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1.
Jorge Olivar Stella Bogino Cyrille Rathgeber Vivien Bonnesoeur Felipe Bravo 《Annals of Forest Science》2014,71(3):395-404
? Context
Modification of stand density by thinning may buffer the response of tree growth and vigor to changes in climate by enhancing soil water availability.? Aims
We tested the impact of thinning intensity on cambial growth of Aleppo pine (Pinus halepensis L.) under semi-arid, Mediterranean conditions.? Methods
A multiple thinning experiment was established on an Aleppo pine plantation in Spain. We analysed the stem growth dynamics of two different crown classes under four different thinning intensities (15 %, 30 %, and 45 % removal of the basal area) for 2 years, based on biweekly band dendrometer recordings. Local relative extractable soil water was derived from the use of a water balance model Biljou© (available at https://appgeodb.nancy.inra.fr/biljou/) and used as an explanatory variable.? Results
Radial growth was mainly controlled by soil water availability during the growing season, and differed by crown class. The growth rates of dominant trees were significantly higher than the growth rates of suppressed trees. Removal of 30 % and 45 % of the initial basal area produced a growth release in both dominant and suppressed trees that did not occur under less intense thinning treatments.? Conclusions
Soil water availability was the main driver of radial growth during the growing season. Forest management confirmed its value for ameliorating the effects of water limitations on individual tree growth. These results may help managers understand how altering stand density will differentially affect diameter growth responses of Aleppo pine to short-term climatic fluctuations, promoting forests that are resilient to future climatic conditions. 相似文献2.
Fire severity as a key factor in post-fire regeneration of Pinus pinaster (Ait.) in Central Portugal
Context
Mediterranean pine woodlands are strongly affected by wildfires; however, there are knowledge gaps in the role of fire severity on Pinus pinaster Ait. regeneration.Objectives
The principal questions were: (a) does post-fire regeneration of Maritime Pine differ where canopies were consumed to a low vs. high degree and (b) which factors, besides fire severity, could explain these differences.Methods
Pine recruitment was monitored from 2 to 36?months after a summer wildfire in 18 25?m2 plots, equally divided over two fire severity classes based on crown consumption. Besides the degree of crown consumption, three quantitative fire severity indices as well as post-fire site conditions, seed input and understory vegetation recovery were measured.Results
Pine seedling densities were consistently higher in the plots with low than high crown consumption but due to marked spatial variability, they were only significantly different at two out of the eight sampling occasions. This variability could be explained by a quantitative index based on the diameter of twigs, as well as by seed input and post-fire site conditions (ash and litter cover).Conclusion
Fire severity was found to strongly affect the pine recruitment following wildfire, using both a qualitative visual severity index and a quantitative, more labour-intensive one. 相似文献3.
Context
Excessive recruitment in post-fire regeneration of Pinus halepensis can require pre-commercial thinning. The 1994 Moratalla fire (Spain) and the thinning applied there since 2004 provided good conditions for testing pre-commercial thinning effectiveness.Aims
To analyse pre-commercial thinning effects on tree size, reproductive potential, stem biomass and annual growth in 15-year-old saplings.Methods
Twenty nine circular plots (5 m radius) were established based on (1) years since thinning and (2) aspect. Non-thinned plots were included as controls. Sapling variables were measured and pine cones were counted and classified according to maturity. Three saplings per plot were felled, five sample disks were extracted from each and tree rings were analysed with Windendro® software.Results
Sapling size, annual growth and stem biomass were higher in thinned plots than in control plots 2–4 years after thinning. Annual growth declined five years after thinning. Cone counts were higher 2–5 years after thinning, depending on the cone type. Aspect influenced some variables.Conclusion
Thinning accelerated stand maturity but positive effects were indiscernible until 2–5 years later. Serotinous cones, which are fundamental to regeneration after fire, increased in number after 5 years. The effects of aspect were inconclusive due to interactions with years since thinning. 相似文献4.
Iciar Alberdi Isabel Cañellas Laura Hernández Sonia Condés 《Annals of Forest Science》2013,70(3):277-285
Context
Old-growth trees play a very important role in the maintenance of biodiversity in forests. However, no clear definition is yet available to help identify them since tree age is usually not recorded in National Forest Inventories.Aims
To develop and test a new method to identify old-growth trees using a species-specific threshold for tree diameter in National Forest Inventories.Methods
Different nonlinear mixed models for diameter–age were generated using data from the Spanish Forest Inventory in order to identify the most appropriate one for Aleppo pine in its South-western distribution area. The asymptote of the optimal model indicates the threshold diameter for defining an old-growth tree. Additionally, five site index curves were examined to analyze the influence of site quality on these models.Results
The Hossfled III mixed model was found to be the most appropriate to fit diameter–age curves for Aleppo pine trees. The overall diameter at breast height threshold for old-growth trees was 40.6 cm, although over a range of sites with increasing site quality, the threshold figure was 36.0, 38.0, 40.4, 43.1, and 46.3 cm, respectively.Conclusions
This method allows the identification of old-growth trees and therefore of biodiversity hotspots, thus providing decision makers with a useful tool for management purposes. 相似文献5.
? Context
Maritime pine (Pinus pinaster Aiton) is one of the most important Portuguese species, growing in pure stands ranging from even-aged to multi-aged structures. Current growth and yield models were developed only for even-aged, managed stands and/or for very specific regions of Portugal.? Aims
This paper focuses on the validation of the existing size-class model PBRAVO, adapted to even-aged stands, and on the subsequent development of a single tree distance-dependent growth and yield model (PBIRROL), both in distance-independent and distance-dependent versions, for uneven-aged stands.? Methods
The new model is composed of four modules, each with a set of sub-models for: tree variable prediction, tree volume prediction, future tree list prediction and growth projection.? Results
The evaluation of the PBRAVO and PBIRROL models showed that the new model gives more accurate predictions. Moreover, medium-term simulations provided consistent and logical predictions.? Conclusion
It was verified that individual tree models are more suited to simulate poorly managed uneven-aged stands than diameter distribution models. No clear superiority of distance-dependent models was found over models using just distance-independent measures of inter-tree competition. 相似文献6.
Survival to prescribed fire of plantation-grown Corsican black pine in northern Portugal 总被引：1，自引：0，他引：1
Context
The current fire regime threatens black pine (Pinus nigra Arn.) persistence in the Mediterranean Basin, which recommends larger-scale fuel treatments. Prescribed burning is an option for stand protection but its use in young stands (which are particularly at risk) is hindered by the scarce knowledge on post-fire tree survival.Aims
The objectives were to characterize bark thickness as a fire-resistance trait in P. nigra and to describe how post-fire tree survival responds to tree size and fire effects in a 16-year-old plantation.Methods
Bark thickness was related to diameter at breast height and height in the stem. Metrics describing tree size and stem and crown damage were measured 1?year after prescribed burning in 259 trees. Tree survival was modeled with logistic regression and Classification and Regression Tree analysis.Results
Bark thickness increased linearly with diameter at breast height (dbh) and decreased with height in the stem. Tree survival was primarily a function of crown injury. Stem damage was an influent factor in small trees.Conclusion
Due to thinner bark and lower tolerance to crown damage, young P. nigra trees are less fire-resistant than other Mediterranean pines, e.g., Pinus pinaster. Prescribed fire should not be attempted if dbh <10?cm. Mechanical clearing is the treatment of choice in young stands with a significant shrub layer. 相似文献7.
? Context
The effects of leaf inclination on plant light capture, growth, and water balance of monospecific canopies are well documented, but we still lack information on such effects in the case of multispecific canopies.? Aims
We investigated the effects of leaf inclination on the absorption of photosynthetically active radiation (PAR) of a mixed forest.? Methods
We ran a 3D mechanistic radiation transfer model for a Mediterranean forest where Pinus halepensis makes the upper strata while Quercus ilex occupies the lower strata. As factors, we included (1) the distributions of leaf inclinations that ranged from vertical to horizontal (including the actual inclinations), (2) the fraction of diffuse light, sun position, and leaf area index (LAI), and (3) the Pinus/Quercus LAI ratio.? Results
Simulated PAR absorption was more than twice as sensitive to leaf inclination in oaks than in pines because oaks depended on PAR transmitted below the pine layer. The extent of the effect depended on season, fraction of diffuse light, LAI, and vegetation spatial structure. None of the observed inclinations maximized PAR absorption, suggesting a trade-off with water economy.? Conclusion
Erroneous assumptions about leaf inclination lead to larger errors when modelling heterogeneous, mixed canopies. This also highlights potential caveats when using models that do not account for the spatial structure of canopies. 相似文献8.
Context
In the Mediterranean area, different post-fire management strategies are used for coppices of resprouting species to promote a more regular forest structure, enhance plant growth, and reduce fire risk. However, the effects of these management treatments on forest-associated fauna are unknown, which in turn could be limiting their beneficial effects.Aims
The aim of this work was to determine whether forest management of a recently burned area dominated by a vigorous resprouting tree species (Arbutus unedo L.) affects ant communities.Methods
Ant communities, sampled using pitfall traps, were examined from unmanaged and selective thinning coppices of A. unedo. Ants are here used as bioindicators of ecosystem health and surrogates for other animal groups.Results
Very limited effects of these post-fire management strategies on the structure and composition of ant communities were found. The lack of effects could be due to the reported small changes in physical conditions among treatments; or either, the most sensitive ant species to these post-fire management treatments might be the same ones affected by fire and, consequently, the ant species that would potentially be affected most were no longer in the study area.Conclusion
The lack of any significant effects caused by these post-fire management practices on the associated fauna of A. unedo coppices points out the suitability of these treatments in these circumstances. 相似文献9.
Teresa Valor Míriam Piqué Bernat C. López José Ramón González-Olabarria 《Annals of Forest Science》2013,70(5):503-513
Context
After wildfire, surviving trees are of major ecological importance as they can help in the post-fire regeneration process. Although these trees may be damaged, they may also benefit from reduced fuel hazard and competition. However, little is known about the long-term growth response of surviving trees.Aims
This study aims to explain short- to long-term variations in the postfire growth of surviving black pines in an area burnt in 1994, focusing on levels of fire severity and tree sizes.Methods
Relative basal area increments were used to detect time-course variations in postfire radial tree growth depending on fire severity. Linear mixed-effects models were used to describe the factors affecting postfire ring growth.Results
In the short term, fire caused stronger reduction in growth in small trees with increasing bole char height. However, as time since fire increased, a positive effect of fire on growth due to reduced competition counteracted the short-term fire impacts. Indeed, small surviving trees demonstrated a surge in growth 15 years after the fire.Conclusion
It was concluded that reduced competition might offset the short-term negative effects of fire in surviving black pines. 相似文献10.
Context
Although drought is generally considered the main environmental constraint in Mediterranean environments, the ability to acclimate to and tolerate frost in early developmental stages can be a determinant for seedling survival of many Mediterranean tree species like stone pine (Pinus pinea L.).Aims
The aim of this study was to assess the impact of the developmental stage of naturally regenerated stone pine individuals on tolerance to low temperature (LT) from summer to late autumn and in spring, at a highly continental site in central Spain. Specifically, we tested to what extent the differences in tolerance are related to shoot heteroblasty.Methods
We assessed LT tolerance of needles from individuals at three age classes (class C1: seedlings, class C2: 4- to 8-year-old saplings and class C3: >9-year-old saplings) over nine dates from summer to spring.Results
LT tolerance displayed severe seasonal trends and differed between age classes. It usually increased with sapling age. Such differences were tightly related to heteroblasty of the shoots. Our results point to a higher LT tolerance associated with larger leaf dry mass per unit area (LMA) values. No impact of late frosts on shoot growth rates was detected during this study.Conclusions
Developmental changes during early plant growth seem to play a role in frost tolerance of stone pine seedlings, a finding which furthers our understanding of regeneration dynamics in this species in areas with continental influence. 相似文献11.
Models of the longitudinal distribution of ring area as a function of tree and stand attributes for four major Canadian conifers 总被引：1，自引：0，他引：1
Context
It is widely accepted that ring area increment generally increases from the tree apex to the crown base and is more-or-less constant below the crown base (Pressler’s law), but few quantitative models of this distribution have been developed.Aims
The aim of this study was to develop a model of ring area increment using easily obtained crown features and other tree or stand characteristics in order to further the understanding and prediction of tree growth, form, and wood quality.Methods
The models were fit to stem analysis observations from white spruce, black spruce, balsam fir, and lodgepole pine.Results
In the final model, which includes tree crown and stand variables, ring area increment within the crown region was slightly curvilinear, the slope of ring area increment below the crown was non-zero, and the effect of butt swell was appreciable up-to a relative height of 0.10.Conclusions
The high accuracy of the mixed effects model suggests that the three-component model form is appropriate for describing ring area profiles, whereas some tree-to-tree variation remains unexplained. The tree and stand variables used in these models can be easily measured in the field or obtained from remote sensing techniques. 相似文献12.
Using density management diagrams to assess crown fire potential in Pinus pinaster Ait. stands 总被引：1，自引：0，他引：1
Ibán Gómez-Vázquez Paulo M. Fernandes Manuel Arias-Rodil Marcos Barrio-Anta Fernando Castedo-Dorado 《Annals of Forest Science》2014,71(4):473-484
Context
Density management diagrams (DMDs) are useful for designing, displaying and evaluating alternative density management regimes for a given stand-level management objective. The inclusion of variables related to crown fire potential within DMDs has not previously been considered.Aims
The aim of this study was to include isolines of variables related to crown fire initiation and spread in DMDs to enable identification of stand structures associated with different types of wildfire.Methods
Biometric and fuel data from maritime pine (Pinus pinaster Ait.) stands in NW Spain were used to construct DMDs. Different surface and crown fire behaviour models were used together to estimate crown fire potential.Results
The crown fire potential varied greatly throughout development of the maritime pine stands. Low stands were more prone to crowning. The type of crown fire was mainly determined by stand density.Conclusion
The DMDs developed can be used to identify relationships between stand structure and crown fire potential, thus enabling the design of thinning schedules aimed at reducing the likelihood of crowning. 相似文献13.
Rafał Ważny 《Annals of Forest Science》2014,71(7):801-810
Context
The requirement for rebuilding forecrop stands besides replacement of meadow vegetation with forest plants and formation of soil humus is the presence of a compatible ectomycorrhizal (ECM) fungal community.Aims
This study aims to assess ectomycorrhizal fungi diversity associated with silver fir (Abies alba Mill.) seedlings regenerating in silver fir stands and Scots pine forecrops.Methods
One-year-old seedlings were sampled in six study sites: three mature fir forests and three pine forests. ECM fungi were identified by polymerase chain reaction amplification and sequencing of the internal transcribed spacer of rDNA.Results
The mean mycorrhizal colonization exceeded 90 %. Thirty-six ectomycorrhizal taxa were identified in fir stands and 23 in pine forecrops; ten out of these species were common to both stands. The fungal communities were different between study sites (R?=?0.1721, p?=?0.0001). Tomentella stuposa was the only species present at all sites.Conclusion
Silver fir seedlings in Scots pine forecrops supported smaller ECM fungal communities than communities identified in mature silver fir stands. Nevertheless, fungal colonization of seedling roots was similar in both cases. This suggests that pine stands afforested on formerly arable land bear enough ECM species to allow survival and growth of silver fir seedlings. 相似文献14.
Context
Regular mortality (or self-thinning) is an integral part of woody plant dynamics, and the mortality model is one of the most important transition functions within dynamic growth models.Aims
The main objectives of the present study were to derive one-step stand-level mortality models for estimating the reduction in stand density with stand dominant height growth and to examine the applicability of the models in predicting regular mortality trends in even-aged natural stands and plantations.Methods
Four model formulations based on graphical examination of the main trends in the data and on defined desired properties of the selected mathematical functions were proposed and tested in algebraic difference equation form with two datasets from pine plantations and two datasets from natural broadleaved even-aged stands.Results
The mortality patterns of three of the datasets were characterized by reverse sigmoid curves, while the forth dataset exhibited a reverse J-shaped trajectory. The models analyzed adequately represented the density decrease trends of the data through sets of polymorphic curves with multiple asymptotes.Conclusion
The dominant height-dependent mortality equations provide a simple and reliable approach to modeling the regular density decrease trends and can be considered for incorporation as a submodel within the framework of a dynamic Stand Density Management Diagram. 相似文献15.
Context
The development of multiple trait selection indices for solid (structure) wood production in the Scots pine (Pinus sylvestris L.) breeding program requires genetic variances and covariances estimated among wood quality traits including stiffness.Aims
Genetic control and relationships among Scots pine growth, fiber, and wood quality traits were assessed by estimating heritability, phenotypic and genetic correlation using a Scots pine full-sib family trial.Method
Wood quality traits including clearwood and dynamic acoustic stiffness were measured using SilviScan and Hitman in a 40-year-old progeny trial and by sampling increment cores of 778 trees of 120 families. Genetic parameters were estimated using the mixed model by the ASReml software.Results
Heritability ranged from 0.147 to 0.306 for growth, earlywood, transition wood and latewood proportion traits and from 0.260 to 0.524 for fiber dimension, wood density, MFA and stiffness traits. The highly unfavorable genetic correlation between diameter and whole core density (?0.479) and clearwood stiffness (?0.506) and dynamic acoustic stiffness (?0.382) was observed in this study.Conclusion
The unfavorable genetic correlations between growth traits and stiffness indicate that multiple traits selection using optimal economic weights and optimal breeding strategies are recommended for the advanced Scots pine breeding program. 相似文献16.
Daniel Moreno-Fernández Isabel Cañellas Rafael Calama Javier Gordo Mariola Sánchez-González 《Annals of Forest Science》2013,70(8):761-768
Context
Edible stone pine (Pinus pinea L.) nut is a forest product which provides the highest incomes to the owners of stone pine forests.Aim
The objective of this work is to evaluate the effect of first thinning on growth and cone production in an artificially regenerated stand in order to determine optimum intensity.Methods
A thinning trial was installed in 2004 to compare two thinning regimes (heavy and moderate) and a control treatment. From 2004 to 2012, six inventories of forest attributes were carried out, and the cone crop was harvested annually. We evaluated the effect of thinnings on growth using repeated measures analysis of variance with a mixed model approach. With regards to cone production, we first estimated the probability of finding cones in a tree by applying a generalized mixed model and then estimated cone production by using a mixed model, including climatic variables.Results
We found that thinning had a positive influence on tree diameter increment. Thinning increased the probability of finding cones and cone production. However, significant differences between heavy and moderate thinnings were not found.Conclusion
We recommend early silvicultural treatments in stone pine stands to favor the development of trees and larger edible pine nut production. 相似文献17.
Context
Families of the Gumbel (type I), Fréchet (type II) and Weibull (type III) distributions can be combined in the generalized extreme value (GEV) family of distributions. Maximum and minimum values of diameters in forest stands can be used in forest modelling, mainly to define parameters of the functions used in diameter class models as well as in some practical cases, such as modelling maximum diameters for sawing and processing purposes.Aims
The purpose of this study was to examine and compare two extreme value distribution functions (the Gumbel and the Weibull functions) in modelling the distribution of the minimum and the maximum values of representative sets of tree diameter samples. Both of these functions were applied to the lower and upper values of the diameter distributions of the main forest species in northwest Spain: Quercus robur L., Betula pubescens Ehrh., Pinus radiata D. Don, Pinus pinaster Ait. and Pinus sylvestris L.Methods
Parameters of the Gumbel function were estimated using the mode and the moments of the distributions, and parameters of the Weibull function were estimated using the moments method.Results
In general, the Weibull distribution was the most suitable model for describing the maximum diameters. The mode method of the Gumbel yielded the best results for minimum diameters of birch and Monterrey pine. The Gumbel distribution, fitted by either the mode- or moments-based methods, proved more suitable than the Weibull distribution for describing the minimum diameters in maritime pine and Scots pine stands.Conclusion
In some cases, better results were obtained with the Gumbel than the Weibull distribution for describing the distribution of extreme diameter values in forest stands in northwest Spain. This is the first example of the application of the Gumbel distribution in forest modelling. 相似文献18.
Ecology and evolution of pine life histories 总被引：1，自引：0，他引：1
Jon E. Keeley 《Annals of Forest Science》2012,69(4):445-453
Introduction
Pinus is a diverse genus of trees widely distributed throughout the Northern Hemisphere. Understanding pine life history is critical to both conservation and fire management.Objectives
Here I lay out the different pathways of pine life history adaptation and a brief overview of pine evolution and the very significant role that fire has played.Results
Pinus originated ~150?Ma in the mid-Mesozoic Era and radiated across the northern continent of Laurasia during the Cretaceous Period. Pines have followed two evolutionary strategies interpreted as responses to competition by the newly emerging angiosperms. The Strobus lineage mostly has radiated into stressful sites of low nutrient soils and extremes in cold or heat. The Pinus (subgenus) lineage has radiated into fire-prone landscapes with diverse fire regimes. Examination of life history traits illustrates syndromes associated with fire-avoider, fire-tolerater, fire-embracer, and fire-refuge strategies.Conclusion
Understanding the current pattern of pine distribution requires interpreting their evolution in terms of climate, geology, and fire. All three of these factors have played a role since the Mesozoic origin of the genus. All are important to the appropriate management of these resources. 相似文献19.
Sergio de-Miguel Timo Pukkala Nabil Assaf Zuheir Shater 《Annals of Forest Science》2014,71(1):101-112
? Context
Biomass prediction is important when dealing for instance with carbon sequestration, wildfire modeling, or bioenergy supply. Although allometric models based on destructive sampling provide accurate estimates, alternative species-specific equations often yield considerably different biomass predictions. An important source of intra-specific variability remains unexplained.? Aims
The aims of the study were to inspect and assess intra-specific differences in aboveground biomass of Pinus brutia Ten. and to fill the gap in knowledge on biomass prediction for this species.? Methods
Two hundred one trees between 2.3 and 55.8 cm in diameter at breast height were sampled throughout the eastern- and southernmost natural distribution area of P. brutia, in Middle East, where it forms different stand structures. Allometric equations were fitted separately for two countries. The differences in biomass prediction at tree, stand, and forest level were analyzed. The effect of stand structure and past forest management was discussed.? Results
Between-country differences in total aboveground biomass were not large. However, differences in biomass stock were large when tree components were analyzed separately. Trees had higher stem biomass and lower crown biomass in dense even-aged stands than in more uneven-aged and sparse stands.? Conclusion
Biomass and carbon predictions could be improved by taking into account stand structure in biomass models. 相似文献20.
José Ramón González-Olabarria Jordi Garcia-Gonzalo Blas Mola-Yudego Timo Pukkala 《Annals of Forest Science》2017,74(3):52