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1.

Context

Loblolly pine is often grown in intensively managed plantations for wood production. In order to fully evaluate the effects of management practices on wood quality and ultimately value, it is necessary to relate mechanical properties to management practices.

Aims

The aim of this study was to evaluate the effect of planting density on mechanical properties of lumber recovered from loblolly pine trees from a 27-year-old spacing trial and develop prediction equations for modulus of elasticity and modulus of rupture from stand, tree, and board characteristics.

Methods

Regression methods were applied to sample trees from three planting densities (2,989, 1,682, and 746 trees ha?1) and used to relate mechanical properties of lumber extracted from the trees to stand, tree, and board characteristics.

Results

Initial planting density was found to be correlated with modulus of elasticity and, to a lesser extent, with modulus of rupture. Including board characteristics and utilizing the visual grade and board position as regressors produced improved prediction equations.

Conclusions

The mean modulus of elasticity declines with decreasing planting density while the variability increases, suggesting that planting density is a surrogate for frequency and size of knots. Thus, lower planting densities, while producing more lumber, may produce proportionally fewer boards of greater modulus of elasticity than higher planting densities.  相似文献   

2.

Context

Information on wood properties variation is needed by forest growers and timber processors to best utilise the available forest resource and to guide future management.

Aim

This study aims to quantify the variation in selected properties of Sitka spruce (Picea sitchensis (Bong.) Carr.) structural timber.

Methods

Twelve harvest-age stands were selected, ten trees per site were felled and processed into 301 logs. Dynamic modulus of elasticity (MOEdyn) was measured on each tree and log using portable acoustic instruments. Logs were processed into structural timber and its MOE and bending strength was determined.

Results

Overall, the timber satisfied the MOE, bending strength and density requirements for the C16 strength class. Approximately 25 % of the total variation in timber mechanical properties was attributed to between-stand differences, with the remaining 75 % attributed to within-stand differences. A series of equations were developed to predict site, tree and log-level variation in timber properties.

Conclusion

Knowledge of the site and stand factors that are associated with differences in timber properties can assist with segregation of the current resource. Portable acoustic tools can also be used to increase the stiffness of sawn timber by segregating out individual trees and logs that will yield low stiffness timber.  相似文献   

3.

? Background

A culture/density study was established in 1995 in the Lower Coastal Plain of the southeastern USA to evaluate the effects of intensive silviculture and current operational practices on the growth and yield of loblolly pine plantations across a wide range of planting densities (741–4,448 trees/ha). The operational regime consisted of bedding and herbicide application in site preparation and fertilizer applications at planting and in the eighth and 12th growing seasons. The intensive management regime had additional complete competition control, tip moths control, and more repeated fertilization treatments.

? Methods

The data from 14 locations from this split-plot experiment design with repeated measurements were analyzed with a mixed-effects model approach in terms of average DBH, average height, average dominant height, survival, stand basal area, and stand volume.

? Results

In the first few years after planting, there were no significant effects of management intensity and planting density. In later years, both management intensity and planting density significantly impacted response variables, and their interaction was only significant for average diameter at breast height (DBH). Responses to intensive management in DBH were greatest at the lowest planting densities. Intensive management resulted in larger average DBH, average height, dominant height, stand basal area, and volume. Intensively managed plots had more mortality at age 12. There were negative average DBH, average height, dominant height, and survival responses but positive stand basal area and volume responses to increasing planting density. However, there were no significant differences for planting densities above 2,224 trees/ha.

? Conclusions

The results demonstrate that both management intensity and planting density significantly affect loblolly pine productivity in the Lower Coastal Plain, and their effects are additive in nature due to the general lack of interactions.  相似文献   

4.
  • ? Direct assessment of modulus of elasticity (MOE) on standing trees is attractive for breeders to evaluate genotypes prior to selection: this can be done using the Rigidimeter, a bending-based measurement device.
  • ? In this study, we tested its reliability to properly rank genotypes by relating trunk MOE with MOEs estimated with a vibrating analysis system (Bing) on different types of conditioned wood specimens from the same trees (boards and standardised 2×2×30 cm-clear-wood specimens). One hundred and ten trees from different genotypes of hybrid larch (Larix × eurolepis) were tested.
  • ? Mean trunk MOE was 7 300 MPa with a similar value obtained for sawn boards. Clear-wood specimens MOE increased from pith to bark from less than 6 000 MPa to nearly 9 000 MPa. Moderate correlations (r = 0.48–0.61) were found at the individual tree level between trunk MOE and MOE of wood samples.
  • ? Single specimen MOE was shown to be strongly related to a linear combination of trunk MOE and sample position.
  • ? At the genotype mean level, trunk MOE was highly correlated with wood samples MOE (r = 0.80–0.91). Ranking of genotypes based on trunk MOE was mostly consistent with that based on standardised specimens.
  • ? It was concluded that besides other operational advantages which are discussed, the Rigidimeter is a valuable tool for breeders to routinely evaluate and rank genotypes for stiffness prior to further selection.
  •   相似文献   

    5.

    ? Context

    Mechanical wood properties are increasingly relevant for structural applications and are influenced by growing space availability. Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) has an increasing market share in Europe and is mainly processed to sawn timber.

    ? Aim

    A sample of 164 thinning trees was taken from two Douglas-fir long-term forestry research plots in Germany. The end-use quality of about 2,000 side and center boards was analyzed as a function of initial plant density (1,000, 2,000, and 4,000 trees per hectare) and log position within the stem.

    ? Methods

    Sawn timber quality was described by knottiness, density, modulus of elasticity, and strength. Explanatory parameters were radial position, longitudinal position, and initial plant density. All boards were strength graded visually and by the grading machine GoldenEye-706 using both X-rays for detecting densities and size as well as position of knots and laser interferometry for detecting eigenfrequency (DIN 4074, DIN 2012; EN 14081-2, CEN 2010).

    ? Results

    High plant density led to better mechanical sawn timber quality. Significant differences were especially observed between 1,000 and 2,000 trees per hectare. The yield of machine strength-graded center boards of strength class C24 increased from 50 to 89 % at low and high initial plant density, respectively.

    ? Conclusion

    Foresters are able to improve end-product quality by controlling planting density in particular. The roundwood price that foresters get should be based on the proportion of higher strength classes within logs to give incentives for a more quality-oriented forest management.  相似文献   

    6.
  • ? Environmental determinants of wood properties variation were examined in Eucalyptus globulus, a globally important hardwood plantation species, in southern Tasmania, Australia.
  • ? Radial variation in wood properties, measured with the SilviScan system, were re-scaled from distance to time abscissa using stem radial growth data measured with dendrometers. With this re-scaled data it was possible to evaluate how water availability and temperature affected wood density, microfibril angle (MFA) and fibre and vessel transverse dimensions in irrigated and non-irrigated trees.
  • ? Wood density, fibre radial diameter and MFA were sensitive to water availability. Wood density increased and fibre radial diameter decreased in response to reduced water availability. When high water availability was maintained, wood density was negatively correlated with temperature. Together, temperature and soil matric potential explained about 60% of temporal variation in wood density variation. In contrast MFA was not related to temperature but decreased with increasing water stress. Slower growing trees also had lower MFA than faster growing trees. Slower growing trees had a larger number of vessels per unit area of wood than faster growing trees within this even aged stand. However, vessel radius to the 4th power was significantly higher in faster growing trees than in slower growing trees.
  • ? Overall, E. globulus wood properties were sensitive to temporal changes in environmental conditions (particularly water availability) and associated growth rates. The data provided support for the hypothesis that growth rates are hydraulically mediated.
  •   相似文献   

    7.

    Context

    Avoidance or control of epicormic shoots is among the major silvicultural challenges for the production of high-quality oak timber. In northern Europe, contemporary oak silviculture aims to produce valuable timber on a relatively short rotation, applying early, heavy thinning combined with artificial pruning.

    Aims

    The aim of this study was to analyse the effects of pruning and stand density on the production of new epicormic shoots on young trees of pedunculate oak (Quercus robur L.).

    Methods

    The study was based on two field experiments in even-aged stands of pedunculate oak subjected to different thinning practices and early selection of potential future crop trees. From ages 13 to 15 years, stem density was reduced to 300 trees ha?1, 1,000 ha?1 or stands remained unthinned. Pruning was conducted on selected trees at ages 22–24 years. At that age, the stem density in unthinned control plots ranged from 2,500 to 3,100 ha?1. All treatments were replicated twice within each experiment.

    Results

    Pruning led to an overall increase in the total production of new epicormic shoots. More epicormic shoots were produced in the lower part of the stem (0–3 m in height) than in the upper part (3–6 m). The number of new epicormic shoots increased with increasing stand density.

    Conclusion

    Early, heavy thinning combined with high pruning at regular intervals may help shorten the rotation length for pedunculate oak without further reduction in wood quality than that which is caused by wider annual growth rings.  相似文献   

    8.

    Context

    Recruitment is an important process in forest stand dynamics, especially in uneven-aged stands. Continuous recruitment is a prerequisite for diverse, uneven-aged silvicultural systems, but patterns may vary significantly.

    Aims

    The main goals of the study were to examine the recruitment of the main tree species in selection and irregular shelterwood stands in silver fir?CEuropean beech?CNorway spruce forests and to determine the main predictors of the recruitment occurrence.

    Methods

    Data from 5,486 permanent inventory plots were used to study recruitment of saplings into the tree layer (diameter at breast height ??10?cm).

    Results

    Recruitment rate differed significantly between selection (7.6?trees?ha?1?year?1) and irregular shelterwood (26.1?trees?ha?1?year?1) stands. Shade-tolerant fir and beech recruited with higher probability in selection stands, while light-dependent sycamore recruited with higher probability in irregular stands. In addition, forest types, soil pH, stand basal area, mean diameter, and the basal area of the same tree species with respect to recruitment were found to be important predictors of recruitment occurrence.

    Conclusions

    The application of different uneven-aged silvicultural systems and their forms makes it possible to considerably influence the future tree species composition of uneven-aged forests.  相似文献   

    9.

    ? Context

    Modification of stand density by thinning may buffer the response of tree growth and vigor to changes in climate by enhancing soil water availability.

    ? Aims

    We tested the impact of thinning intensity on cambial growth of Aleppo pine (Pinus halepensis L.) under semi-arid, Mediterranean conditions.

    ? Methods

    A multiple thinning experiment was established on an Aleppo pine plantation in Spain. We analysed the stem growth dynamics of two different crown classes under four different thinning intensities (15 %, 30 %, and 45 % removal of the basal area) for 2 years, based on biweekly band dendrometer recordings. Local relative extractable soil water was derived from the use of a water balance model Biljou© (available at https://appgeodb.nancy.inra.fr/biljou/) and used as an explanatory variable.

    ? Results

    Radial growth was mainly controlled by soil water availability during the growing season, and differed by crown class. The growth rates of dominant trees were significantly higher than the growth rates of suppressed trees. Removal of 30 % and 45 % of the initial basal area produced a growth release in both dominant and suppressed trees that did not occur under less intense thinning treatments.

    ? Conclusions

    Soil water availability was the main driver of radial growth during the growing season. Forest management confirmed its value for ameliorating the effects of water limitations on individual tree growth. These results may help managers understand how altering stand density will differentially affect diameter growth responses of Aleppo pine to short-term climatic fluctuations, promoting forests that are resilient to future climatic conditions.  相似文献   

    10.
    11.

    Context

    Cluster planting has become a conventional establishment method for oaks in Central Europe, where the spacing of seedlings within clusters varies between ‘nests’ (0.2?×?0.2 m) and ‘groups’ (1?×?1 m). Although the space between clusters is expected to fill with voluntary regeneration, its competitive effect on oak growth and quality had not been studied yet.

    Aims

    The aim of the study was to analyse the effects of inter- and intraspecific interactions on growth and quality of oaks grown in cluster plantings by quantifying the influence of neighbouring trees. In addition, we analysed whether the spatial position of oaks within groups (inner section or periphery) influenced their quality development.

    Methods

    Using Hegyi’s competition index, the influence of competition from intra- and interspecific trees from early, mid- and late-successional species, on diameter, height, slenderness and quality (length of branch-free bole) of 10- to 26-year-old oaks grown in cluster planting stands was quantified at seven sites in Baden-Württemberg and Hessen, Germany.

    Results

    In general, mid- and late-successional trees exerted a stronger competitive influence on growth of target oaks in clusters than the conspecific oaks and pioneer tree species. Oak quality development benefited from intraspecific competition, but self-pruning was not further promoted through additional interspecific competition. Within groups, inner oaks had a higher probability of developing into potential future crop trees than outer oaks.

    Conclusion

    Our study showed that intra- and interspecific competition had different effects on target oak trees and that these effect differed between nest and group plantings. The development of naturally regenerated and planted trainer trees in group plantings should be monitored carefully and if necessary be controlled through thinning or pollarding.  相似文献   

    12.

    ? Context

    There are many stand property–density relationships in ecology which represent emergent properties of plant populations. Examples include self-thinning, competition–density effect, constant final yield, and age-related decline in stand growth. We suggest that these relationships are different aspects of a general framework of stand property–density relationships.

    ? Aims

    We aim to illustrate the generalities and ecological implications of stand property–density relationships, and organize them in a comprehensive framework.

    ? Methods

    We illustrate relationships between stand property and density (1) at one point in time, (2) over time, and (3) independent of time. We review the consequences of considering different variables to characterize stand property (mean tree size, mean tree growth, stand growth, stand yield, stand leaf area).

    ? Results

    We provide a framework that integrates the broad categories of stand property–density relationships and individual expressions of these relationships. For example, we conclude that constant final yield is a special case of the growth–growing stock relationship for life forms were yield is a reasonable approximation of growth (non-woody plants).

    ? Conclusion

    There is support in the literature for leaf area being broadly integrative with respect to various expressions of stand property–density relationships. We show how this is and suggest implications for plant population ecology and forest management.  相似文献   

    13.
  • ? In the tropical intertidal zones, little is known on water uptake by mangroves. Transpiration rates are generally measured at leaf level, but few studies exist on water use at tree or stand levels.
  • ? The objective of this study was to measure sap flow in trees of different sizes to appreciate the range of variation in water use that may exist in a site dominated by 80% mature Avicennia germinans.
  • ? The results showed that from the dry to the wet season the mean water use increased from 3.2 to 5.3 dm3 d?1 in small trees (DBH ~ 13 cm), from 11.5 to 30.8 dm3 d?1 in medium trees (~24 cm) and from 40.8 to 64.1 dm3 d?1 in large ones (~45 cm).
  • ? Sapwood remained active up to a depth of 8 cm with radial variations within the stem. Weak correlations were obtained with VPD and net radiation.
  • ? This study confirmed that transpiration was larger under low levels of salinity. Water use at stand level (~1900 living stems ha?1) was estimated to be in the range of 5.8 to 11.8 m3 ha?1 d?1 according to the season.
  •   相似文献   

    14.

    Context

    The reforestation of degraded tropical pastures requires innovative tree planting systems that consider land user needs.

    Aim

    We established a silvopastoral reforestation system and assessed the effects of companion trees on the native timber tree Tabebuia rosea in Panama. Timber tree individuals were established in (1) solitary plantings (TSol) or in companion plantings with (2) Guazuma ulmifolia (TGua) or (3) the nitrogen-fixing Gliricidia sepium (TGli).

    Methods

    We quantified growth characteristics and herbivory of T. rosea, and analyzed leaf chemistry (including the stable isotopes ??15N and ??13C) and structure (leaf mass per area).

    Results

    Companion trees significantly affected stem diameter growth of T. rosea. Stem diameter growth was as high in TGli trees as in TSol trees but was reduced in TGua trees. Furthermore, TGua trees had higher leaf water content, and lower ??13C and lower leaf mass-to-area ratio than TGli trees, suggesting there were effects of shading by G. ulmifolia on T. rosea. Herbivory was high but not affected by planting regimes. Leafing phenology did not differ between planting regimes and G. sepium did not increase nitrogen content in T. rosea leaves.

    Conclusion

    Companion tree planting can support timber tree growth in silvopastoral reforestations, but adequate species selection is crucial for successful implementation of this planting system. Tree?Ctree interactions seem to be more relevant for timber tree growth than herbivory in the studied system.  相似文献   

    15.

    Context

    Reliable information on tree stem diameter variation at local spatial scales and on the factors controlling it could potentially lead to improved biomass estimation over pine plantations.

    Aims

    This study addressed the relationship between local topography and tree diameter at breast height (DBH) within two even-aged radiata pine plantation sites in New South Wales, Australia.

    Methods

    A total of 85 plots were established, and 1,302 trees were sampled from the two sites. Airborne light detection and ranging (LiDAR) was used to derive slope and aspect and to link them to each individual tree.

    Results

    The results showed a significant relationship between DBH and local topography factors. At both sites, trees on slopes below 20° and on southerly aspects displayed significantly larger DBHs than trees on steeper slopes and northerly aspects. Older trees with similar heights also exhibited a significant relationship between DBH and aspect factor, where greater DBHs were found on southerly aspects.

    Conclusions

    The observed correlation between tree DBH and LiDAR-derived slope and aspect could contribute to the development of improved biomass estimation approaches in pine plantations. These topographical variables are easily attained with airborne LiDAR, and they could potentially improve DBH predictions in resource inventories (e.g. stand volume or biomass) and support field sampling design.  相似文献   

    16.

    Context

    In N-saturated forests nitrate concentrations in seepage water ( $ {\text{N}}{{\text{O}}_3}{^{ - }_{\text{seepage}}} $ ) regularly show high spatial variability even within homogeneous stands. Up to now the reasons of this variability are not fully understood.

    Aims

    The main objective was to identify the crucial parameters that control spatial variability of $ {\text{N}}{{\text{O}}_3}{^{ - }_{\text{seepage}}} $ at the H?glwald site.

    Methods

    We investigated a multitude of parameters (e.g. N turnover, root biomass, soil chemistry, soil physics, stand parameters) and related them to $ {\text{N}}{{\text{O}}_3}{^{ - }_{\text{seepage}}} $ , measured in 40?cm depth with suction cups.

    Results

    A small number of biological parameters (net N mineralization, root distribution, and stand density) explained up to 93?% of the variability of $ {\text{N}}{{\text{O}}_3}{^{ - }_{\text{seepage}}} $ in linear regression models. Net N-mineralization rates in the humus layer and fine root biomass in the upper mineral soil influenced $ {\text{N}}{{\text{O}}_3}{^{ - }_{\text{seepage}}} $ positively. Fine root biomass in deeper soil layers (30?C40?cm depth) and stand density had a negative influence.

    Conclusion

    The rate of net N mineralization in the organic layer is decisive for the nitrate production in the soil. Roots in the upper mineral soil increase $ {\text{N}}{{\text{O}}_3}{^{ - }_{\text{seepage}}} $ by intensive water uptake but excluding nitrate at the same time. The variation of these two parameters is responsible for most of the small-scale variability of $ {\text{N}}{{\text{O}}_3}{^{ - }_{\text{seepage}}} $ .  相似文献   

    17.
  • ? Empirical observations suggest that in pure even-aged forests, the mean diameter of forest trees (D, diameter at breast height, 1.3 m above ground) tends to remain a constant proportion of stand height (H, average height of the largest trees in a stand) divided by the logarithm of stand density (N, number of trees per hectare): D = β(H ? 1.3)/ ln(N).
  • ? Thinning causes a relatively small and temporary change in the slope β, the magnitude and duration of which depends on the nature of the thinning.
  • ? This relationship may provide a robust predictor of growth in situations where scarce data and resources preclude more sophisticated modelling approaches.
  •   相似文献   

    18.

    ? Context

    Teak??s wood color is considered an important attribute in the marketing phase and it has been influenced by environmental setting, stand conditions and management, plant genetic source, and age. However, there is a lack of understanding about how the environmental factors might affect the teak??s wood color planted in short-rotation forest plantations.

    ? Aims

    The aim of this study is to understand the relationship, gathered from generated information, between edaphic and climatic variables and their effects in the wood color variation of Tectona grandis from trees in forest plantations.

    ? Methods

    Twenty-two plots were grouped in five cluster sites that shared similar climatic and soil conditions. Data about soil??s physical?Cchemical properties and climatic variables were collected and analyzed. Representative trees were harvested next to each plot in order to obtain a wood sample per tree at a diameter breast height. Wood color was measured using standardized CIELab??s chromaticity system.

    ? Results

    After comparing the wood change color index (?E*) in the five studied clusters, it was found that heartwood produced from drier and fertile sites had more yellowish-brown color. The heartwood b* color index resulted with significant correlations (R?>?0.5, P?<?0.05) among nine climatic and eight edaphic variables.

    ? Conclusion

    It was concluded that climatic variables should be considered as the first-order causal variables to explain wood color variation. Hence, darker b* wood color was associated with dry climates; also, with deeper and fertile sites.  相似文献   

    19.
  • ? The physical properties of wood and the associated variations within and between trees were evaluated by analysing 770 small specimens of clear wood from 11 Pinus sylvestris L. trees thinned from 3 plantations.
  • ? Within-tree variations in basic density or volumetric shrinkage increased with cambial age and decreased with increasing ring width. The effect of the height in the stem on wood properties was considered indirect and height was not included as an explanatory variable in the mixed models proposed to estimate basic density and volumetric shrinkage.
  • ? The models had random components for the intercept parameter and explained 52.5% of the total variance in basic density and 56% of the total variance in volumetric shrinkage. Linear shrinkage in the direction of the grain was extremely variable.
  • ? Between-tree variation and between-plot variation in the physical properties of wood were high, considering that all trees sampled were growing in similar sites and stands. It would then be desirable to predict physical properties of wood on living trees in order to use the quality of wood as a criterion for timber tree selection in thinnings.
  •   相似文献   

    20.

    ?Context

    Selective logging followed by natural regeneration is rarely employed for restocking subtropical evergreen broad-leaved forests in East Asia compared with the use of clear-cutting.

    ?Aims

    To clarify the succession of these forests, the effects of selective logging on stand structure, species diversity, and community similarity were studied in a mature and regenerating forest in Okinawa, Japan.

    ?Methods

    Four study plots were established, and trees ≥1.2 m height were identified by species name, tree height, and diameter at breast height.

    ?Results

    The results showed that the species composition of regenerating forest was similar to mature forest; however, the former had a greater species density and Shannon–Wiener index than the latter. Castanopsis sieboldii and Distylium racemosum, the predominant trees in the mature forest, continued to dominate the regenerating forest, with a broad layer distribution. High Sørensen and Jaccard community similarity indices for mature and regenerating forest indicated that the regeneration occurred in a progressive succession.

    ?Conclusion

    The similar species composition and stand structure for both mature and regenerating forest, and the higher species diversity for the latter, provided no evidence of forest degeneration and suggested that the regenerating forest may develop into a stand similar to preselective logging forest.  相似文献   

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