conifers and deciduous broad-leaved trees tended to
be spatially segregated from evergreen broad-leaved trees, which formed thickets in the understorey. The regeneration of understorey
evergreen broad-leaved trees was affected by canopy gaps. The recruitment of conifers and deciduous broad-leaved species was
not observed during the four growing seasons. This suggests that regeneration is sporadic and the present environmental conditions
are not favorable for these canopy species. The mortality and unsuccessful recruitment of conifers and deciduous trees appeared
to cause fluctuations in the productivity of the stand. However, an abundance of canopy gaps accelerates the regrowth of shorter
species, and the fluctuation of productivity resulting from the population dynamics of canopy species would be partly mitigated
by the regeneration of evergreen understorey species. The horizontal and vertical heterogeneity of the temperate mixed forest
was a result of the patch structures of the three growth-form groups. The different regeneration patterns among the three
groups, which were driven by interactions of species-specific regeneration niches and disturbance regimes, might be an important
factor in maintaining the aboveground productivity in a transitional mixed forest between warm-temperate and cool-temperate
zones. 相似文献
9.
Subtropical forests in the Ryukyu Islands have been degraded by silvicultural practices, and thus their structural attributes
are being shifted to other states dominated by a few tree species. This study clarified the mechanisms of the change, and
examined the effect of clear logging on the resilience of a subtropical forest. Sprouting regeneration and typhoon disturbance
were introduced into an individual-based model, SEIB-DGVM, for describing stand development and succession. The regeneration
dynamics from young secondary to old-growth stands were reproduced fairly well with the model. Sprouting recruitment produced
high stem density at the beginning of stand development, which caused a self-thinning trajectory following the −3/2 power
law. In the late development stage after 70 years, tree species diversity fluctuated because of the regenerative response
of sprouting species and the facilitatory effect of typhoon disturbance on the coexistence of subordinate species. The death
of canopy trees because of typhoon disturbances reduced the dominance of Castanopsis sieboldii, and depressed its dominance in the understory. Consequently, the understory species could establish by virtue of fallen
canopy trees, and tree species diversity increased at the stand level. Clear logging experiments in the model revealed that
species diversity deteriorated, especially in the stand dominated by sprouting species. Resilience of subtropical forests
was determined by initial species composition before clear logging. Our simulation results suggest that repeated logging drives
subtropical forests with high species diversity to a stand monopolized by C. sieboldii. 相似文献
10.
Mycorrhizal associations of important tree species were investigated within the research area of the Tropenbos Cameroon Programme (TCP), situated on the western portion of the Atlantic Biafrean forest of south Cameroon. Ninety-seven tree species of economic, social, and ecological importance, and three lianas were selected in three sites that differed in altitude, soil clay content, and soil pH. In each site plots were laid out in undisturbed forest. In each plot, seedlings, saplings, juvenile, and mature trees were identified to species level and counted; girth at breast height measured and basal area calculated; root samples were taken and examined for mycorrhizal type and extent of mycorrhizal colonization.
All 100 species investigated were mycorrhizal. Tree of 74 species formed exclusively arbuscular mycorrhizas (AM); 23 trees and three Gnetum species formed ectomycorrhizas (ECM). Five of these ECM plants also harbored AM structures. Extent of mycorrhizal root colonization showed large differences for various AM trees; however, colonization of more than half of these trees was less than 25%. Colonization of ECM trees was often higher than 75%. The contribution of ECM trees to basal area varied between 19 and 35%. ECM trees often occurred in small to large clumps. In sustainable forest management plans, existing ECM forest clumps should be given special conservation value. 相似文献
11.
亚热带常绿阔叶林结构、树种多样性和土壤特性的研究(英文) 总被引：1，自引：0，他引：1
采用样地调查法,对日本冲绳岛北部的亚热带常绿阔叶林的林分结构,树种组成以及土壤特性等进行了调查分析。结果表明,该亚热带林具有林冠低矮、小径木多等特点。林分的平均林冠高度仅10m,平均立木密度达5400株·hm2(≥3.0cmDBH),其中胸径小于10cm的立木占64%。林分胸高断面积为54.4m2·hm-2。30个调查样地共出现树种数(≥3.0cmDBH)达80种,分属31个科。根据树种的重要值,Castanopsis sieboldii和Schima wallichii分别为优势树种和亚优势树种。树种多样性指数(H’)平均达3.63,均衡度指数(J’)为0.71,丰富度指数(S’)为4.72。这些多样性指数都随着优势树种重要值的增加而显著降低。土壤养分测定结果表明,该森林的土壤肥力低,异质性大,并可能存在Al毒害。由相关分析知,林分密度和优势树高均与土壤pH值呈显著正相关关系;多样性指数则与土壤交换性K ,Ca2 以及Ca2 /Al3 比呈显著的正相关,而与土壤有机碳,全氮和磷呈显著的负相关。这些结果说明土壤性质是影响冲绳亚热带常绿阔叶林林分结构和树种组成多样性的重要因素。图3表4参38。 相似文献
12.
Impact of bark stripping by sika deer, Cervus nippon, on subalpine coniferous forests in central Japan 总被引：7，自引：0，他引：7
Shotaro Yokoyama Ikuyo Maeji Tsuyoshi Ueda Masaki Ando Ei&#x;ichi Shibata 《Forest Ecology and Management》2001,140(2-3):93-99
We investigated the effect of bark stripping by sika deer, Cervus nippon, on forest regeneration in subalpine coniferous forests on Mt. Ohdaigahara and in the Ohmine Mountains of central Japan. Bark stripping by sika deer occurred in five major tree species: Abies homolepis; Abies Veitchii; Tsuga diversifolia; Picea jezoensis var. hondoensis; and Chamaecyparis obtusa. The percentage of damaged trees on Mt. Ohdaigahara was higher than in the Ohmine Mountains, probably because of the higher deer density. On Mt. Ohdaigahara, the DBH distributions of stems for P. jezoensis var. hondoensis, A. homolepis, T. diversifolia andC. obtusa were bell-shaped with fewer smaller and larger trees. On the other hand, in the Ohmine Mountains the distributions for P. jezoensis var. hondoensis and A. Veitchii showed a reverse-J shaped with more smaller trees. Larger overstory conifers on Mt. Ohdaigahara were killed by bark stripping when 100% barked, although in the Ohmine Mountains ca. 50% of the trees survived even when 100% barked. After the disappearance of the overstory conifers on Mt. Ohdaigahara, the dwarf bamboo, Sasa nipponica, expanded into the forest floor due to changes in light reaching the forest floor. Since S. nipponica is the main forage of deer in this area, this increase caused a corresponding increase in the deer population, which in turn, could cause a further decline in the coniferous forests. 相似文献
13.
亚高山3种森林群落的结构特征与物种多样性 总被引：1，自引：0，他引：1
本文采用常规的群落学调查方法,研究了亚高山林区次生白桦（BF）、岷江冷杉（FF）和紫果云杉林（SF）的物种组成、叶面积指数（LAI）、年龄结构、胸径（DBH）分布和多样性指数等群落综合特征。结果表明：（1）BF群落中出现的雏管束植物有27科57属71种,FF群落中出现的维管柬植物有28科50属60种,SF群落中出现的维管束植物有27科36属39种;（2）BF、FF、SF群落的LAI分别为0．84（±0．11）、1．25（±0．30）和3．53（±0．22）;（3）FF、SF群落内的岷江冷杉、紫果云杉和方枝柏的树高、冠幅与胸径大小之间有显著的线性关系（R^2〉0．84）,BF群落内的白桦树高、冠幅与胸径之间没有显著的线性关系（R^2〈0．10）。处于演替早期的森林群落物种多样性高于演替后期的物种多样性,演替后期的群落LAI高于演替初期的群落圳。可利用胸径大小与树高、冠幅之间的显著线性关系来评估亚高山“针叶树种”的冠幅和树高。 相似文献
14.
Changes with altitude of the stand structure of temperate forests on Mount Norikura,central Japan 总被引：1，自引：0，他引：1
Changes in stand structure with altitude have been studied at ten sites between 800 and 2,500 m above sea level (a.s.l.) in
temperate forests on Mount Norikura (36°06′N, 137°33′E, 3,026 m a.s.l.) in central Japan. Vegetation in this range of altitudes
was roughly classified as a montane deciduous broad-leaved forest zone between 800 and 1,600 m a.s.l. and a subalpine coniferous
forest zone between 1,600 and 2,500 m a.s.l. at the timberline. The frequency distribution of trunk height was an L-shaped
pattern at 800 m a.s.l., changed to a flat pattern with increasing altitude up to 2,000 m a.s.l., but changed to an L-shaped
pattern again from 2,000 to 2,500 m a.s.l. This change in frequency distribution of trunk height with altitude was related
to the change in maximum trunk height with altitude. The maximum trunk height did not change with altitude between 800 and
2,000 m a.s.l. but it decreased markedly from 2,000 to 2,500 m a.s.l. Although mechanical damage to conifer trunks and branches
was not observed between 800 and 2,000 m a.s.l., the proportion of damaged trees increased from 2,000 to 2,500 m a.s.l., suggesting
that subalpine conifers cannot grow in height near the timberline because of mechanical damage. The increase in the number
of small trees from 2,000 m a.s.l. to the timberline was therefore because of a less developed canopy structure, i.e. small
trees can grow without shading by canopy trees. This study suggests that mechanical damage, probably because of strong winds
in the winter, affects changes with altitude in regeneration and formation of the timberline. 相似文献
15.
Competition for canopy space is a fundamental structuring feature of forest ecosystems and remains an enduring focus of research attention. We used a spatial neighborhood approach to quantify the influence of local competition on the size of individual tree crowns in north-central British Columbia, where forests are dominated by subalpine fir (Abies lasiocarpa), lodgepole pine (Pinus contorta) and interior spruce (Picea glauca × engelmanii). Using maximum likelihood methods, we quantified crown radius and length as functions of tree size and competition, estimated by the species identity and spatial arrangement of neighboring trees. Tree crown size depended on tree bole size in all species. Given low levels of competition, pine displayed the widest, shortest tree crowns compared to the relatively long and narrow crowns found in spruce and fir. Sensitivity to crowding by neighbors declined with increasing tree height in all but the pine crown radius model. Five of the six selected best models included separate competition coefficients for each neighboring tree species, evidence that species generally differ in their competitive effects on neighboring tree crowns. The selected crown radius model for lodgepole pine, a shade-intolerant species, treated all neighbors as equivalent competitors. In all species, competition from neighbors exerted an important influence on crown size. Per-capita effects of competition across different sizes and species of neighbors and target trees varied, but subalpine fir generally displayed the strongest competitive effects on neighbors. Results from this study provide evidence that species differ both in their response to competition and in their competitive influence on neighbors, factors that may contribute to maintaining coexistence. 相似文献
16.
We experimentally investigated interacting effects of canopy gaps, understory vegetation and leaf litter on recruitment and mortality of tree seedlings at the community level in a 20-year-old lowland forest in Costa Rica, and tested several predictions based on results of previous studies. We predicted that experimental canopy gaps would greatly enhance tree seedling recruitment, and that leaf litter removal would further enhance recruitment of small-seeded, shade-intolerant seedlings in gaps. We created a large (320–540 m2) gap in the center of 5 out of 10 40 m × 40 m experimental plots, and applied the following treatments bimonthly over a 14-month-period in a factorial, split–split plot design: clipping of understory vegetation (cut, uncut), and leaf litter manipulations (removal, addition, control). As expected, experimental gaps dramatically increased tree seedling recruitment, but gap effects varied among litter treatments. Litter addition reduced recruitment in gaps, but enhanced recruitment under intact canopy. Species composition of recruits also differed markedly between gap treatments: several small-seeded pioneer and long-lived pioneer species recruited almost exclusively in gaps. In contrast, a few medium-to-large-seeded shade-tolerant species recruited predominantly under intact canopy. Leaf litter represents a major barrier for seedling emergence and establishment of small-seeded, shade-intolerant species, but enhances emergence and establishment of large-seeded, shade-tolerant species, possibly through increased humidity and reduced detection by predators. Periodic clipping of the understory vegetation marginally reduced tree seedling mortality, but only in experimental gaps, where understory vegetation cover was greatly enhanced compared to intact canopy conditions. Successful regeneration of commercially valuable long-lived pioneer trees that dominate the forest canopy may require clear-cutting, as well as weeding and site preparation (litter removal) treatments in felling clearings. Management systems that mimic natural canopy gaps (reduced-impact selective logging) could favor the regeneration of shade-tolerant tree species, potentially accelerating convergence to old-growth forest composition. In contrast, systems that produce large canopy openings (clear-cutting) may re-initiate succession, potentially leading to less diverse but perhaps more easily managed “natural plantations” of long-lived pioneer tree species. 相似文献
17.
Composition, structure, and species-specific patterns of recruitment and growth were characterized in two yellow birch (Betula alleghaniensis Britt.)–conifer stands in Quebec, Canada, to improve our understanding of the dynamics of these complex ecosystems. The mixture of mid- and shade-tolerant species in the canopy, the inverse J-shape stem diameter distribution, and the age distribution were indicative that the two stands were in a late-successional stage. Recruitment of mid-tolerant species above 1.3 m in height appeared to be periodic and synchronized with historical spruce budworm (Choristoneura fumiferana Clem.) outbreaks, while the coniferous component of these mixedwood stands recruited continuously. Results suggest that recruitment of yellow birch and red maple (Acer rubrum L.) requires disturbances of a certain intensity that affect at least 25 % of the forest cover. In contrast, balsam fir (Abies balsamea (L.) Mill.) and red spruce (Picea rubens Sarg.) can recruit under the canopy without relying on moderate or large canopy disturbances. Results suggest that the historical disturbance regime, and differences in shade tolerance between species, largely govern the contemporary composition of these stands. This study improves the comprehension of mechanisms that regulate the dynamics of yellow birch-conifer stands and will be useful for the subsequent elaboration of forest management strategies. 相似文献
18.
King DA 《Tree physiology》1991,9(3):369-381
Relationships between tree height and crown dimensions and trunk diameter were determined for shade-tolerant species of old-growth forests of western Oregon. The study included both understory and overstory species, deciduous and evergreen angiosperms and evergreen conifers. A comparison of adult understory species with sapling overstory species of similar height showed greater crown width and trunk diameter in the former, whether the comparison is made among conifers or deciduous trees. Conifer saplings had wider crowns than deciduous saplings, but the crown widths of the two groups converged with increase in tree height. Conifer saplings had thicker trunks than deciduous saplings of similar crown width, possibly because of selection for resistance to stem bending under snow loads. The results suggest that understory species have morphologies that increase light interception and persistence in the understory, whereas overstory species allocate their biomass for efficient height growth, thereby attaining the high-light environment of the canopy. The greater crown widths and the additional strength requirements imposed by snow loads on conifer saplings result in less height growth per biomass increment in conifer saplings than in deciduous saplings. However, the convergence in crown width of the two groups at heights greater than 20 m, and the proportionately smaller effect of snow loads on large trees, may result in older conifers equalling or surpassing deciduous trees in biomass allocation to height growth. 相似文献
19.
Jian Rang Wang Tony Letchford Phil Comeau J. P. Kimmins 《Forest Ecology and Management》2000,130(1-3):17-26
Stand structure, height and diameter growth, above- and below-ground biomass, and nutrient concentrations and content were determined for a 35-year-old fire-origin paper birch (Betula papyrifera) and subalpine fir (Abies lasiocarpa) mixed-species stand in the Sub-Boreal Spruce (SBS) zone of British Columbia. Paper birch, which formed the dominant overstory following the 1961 fire, had normal distributions of height and diameter classes. Subalpine fir, which dominated the understory, had the reverse J-shaped height and diameter distributions that are expected of a shade tolerant, climax species. Paper birch grew more than three times the height of subalpine fir. Growing in the summer shade of the birch, subalpine fir had slow but steady height growth during the first 10–15 years, after which height growth declined somewhat. Allometric equations, relating dry weight of foliage, branches, stemwood, stembark, roots, and total biomass to diameter at breast height (DBH), were developed to estimate above- and below-ground biomass. Total biomass of paper birch reached 83.2 t ha−1, while subalpine fir biomass was 26.7 t ha−1. Subalpine fir allocated more biomass to foliage and branches compared to paper birch. Foliage of paper birch had higher nutrient concentrations of N, P, K, Ca, and Mg than subalpine fir foliage. Branches and stembark of subalpine fir had higher P, and Ca concentrations than paper birch. Subalpine fir branches contained more of all examined nutrients than paper birch branches. This is a significant component in nutrient cycling of the mixed-species forest. 相似文献
20.
Many forest plantations in the humid neo-tropics are established on degraded soils in abandoned pasture land and, with some exceptions, the species planted have not grown successfully. Studies of adaptability and growth under these conditions are scarce, particularly for native species. In this paper we present data on growth and tree form at 3 years of age for 11 species planted in abandoned pastures. The research plantation was established at La Selva Biological Station, in the Atlantic lowlands of Costa Rica (10° 26′N, 83° 59′W). Survival, diameter at breast height (DBH), total height, basal area, volume index and tree form were used to evaluate the species performance. After 3 years, Vochysia guatemalensis displayed a survival significantly higher (98%) than that of the other species. The lowest survival was 75% for Pinus tecunumanii. There were significant differences in growth measures among species. Acacia mangium exhibited significantly higher growth rates than the other species. Although this species grew rapidly, it did not develop straight single stems. About 14% of trees of A. mangium had bifurcation below DBH and more than 50% showed multiple axes or branches as large in diameter as the principal axis. The native species with the highest growth rate was Vochysia guatemalensis (DBH 12.7 cm, total height 7.5 m, volume index 55 m3 ha−1 at 3 years of age). Vochysia ferruginea exhibited a slightly lower growth rate. Both Vochysia species were ranked highly with regard to tree form, with more than 80% straight single stemmed trees. The lowest growth rates were exhibited by Pentaclethra macroloba, Pithecellobium macradenium and Virola koschnyi; however, the latter two species formed straight single terms. Pentaclethra macroloba and Inga edulis failed to form straight single stems. In general, the growth rates of the species studied were high in comparison with results reported from other tree plantations in the tropics. In spite of this, it is not prudent to draw final conclusions on the basis of these 3 year measurements. 相似文献