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1.
Between 1987 and 2000, a spruce beetle (Dendroctonus rufipennis) outbreak infested 1.19 million ha of spruce (Picea spp.) forests in Alaska, killing most of the large diameter trees. We evaluated whether these forests would recover to their pre-outbreak density, and determined the site conditions on which spruce germinated and survived following the spruce beetle outbreak in forests of the Anchor River watershed, Kenai Peninsula, Alaska. White spruce (Picea glauca) and Lutz's spruce (Picea × lutzii), a hybrid between white and Sitka spruce (Picea sitchensis), dominate the study area. We measured the pre- and post-outbreak density of spruce in 108 3 m × 80 m plots across the study area by recording all live trees and all dead trees >1.5 m tall in each plot. To determine the fine scale site conditions on which spruce germinated and survived, we measured ground surface and substrate characteristics within 20 cm circular plots around a subset of post-outbreak spruce seedlings. The density of post-outbreak spruce (855/ha) was adequate to restock the stands to their pre-outbreak densities (643/ha) for trees >1.5 m tall. We could not accurately estimate recovery for pre-outbreak spruce seedlings because dead seedlings may have decayed in the 5–18 years since the beetle outbreak occurred. At the fine scale, spruce that germinated post-outbreak grew on a wide variety of substrates including downed log, stump, mesic organic mat, peat, hummocks and mineral soil. They exhibited a strong preference for downed logs (53%) and stumps (4%), and most (91%) of the downed logs and stumps that spruce rooted on were heavily decayed. This preference for heavily decayed logs and stumps was especially evident given that their combined mean cover was only 2% in the 3 m × 80 m plots. Within the 3 m × 80 m plots, spruce seedling survival was negatively correlated with bluejoint (Calamagrostis canadensis) litter cover.  相似文献   

2.
The nutrient status of Norway spruce in pure and in mixed-species stands   总被引：1，自引：0，他引：1  
Atmospheric deposition of N and S appears to have caused nutrient imbalance in Norway spruce stands in southern Sweden. This calls for a change of forest management to procedures that promote nutrient balance. Studies have shown lower soil acidity in Norway spruce/deciduous mixed stands than in spruce monocultures, but the tree nutrient status in such mixtures has not been much investigated so far.

The nutrient status of Norway spruce foliage and top mineral soil chemistry in monocultures and in stands mixed with beech, birch, or oak was investigated through paired comparisons on 30 sites in southern Sweden (27 sites) and eastern Denmark (three sites). In total, 45 mixed stands and 34 pure stands were included in the study.

Spruce needles from mixed stands had higher concentrations and ratios to N of K, P, and Zn than needles from pure spruce stands. Among the mixed stands, the K status appeared to be positively correlated with the percentage of deciduous tree basal area. Soil samples from mixed stands had a higher Mg concentration, base saturation, and BC/Al ratio than soil samples from pure stands. The spruce needle nutrient status was comparable in pure stands on fertile sites and in mixed stands on poor sites. We did not detect any differences in spruce tree growth between pure and mixed stands.

This paper discusses possible reasons for a positive effect on the tree nutrient status in mixed-species stands and the possibility of using mixed-species stands as a forest management procedure to avoid nutrient imbalance.  相似文献   


3.
Soil properties were compared in adjacent 50-year-old Norway spruce, Scots pine and silver birch stands growing on similar soils in south-west Sweden. The effects of tree species were most apparent in the humus layer and decreased with soil depth. At 20-30 cm depth in the mineral soil, species differences in soil properties were small and mostly not significant. Soil C, N, K, Ca, Mg, and Na content, pH, base saturation and fine root biomass all significantly differed between humus layers of different species. Since the climate, parent material, land use history and soil type were similar, the differences can be ascribed to tree species. Spruce stands had the largest amounts of carbon stored down to 30 cm depth in mineral soil (7.3 kg C m−2), whereas birch stands, with the lowest production, smallest amount of litterfall and lowest C:N ratio in litter and humus, had the smallest carbon pool (4.1 kg C m−2), with pine intermediate (4.9 kg C m−2). Similarly, soil nitrogen pools amounted to 349, 269, and 240 g N m−2 for spruce, pine, and birch stands, respectively. The humus layer in birch stands was thin and mixed with mineral soil, and soil pH was highest in the birch stands. Spruce had the thickest humus layer with the lowest pH.  相似文献   

4.
The profitability of growing a naturally emerged birch (Betula pendula Roth or Betula pubescens Ehrh.) overstory in a young Norway spruce (Picea abies (L.) Karst.) plantation was examined with empirical stand structure, growth and yield, logging cost, and logging damage models. In the projected alternatives, an overstory of either birch species was thinned to 200–1000 stems per ha at the age of 15 years and retained for another 15 years. Development of the remaining spruce stand was simulated up to rotation age (70–85 years). Alternative treatments included removing the overstory completely at 15 years, and managing a pure spruce stand that was kept free of birch throughout.

Growing a birch overstory of 200–1000 stems per ha up to age 30 years resulted in a 61–93 m3 ha−1 or 9.1–16.8% yield loss for the spruce stand due to growth retardation, and a mortality of 382–498 out of 1900 stems per ha through logging damage. This was compensated for or exceeded by the additional yield of the birch (54–173 m3 ha−1) except for the lowest stocking (200–400 stems per ha) alternatives with B. pubescens. Treatment regimes with a birch overstory were clearly the most profitable alternatives, yielding up to 151% (B. pendula) and 113% (B. pubescens) of the net present value of the pure spruce alternative, at 4% interest rate. Removing the birch overstory already at 15 years was the least profitable alternative with 79 and 83% net present values, respectively. The most profitable treatment with current technology, price, and cost structure appears to be to grow 500–800 birch per ha up to about age 40 years for B. pendula and 45–50 years for B. pubescens.  相似文献   


5.

Mean age, mean and top heights and yield were studied in 20 mixed stands of birch ( Betula pubescens Ehrh. and B. pendula Roth) and Picea abies (L.) Karst. and nine mixed stands of birch and Pinus sylvestris L. in south-eastern Norway. Each mixed stand and the adjacent pure coniferous stand (control) were growing under the same site conditions and had not been commercially thinned. There were no significant differences in mean age at breast height or in top heights between birch and conifers in the mixed stands, while mean height was significantly higher for birch than for spruce. A growth index was calculated based on total volume and age at breast height. For the spruce sites the growth in young mixed stands (birch < 17 m) was superior to that of pure spruce, while the difference was insignificant in older stands. The growth index correlated positively with the ratio between generatively and vegetatively regenerated birch trees, and negatively with the age of the oldest species in the mixture and with site quality. For the pine sites there was no significant difference in the growth index between mixed birch-pine and pure pine stands. A mixture effect of birch on the volume yield of spruce or pine could not be demonstrated.  相似文献   

6.
A process-based ecosystem model was used to assess the impacts of changing climate on net photosynthesis and total stem wood growth in relation to water availability in two unmanaged Norway spruce (Picea abies) dominant stands with a mixture of Scots pine (Pinus sylvestris) and birch (Betula sp.). The mixed stands were grown over a 100-year rotation (2000-99) in southern and northern Finland with initial species shares of 50, 25 and 25% for Norway spruce, Scots pine and birch, respectively. In addition, pure Norway spruce, Scots pine and birch stands were used as a comparison to identify whether species' response is different in mixed and pure stands. Soil type and moisture conditions (moderate drought) were expected to be the same at the beginning of the simulations irrespective of site location. Regardless of tree species, both annual net canopy photosynthesis (P(nc)) and total stem wood growth (V(s)) were, on average, lower on the southern site under the changing climate compared with the current climate (difference increasing toward the end of the rotation); the opposite was the case for the northern site. Regarding the stand water budget, evapotranspiration (E(T)) was higher under the changing climate regardless of site location. Transpiration and evaporation from the canopy affected water depletion the most. Norway spruce and birch accounted for most of the water depletion in mixed stands on both sites regardless of climatic condition. The annual soil water deficit (W(d)) was higher on the southern site under the changing climate. On the northern site, the situation was the opposite. According to our results, the growth of pure Norway spruce stands in southern Finland could be even lower than the growth of Norway spruce in mixed stands under the changing climate. The opposite was found for pure Scots pine and birch stands due to lower water depletion. This indicates that in the future the management should be properly adapted to climate change in order to sustain the productivity of mixed stands dominated by Norway spruce.  相似文献   

7.
A rain shelter experiment was conducted in a 90‐year‐old Norway spruce stand, in the Kysucké Beskydy Mts (Slovakia). Three rain shelters were constructed in the stand to prevent the rainfall from reaching the soil and to reduce water availability in the rhizosphere. Fine root biomass and necromass were repeatedly measured throughout a growing season by soil coring. We established the quantities of fine root biomass (live) and necromass (dead) at soil depths of 0–5, 5–15, 15–25 and 25–35 cm. Significant differences in soil moisture contents between control and drought plots were found in the top 15 cm of soil after 20 weeks of rainfall manipulation (lasting from early June to late October). Our observations show that even relatively light drought decreased total fine root biomass from 272.0 to 242.8 g m?2 and increased the amount of necromass from 79.2 to 101.2 g m?2 in the top 35 cm of soil. Very fine roots (VFR), that is, those with diameter up to 1 mm, were more affected than total fine roots defined as 0–2 mm. The effect of reduced water availability was depth‐specific; as a result, we observed a modification of vertical distribution of fine roots. More roots in drought treatment were produced in the wetter soil horizons at 25–35 cm depth than at the surface. We conclude that fine and VFR systems of Norway spruce have the capacity to re‐allocate resources to roots at different depths in response to environmental signals, resulting in changes in necromass to biomass ratio.  相似文献   

8.
The purpose of this study was to examine the effects of type and amount of ground vegetation and substrate on the germination and early survival of Norway spruce (Picea abies (L.) Karst) seedlings in uneven-aged stands on mineral soils in Southern Finland. The study was conducted on two permanent plots representing average site quality for spruce stands in the area. A gradient in selection cut intensity and residual stand density was imposed across each of the two permanent plots. Regeneration and vegetation surveys were repeated annually on a grid of unmanipulated microplots and on a set of soil treatment microplots. The results indicated that the number of spruce germinants per unit area was positively correlated with herb cover but negatively correlated with total plant cover. Germinant density was also related to vegetation-free seedbed conditions, high overstory competition index and presence of larger spruce seedlings. Regeneration microsites dominated by moss or with plants with a relatively high light demand were associated with low germinant numbers. Bare humus layer and decayed stumps seemed to promote germinant emergence. Exposure of mineral soil resulted in germination rates that were 4.3–6.9 times higher than in undisturbed conditions during the first growing season.  相似文献   

9.
The effect of tree species mixture on stand volume yield and on tree-species-specific diameter and height growth rates were analysed in managed mixed stands of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Ehrn.).Data were obtained from 14 repeatedly measured stands located in Southern Finland on mineral soil sites with varying admixture of Scots pine and silver birch. Statistical analysis was carried out for studying the effect of species mixture on the development of stand characteristics. For the analysis, the plots were categorised into three groups (plot types) according to the species dominance. In order to analyse species-specific growth rates, individual-tree mixed linear growth models for tree diameter and height growth were developed for both tree species.The results clearly show that the yield of the managed mid-rotation, mixed stands was greater for stands dominated by Scots pine than for stands dominated by birch, and the stand volume increment decreased with an increasing proportion of silver birch. Analysis of diameter and height growth by tree species revealed that the main reason for this pattern is the negative impact of birch competition on the growth of pine trees. The increase in diameter of pine was clearly hampered if the proportion of birch was high. An abundance of birch also slightly decreased the growth in height of Scots pine, although the effect was less than on diameter growth. Species mixture did not affect the diameter growth of birch but did have a significant effect on height development. Height growth of birch was considerably greater in pine-dominated stands than in birch-dominated stands. In pine-dominated mixed stands, the height growth of birch was quite close to that of dominant pine trees, and birches can endure in competition with pines for light.The results apply for even-aged and single-storey managed stands, where stocking density and structure are controlled with pre-commercial and commercial thinnings. The results are not applicable to unmanaged mixed stands undergoing self-thinning. This study provides new information on mixed stands from a silvicultural perspective, which can be applied in decisions involving the management of mixed stands.  相似文献   

10.
Structure classes and changes in the vegetation attributable to the altitude of the terrain were studied in the Hammastunturi virgin coniferous forest area. The area is located in Finnish Lapland, approximately 250–350 km to the north of the Arctic Circle (68°15'N; 68°45'N). The following forest classes were formed on the basis of the ground vegetation and stand coverage: (1) Pine stands, with birch admixture (n=14), (2) Vaccinium‐vitis‐idaea‐type pine stands (n = 24), (3) Vaccinium myrtillus‐type pine stands (n = 17), (4) mixed stands, dominated by pine (n = 8), (5) mixed stands, dominated by spruce (n=18), (6) paludified spruce stands (n = 3) and (7) vigorously growing spruce stands (n = 6). The forest structure classes were depicted using DCA ordinates. The differences were more distinct than those between the normal forest type classifications of the same sample plots. The basal area and volume of pine decreased with increasing altitude, while the proportion of spruce on those sample plots where it was present increased. Birch was present at an equal frequency at all altitudes. The floral composition and between‐species abundance changed along with increasing altitude. The change in floral frequencies could not be designated to a specific altitude zone.  相似文献   

11.
Morphology and vertical distribution patterns of spruce and beech live fine roots (diameter ≤2 mm) were studied using a soil core method in three comparable mature stands in the Solling: (1) pure beech, (2) pure spruce and (3) mixed spruce–beech. This study was aimed at determining the effects of interspecific competition on fine root structure and spatial fine root distribution of both species. A vertical stratification of beech and spruce fine root systems was found in the mixed stand due to a shift in beech fine roots from upper to lower soil layers. Moreover, compared to pure beech, a significantly higher specific root length (SRL, P<0.05) and specific surface area (SSA, P<0.05) were found for beech admixed with spruce (pure beech/mixed beech SRL 16.1–23.4 m g−1, SSA 286–367 cm2 g−1). Both indicate a flexible ‘foraging’ strategy of beech tending to increase soil exploitation and space sequestration efficiency in soil layers less occupied by competitors. Spruce, in contrast, followed a more conservative strategy keeping the shallow vertical rooting and the root morphology quite constant in both pure and mixed stands (pure spruce/mixed spruce SRL 9.6/7.7 m g−1, P>0.10; SSA 225/212 cm2 g−1, P>0.10). Symmetric competition belowground between mixed beech and spruce was observed since live fine roots of both species were under-represented compared to pure stand. However, the higher space sequestration efficiency suggests a higher competitive ability of beech belowground.  相似文献   

12.
This study evaluates the effect of silvicultural and exploitative interventions on soil organic carbon (SOC) in Chilean Lenga (Nothofagus pumilio (Poepp et Endl.) Krasser) forests in south Patagonia. We analyzed SOC and the organic soil horizons in five stands at different stages of development: intact native forest (NI); a 3-year-old shelterwood stand (S3); an 8-year-old shelterwood stand (S8); a 14-year-old stand that was initially treated with shelterwood and subsequently final cut (10 years after the first intervention) (S14), and a 25-year-old stand subject to a exploitative intervention (E25). The SOC under the forest stands, down to a depth of 50 cm (including the Oi horizon), was 60, 55, 71, 85, and 67 Mg ha−1 for the NI, S3, S8, S14, and E25 forest stands, respectively. A significant decrease in SOC occurred 3 years after an intensive shelterwood cut (S3), particularly in the first 5 cm of the mineral soil. Slightly higher carbon contents were observed in the upper horizons of the mineral soil in both the S8 and S14 stands. Consequently, the applied shelterwood system appears to generate only short-term losses of SOC in the Lenga forest. Soil organic carbon increased over the medium term but decreased to the level observed in intact native forests over the long term. Regeneration, which influences stand microclimate (a factor in SOC storage) and provides an important source of organic soil material, was identified as one of the most important factors influencing SOC.  相似文献   

13.
The growth, technical quality and nutritional status of pure and mixed silver birch (Betula pendula Roth) and downy birch (Betula pubescens Ehrh.) plantations were studied 21 and 22 years after planting on afforested organic soil arable land and on upland forest soil. In mixtures, 50% of both birch species was planted. Silver birch trees grew better, but had higher mortality than downy birch trees on both sites. Mortality of both species was highest, and the difference in their growth smallest, on organic soil. In pure stands on organic soil, downy birch dominant height, diameter and mean volume were 96%, 92% and 82% of those of silver birch and on mineral soil 87%, 84% and 60%, correspondingly. On mineral soil, silver birch had a higher mean annual increment (MAI) (5.8 m3 ha?1a?1) than downy birch (3.9 m3 ha?1a?1), but on organic soil the MAI of both species was similar (3.3–3.4 m3 ha?1 a?1). Planting birches in mixture did not affect the growth of the trees on organic soil. On mineral soil, the mean diameter and mean volume of silver birch trees were higher in mixed than in pure plantations. The technical stem quality of both tree species was low. On mineral soils, pure silver birch is more productive than mixture, but on peat soil the higher growth of silver birch could contribute to increased productivity and downy birch would ensure sufficient survival for future timber production.  相似文献   

14.
【目的】比较秦岭辛家山林场云杉和红桦天然林土壤有机碳密度的估算结果,检验新方法通过扣除根系体积而提高的估算精度。【方法】分别估算矿质土层(表土层、心土层和底土层)和有机土层(凋落物的未/半分解层和完全分解层)的有机碳密度。在现有方法的基础上通过扣除林木根系体积含量来提高矿质土层有机碳密度的估算精度。各层林木根系体积含量的估算方法为:首先,使用前人提出的回归方程估算出单株林木根系生物量,乘以林木生长密度得到单位面积林地的根系总生物量;其次,通过采集部分根系样品测定其生物量和体积,并计算出根系样本的密度以代表整体根系的密度;然后,通过单位面积林地的根系总生物量除以根系的密度计算出单位面积林地的根系总体积;最后,利用前人研究得出的根系沿深度的分布规律,将单位面积根系总体积分配到各土层,计算出根系体积含量。对有机土层有机碳密度的估算,使用林木平均地径估算林木根基部所占面积,将有机土层中含有的林木体积扣除。此外,由于有机土层的各组分分布极不均匀,本研究依据来源器官和物理形态对凋落物(有机土层)中的不同成分进行了细致的分组,分别测定各组分的有机碳密度。【结果】云杉林表土层、心土层和底土层的厚度分别为19.10、14.20和31.03 cm,红桦林则分别为18.57、15.13和28.13 cm;云杉林表土层、心土层和底土层的有机碳含量分别为(44.56±3.72)、(25.63±1.77)和(10.79±2.28)g ·kg^-1 ,红桦林的分别为(34.11±5.46)、(19.06±4.95)和(11.02±3.86)g·kg^-1;2种林分有机土层各组分有机碳含量差异显著(P<0.05),凋落物中枝条、根系、云杉球果和苔藓的有机碳含量均大于600 g·kg^-1 ,叶片次之,云杉林和红桦林分别为(458.90±46.81)和(420.72±55.66)g·kg^-1 ,其余难以分辨的细颗粒含量最低均小于300 g·kg^-1;在矿质土层,云杉林各层每公顷根系体积(及体积比例)分别为表土层66.81(3.5%)、心土层20.69(1.5%)以及底土层9.18(0.3%)m^3,红桦林则分别为50.57(2.7%)、31.75(2.1%)和17.22(0.6%)m^3;使用改进公式估算的云杉林矿质土层有机碳密度为16.58 kg ·m^-2 ,有机土层有机碳密度为4.26 kg ·m^-2 ,完全分解层和半分解层分别占84%和16%,矿质土层和有机土层有机碳密度分别较原方法降低2.13%和0.73%;使用改进公式估算的红桦林矿质土层有机碳密度为 14.06 kg ·m^-2 ,有机土层碳密度为3.49 kg ·m^-2 ,分解层和半分解层分别占90%和10%,矿质土层和有机土层有机碳密度分别较原方法降低1.61%和0.48%。【结论】去除根系体积含量后,云杉林与红桦林的土壤总有机碳密度估算值分别降低1.85%和1.39%,这意味着目前预测的林地土壤碳储量可能普遍偏高。  相似文献   

15.
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16.
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17.
Early management of the regenerated seedlings shapes the future stand properties. To address these issues, pre-commercial thinning (PCT) and control treatments were applied to planted Norway spruce (Picea abies L. Karst) and naturally regenerated birch (Betula pendula Roth., Betula pubescens Ehrh.) stands in forest experiments in southern Sweden (lat. 56–57?N) containing 1.1–5.5?m tall saplings. The treatments were retention of 1000 or 2000?stems?ha?1 of Norway spruce, with no birch or birch at 1000?stems?ha?1. Treatments were replicated with and without annual removal of birch sprouts from stumps. The periodic annual increment (PAI) over five years was calculated for total stand volume and individual trees. The mean PAI of dominant trees was significantly higher both following all PCT treatments than controls, and following low rather than high-density PCT. Birch retention did not affect growth of the dominant trees but PAI was lower in plots with uncontrolled sprouting. The PAI of birch was significantly higher in low-density Norway spruce plots than in control plots and the high-density plots. The treatment response was significant even in stands with initial heights of only 1–2?m.  相似文献   

18.
Variability in short root morphology of the three main tree species of Europe's boreal forest (Norway spruce (Picea abies L. Karst.), Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth)) was investigated in four stands along a latitudinal gradient from northern Finland to southern Estonia. Silver birch and Scots pine were present in three stands and Norway spruce was present in all stands. For three fertile Norway spruce stands, fine root biomass and number of root tips per stand area or unit basal area were assessed from north to south. Principal component analysis indicated that short root morphology was significantly affected by tree species and site, which together explained 34.7% of the total variability. The range of variation in mean specific root area (SRA) was 51-74, 60-70 and 84-124 m(2) kg(-1) for Norway spruce, Scots pine and silver birch, respectively, and the corresponding ranges for specific root length were 37-47, 40-48 and 87-97 m g(-1). The range of variation in root tissue density of Norway spruce, Scots pine and silver birch was 113-182, 127-158 and 81-156 kg m(-3), respectively. Sensitivity of short root morphology to site conditions decreased in the order: Norway spruce > silver birch > Scots pine. Short root SRA increased with site fertility in all species. In Norway spruce, fine root biomass and number of root tips per m(2) decreased from north to south. The differences in morphological parameters among sites were significant but smaller than the site differences in fine root biomass and number of root tips.  相似文献   

19.
From 2002 to 2003, based on the investigation of sample plots and stem analysis of remained plantation communities in the areas of returning farmland to forest in the 1980s in Datong County, Qinghai Province, this paper studies tree productivity and moisture potential productivity of six types of plantations on the land of returning farmland to forest, such as green poplar (Populus cathayana Rehd.) and shrub mixed forest, Asia white birch (Betula platyphylla) and China spruce (Picea asperata) mixed forest, Dahurian larch (Larix gmelinii) pure forest, China spruce pure forest and Asia white birch pure forest and so on. The results show that: in sub-humid region of Loess Plateau, 3 000 trees per hm2 is a proper standard of planting density. Under current condition, the productivity index of green poplar and shrub mixed forest, Asia white birch pure forest, China spruce pure forest, and Asia white birch and China spruce mixed forest with the density of 2 100–3 333 trees per hm2 can serve as potential productivity standard of actual biomass of arbor established forest. In sub-humid area, Thornthwaite Model is adopted to estimate plant climate potential productivity, which is about 8 462 kg&#8226;hm–2&#8226;a–1. The actual potential water productive efficiency of Purplecone spruce (Picea purpurea) and Asia white birch pure established forest are 17.22 and 22.14 kg&#8226;mm–1&#8226;hm–2&#8226;a–1 respectively, and that of green poplar and shrub mixed established forest, and Asia white birch and China spruce mixed established forest are 21.14 and 19.09 kg&#8226;mm–1&#8226;hm–2&#8226;a–1 respectively. The potential productivity of green poplar and shrub mixed forest, Asia white birch and China spruce mixed forest, China spruce pure forest and Asia white birch pure forest which have grown into forest with the density of 3 000 trees per hm2 have attained or been close to that of local climax community, which is local maximum tree productivity at present. These types of forestation models are the developing direction of the returning farmland to forest project.  相似文献   

20.
Fine root distribution and turnover were investigated in ca. 40-year-old pure Norway spruce (Picea abies Karst.) stands in Germany, growing on four sites that differed in soil acidity (Eberg?tzen < Barbis < Fichtelgebirge = Harz). The density of fine root biomass and necromass in different soil horizons differed among the sites. At one of the most acidic sites (Harz), fine root density in the humus layer was more than twice that at the least acidic site (Eberg?tzen). At the two most acidic sites, Fichtelgebirge and Harz, the ratio of biomass to necromass was significantly lower than at Eberg?tzen and Barbis, particularly in the subsoil layer. In each stand, clear vertical gradients in fine root length density and root tip density were observed. Most of the roots and the root tips were in the humus layer and in the first mineral soil horizon (0-10 cm). There was a significantly different decrease in specific root length (cm gDM (-1)) and specific root tip density (root tips gDM (-1)) in the more acidified stands Fichtelgebirge and Harz compared with Eberg?tzen and Barbis. Fine root production estimated by ingrowth cores and a net method was approximately twice as high in the more acidic stands Fichtelgebirge and Harz compared with Eberg?tzen and Barbis. Rates of living fine root biomass turnover were higher at the Fichtelgebirge and Harz sites than at the Eberg?tzen site. Rates of necromass turnover were similar at all sites. The results suggest that the accumulation of necromass was not due to a slower disappearance at the more acid sites, but to earlier root death. Roots contributed 46% to root + needle litter and 32% to root + total aboveground litter at the Harz site in 1997.  相似文献   

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