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1.
? Identification of extractives present in Cupressus lusitanica heartwood has been conducted using GC-MS analyses. The chromatogram of toluene/ethanol extracts indicated the presence of large amounts of benzaldehyde and numerous terpenic compounds such as cedrol, agathadiol, epimanool, bornyl acetate, α-cedrene and β-cedrene. ? The effect of these extractives on the natural durability of cypress wood was investigated on heart wood blocks exposed to pure culture of Poria placenta before or after solvent extraction. Weight losses revealed severe fungal degradations on the extracted blocks compared to unextracted ones. ? Efficiency of heartwood extractives as inhibitors of the growth of Poria placenta on malt/agar test confirms their contribution to cypress natural durability. ? Cypress blocks were treated at 240 °C for different times to reach different levels of thermodegradation to evaluate effect of heat treatment on fungal durability. Results indicate that evaporation of volatile extractives during the first few minutes of heat treatment contribute to decreased wood durability, while longer treatment times lead to the expected improvement of durability. ? This study suggests that the content of extractives, which may be modified during wood drying or weathering processes, could be the origin of the conflicting data described in the literature concerning cypress natural durability. 相似文献
2.
? Decay resistance of larch (Larix sp.) to fungi was evaluated on heartwood samples belonging to 3 species (L. decidua, L. kaempferi and their hybrid), 3 races of European larch (polonica, sudetica and alpine), 13 wood lots (populations) and 313 trees. ? Larch wood appeared, on average, as moderately durable although a high variability was observed. At the sample level as well as at the mean individual tree level, durability ranged from class 1 to 5 according to EN 350-1 standard. At the population level, larch wood varied from ‘durable’ to ‘slightly durable’. Genetics played a major role in decay resistance at the species, provenance and tree levels. Environmental factors such as the position of heartwood samples and the age of trees were also identified as a source of variability. ? The most durable wood was not necessarily from old native alpine stands of European larch: some young larches from faster growing lowland origins also produced durable wood. ? Genetic improvement of larch wood durability appeared therefore likely by the selection of the best populations for decay resistance as well as from the selection of individuals. 相似文献
3.
? Tree age is one of the most important factors to affect the natural durability of wood. ? The purpose of this study was to determine the natural termite resistance of heartwood and sapwood of teak (Tectona grandis Linn. fil.) for trees aged 8, 30 and 51 years. Reticulitermes speratus Kolbe was employed as a test termite using a no-choice feeding method. ? The heartwood and sapwood of all of the trees tested exhibited antitermitic activity. Based on the mean mass loss due to termite activity, the sapwood and heartwood regions of 8-year-old trees are the most susceptible to termites while the heartwood regions of the 30-year-old trees showed termite resistance similar to the termite resistance of 51-year-old trees. ? The mass loss is moderately correlated with n-hexane extractive content, total extractive content, brightness and redness of the wood. 相似文献
4.
? The objective of this study was to explore the distribution of major nutrients (N, P, K, Ca and Mg) in the aboveground compartments of an intensively managed tree species (Pinus pinaster Ait.). A total of 53 trees were cut down in even-aged stands respectively 8, 16, 26, 32 and 40 years old. The nutrient concentrations of the aboveground compartments were analysed. ? Nutrient concentrations of foliage did not vary with any of the variables used, except needle age. Nutrient concentrations of living branches, stem bark, stem sapwood, stem heartwood, stemwood and stem decreased with increasing branch diameter, bark thickness, sapwood thickness and heartwood thickness, respectively. Beyond a certain value of the predictive variable (stem diameter ≈ 15 cm; branch diameter ≈ 2.5 cm), the concentration of all the nutrients stabilised. ? A 50 year-old pine stand was used to obtain a validation dataset for nitrogen concentration. For this nutrient, the regression relationships gave satisfactory estimates for most compartments (mean error = 12–25%) and particularly for the stem. ? A procedure is proposed to estimate the nutrient exports associated with harvests of Pinus pinaster biomass. 相似文献
5.
? Wood colour of Tectona grandis produced from fast-growth plantations is highly variable and the causes of this variation are relatively unknown. ? With the purpose of understanding the colour variation, different fast-growth plantations were sampled with different growth rates, tree ages, and sites. ? Wood colour was measured with a CIELab system, where three variables are estimated: coordinate L* for lightness, coordinate a* defines redness and coordinate b* defines yellowness. ? Results showed only a negative correlation between L* and a*. L* and a* were negatively and positively respectively correlated with pith distance in heartwood, but not for b*. No correlations were found between L* and b* in sapwood and plantation characteristics, while a* was positively correlated with age and height of tree and growth rate. In heartwood, tree age and diameter at breast height were correlated with all colour parameters, but tree height and plantation density were correlated with a* and b*. Cluster site had correlation with L*. Multiple correlation analysis showed that the heartwood is increasing darker (L*) and redder (a*) when the trees are older and bigger. Correlation coefficient shown that sapwood and heartwood with lighter colour (L*) is less resistance to fungal attack, but redness colour (a*) increasing decay resistance. 相似文献
6.
? Eperua grandiflora, which is widely distributed in the French Guiana forest region, shows high variability in decay resistance. Further information concerning this wood quality parameter is necessary, but standard testing methods are complex and time-consuming. We assessed the use of colorimetry to determine durability in heartwood samples from a range of trees. ? Eperua grandiflora colour parameters were measured using a CIELAB system, revealing that the tree effect was greater than the radial position and height effects. ? The wood samples were exposed to Coriolus versicolor and Antrodia sp. according to two European standards (En 350-1 and XP CEN TS 15083-1). Eperua grandiflora is more susceptible to brown rot. These two standards did not give the same durability classes. The high variation in natural durability was due to the tree effect. ? These two properties were found to be correlated and the assessment also distinguished the extreme durability classes but they are not sufficient to classify the class of durability of this species. 相似文献
7.
?The study aimed at clarifying basic life traits (fecundity, mortality and development) of Monochamus galloprovincialis, the vector of Bursaphelenchus xylophilus, the causative agent of the Pine Wilt Disease, in Portugal. ? Mating and oviposition experiments were conducted on P. sylvestris logs under laboratory conditions during two years. Larval development was followed outdoors. ? Fecundity was high, with 138.2 eggs per female. ? For the first time, the number of larval instars (4) in M. galloprovincialis was recognized and instars were described. ? Informations are provided on the shape and size of the larval galleries. Boring of galleries in sapwood and heartwood started at the third instar. All instars could overwinter but adults emerged simultaneously resulting in a univoltine life cycle for the majority of the individuals. ? An obligatory diapause in the forth instar was assumed. However, 8.1% of the insects had a two year development. ? Important differences in mortality were noted between the two experimental years, which could be explained by differences on the size of the breeding logs. ? There is a high biological proximity between M. galloprovincialis populations in France and those in Portugal and Northern Europe. However, a higher fecundity and longevity was observed in France, as well as a higher percentage of insects with a two year development, compared with portuguese populations. 相似文献
8.
? Allometry of sapwood/heartwood is essential for understanding tree growth, water transport and carbon allocation, timber production and use, but such an allometry is lacking for Chinese temperate tree species. ? We studied the allometry and development of heartwood and sapwood for seven Chinese temperate tree species: Korean pine (Pinus koraiensis Sieb. et Zucc), Dahurian larch (Larix gmelinii Rupr.), Japanese elm (Ulmus davidiana Planch var. japonica (Rehd.) Nakai), Manchurian ash (Fraxinus mandshurica Rupr.), Manchurian walnut (Juglans mandshurica Maxim.), Amur cork-tree (Phellodendron amurense Rupr.), and Mongolian oak (Quercus mongolica Fisch.). ? All heartwood parameters investigated, including heartwood radius (HR), heartwood formation rate (HFR), heartwood ring number (HRN), heartwood initiation age (HIA), and heartwood volume ratio (HVR), were positively correlated with tree cambial age (CA). The HR, sapwood width (SW), sapwood area (SA), heartwood and sapwood volumes were significantly related to stem diameter at breast height (DBH) or xylem diameter. There was a polynomial relationship between the sapwood ring longevity (SRL) and sapwood ring number (SRN). However, most of the allometric relationships were species-dependent. The hardwood formation patterns were different between coniferous and broadleaved tree species. A power function was suitable to scale SA from DBH, but the exponent varied from 1.32 for the larch to 2.19 for the cork-tree. ? Our allometry provided a practical means to assess wood development and related physiology for the temperate tree species. 相似文献
9.
? Quinones are primarily responsible for the natural durability of teak. ? The previous paper in this series reported on the natural termite resistance of teak trees of different ages (8-, 30- and 51-year-old trees). In this study, the radial distribution of quinones (tectoquinone, lapachol, desoxylapachol and its isomer) and other components in the ethanol-benzene (1:2) extract were measured by means of gas chromatography. ? Significant differences in desoxylapachol or its isomer content were found among the outer heartwood of 8-, 30- and 51-year old trees, as well as between the inner and outer parts of the heartwood. ? All toxic quinone contents were positively correlated with the total extractive content. The highest correlation degree was measured in the isodesoxylapachol content. ? Although linearly related, only modest correlations were observed between the natural termite resistance parameters and the content of tectoquinone and isodesoxylapachol. 相似文献
10.
? Wood used in outdoor conditions out of ground contact is susceptible to weathering, inducing both fungal decay and leaching of components to the environment. ? This paper presents a methodology to determine these two parameters for untreated, preservative-treated and modified wood. Therefore, the wood was first leached and subsequently exposed to fungal decay of the most prominent wood-rotting fungi. The crustacean Daphnia magna was exposed to the leachates to provide information on their impact on the environment. ? Combining both parameters reveals that preservative-treated wood and modified wood are capable of protecting the wood adequately for application under use class 3 conditions without posing a threat to the environment. ? This proves the suitability of the concept of combining efficacy and ecotoxicology for the evaluation of new types of wood treatments. 相似文献
11.
? At present, the production of wood composites mainly relies on the petrochemical-based and formaldehyde-based adhesives such as phenol-formaldehyde (PF) resins and urea-formaldehyde (UF) resins, which are non-renewable and therefore ultimately limited in supply. ? This paper concerns the decay resistance of wood products bonded with a new, environment-friendly adhesive derived from abundant and renewable cornstarch and tannin. To improve the total resistance of the composite against both Coriolus versicolor and Coniophora puteana rot fungi, borax (di-sodium tetraborate) was added in proportions of 0.5%, 1% and 2% (w/w) to the cornstarch-tannin adhesives. ? The results show that increasing the concentration of borax in the adhesive decreased the mechanical properties of the composite. The best way to avoid this problem was to use wood impregnated with borax. ? Biodegradation studies were conducted on new composites, first without any treatment, followed by borax at 0.5% aqueous solution treatment. The results show that wood impregnated with borax, in the presence of tannin and sodium hydroxide in the adhesive improves the total resistance of the wood composite against both Coriolus versicolor and Coniophora puteana rot fungi. 相似文献
12.
? Mycorrhizal fungi and hydrogels (water-absorbing polymers) can improve water availability for trees. The combination of both factors for plant performance under water limitation has not yet been studied. ? To investigate the influence of the ectomycorrhizal fungus Paxillus involutus, hydrogel and the combination of both factors, a drought-sensitive poplar, Populus euphratica, was examined in this study. ? After 16 weeks of inoculation, no ectomycorrhizas were found. Nevertheless, P. involutus-inoculated poplars displayed increased concentrations of soluble sugars and osmolality, leading to an improved water status. Growth was diminished compared with non-inoculated P. euphratica. The presence of hydrogels in the rooting medium resulted in increased biomass and higher plant water content and decreased the osmolality of plant tissues. Drought markedly decreased water contents in rooting medium and plants, and leaf chlorophyll fluorescence, and stimulated the root growth, concentrations of soluble sugars and osmolality in plants. Under drought conditions, P. euphratica exhibited osmoregulation by accumulation of low-molecular-weight carbohydrates. ? These data indicate that adding hydrogels to soils may improve the plant performance. The reasons for improved osmoregulation by fungi and hydrogels were probably related to their stimulating influence on the formation of soluble carbohydrates under drought conditions. 相似文献
13.
? To investigate the effect of climate on radial growth in young plantation grown teak (Tectona grandis L.), growth ring width was measured in 105 trees and correlated to precipitation and temperature data. ? The social status of trees within the stand was also determined and cross-sectional area (CSA) for the trunk correlated to the proportion of heartwood (HW) within the tree. HW develops asymmetrically in leaning stems of some conifer species, but it is not known if this phenomenon also occurs in broadleaf species. Therefore, we measured HW proportion in leaning and straight stems, along with the number of growth rings in the HW. ? Annual ring width depended strongly on mean monthly temperature during the rainy season and the most significant relationships were found corresponding to the months of June and July. With regard to the weaker relationship between precipitation and radial growth, correlations were highest during the period of bud-break at the beginning of the rainy season. ? The very high stand density affected radial growth, particularly in suppressed trees, which responded little to thinning operations. HW formation was greatest in dominant trees, and was highly regressed with stem CSA. ? Therefore, rapid growth of young stands should be encouraged by reducing stand density. Asymmetric HW formation occurred in both leaning and straight trees, and was significantly greater along the upper sides of leaning stems. It is probable that this eccentric HW formation is linked to mechanical loading on the tree. 相似文献
14.
? Non-destructive detection of fungal decay in living trees is relevant for forest management of valuable species, hazard tree assessment, and research in forest pathology. A variety of tomographic methods, based on stress wave timing, radioactive radiation, or electrical resistivity have been used to detect decay in standing trees non-destructively. But apart from mobile gamma ray computed tomography (Habermehl and Ridder, 1993) which is virtually unavailable, the detection of incipient stages of decay is still not possible. ? Wood moisture and electrolyte content influence the electric resistivity of wood. Both are changed by fungal decay. Therefore electric resistivity tomography (ERT) should detect decay in its early stages. Then it could be used to monitor the spatial and temporal progress of degradation. ? We infected four Fraxinus excelsior trees with Trametes versicolor using wooden dowels and measured two-dimensional electric resistivity tomograms 3, 10, 13 and 21 months after infection. Immediately after the last electric resistivity measurement trees were felled for further analyses of stem cross-sections. Wood moisture content and raw density had significantly increased in infected areas, but dry density had not significantly changed after 21 months. Areas of very low electric resistivity around the infected wounds correlated very well with infected wood in the stem cross-sections. Increasing areas of low electric resistivity around the infected wounds during consecutive measurements indicate increasing areas of infected wood. ? We conclude that the growth of white rot by Trametes versicolor can be monitored with electric resistivity tomography (ERT) beginning from incipient stages, even before wood density decreases. ERT could therefore be a powerful research tool for decay dynamics as well as a method for diagnosing wood decay in forestry and arboriculture. 相似文献
15.
? Studies of allometric relationships between leaf area and the cross-sectional area (CSA) of sapwood in the stem have shed light on the structural and functional relationships between water-conducting and photosynthetic tissues. ? The purpose of this study was to test whether sapwood-leaf area relationships could be extended from stems to roots in coast Douglas-fir (Pseudotsuga menziesii var. menziesii (Mirb.) Franco). Twelve trees were felled, their stumps were excavated, and the CSA of sapwood and heartwood were estimated for individual roots, entire root systems, and stem section. ? Root sapwood CSA was greater than sapwood CSA throughout the stem, and the ratio of leaf area to sapwood CSA (A l :A s ) was accordingly lower for root sapwood. The relationship between sapwood CSA and leaf area was more variable in roots and at groundline compared to crown base. Root A l :A s decreased with relative tree height (tree height/mean stand height). ? The strong allometric relationship between leaf area and the CSA of sapwood in the stem generally holds when extended to roots. The greater CSA of sapwood in roots versus stems may reflect differences in their roles in supporting the tree. 相似文献
16.
? It is of importance, when comparing physiological responses of leaves to environmental constraints among different genotypes, to take into account any effect related to leaf position and age within the canopy that might interfere with the response to the constraints. ? To document such effects, photosynthetic capacity and tolerance to heat and to oxidation were measured on leaves from the top to the bottom of three-month-old single-stem rooted cuttings of Populus deltoides × P. nigra genotypes, ‘Dorskamp’ and ‘Luisa_Avanzo’, thus taking into account a gradient of ages from youngest and still expanding (top) to oldest and fully expanded (bottom) leaves. ? All recorded traits were tightly linked to the age of the leaves. Photosynthetic capacity gradually increased during leaf expansion, in parallel with chlorophyll content and relative nitrogen allocation to RuBisCO and to bioenergetics. On the contrary, dark respiration gradually decreased during leaf expansion until a minimum value was reached at maturity. Compared to expanding leaves, young mature leaves were characterized by a lower sensitivity to heat and a higher one to oxidations generated by methyl-viologen. ? Leaf characteristics appeared to vary along the stem to a larger extent than between the two genotypes that display largely different productivities in plantations. 相似文献
17.
? Aspen (Populus tremuloides Michx.) regenerates at high densities following manual cleaning. ? Ten-year-old stands located near Lac La Biche and Peace River, Alberta were manually cleaned to three densities (0, 500 or 1 500 stems ha?1) at three times (bud set, dormancy or bud flush) to test the hypothesis that maintaining residual aspen reduces regeneration. ? At Lac La Biche up to 98% of the aspen regeneration died in the partially-cleaned plots compared to 67% at Peace River five years post-treatment. A spring frost in the second growing season at Lac La Biche is hypothesized to be the inciting factor predisposing the stump sprouts to infection by decay fungi such as Armillaria root rot, resulting in reduced density and height of the aspen regeneration at Lac La Biche relative to Peace River. Drought and ungulate herbivory provided additional stresses. The high mortality of aspen regeneration at Lac La Biche shifted the understory regeneration from aspen to balsam poplar (Populus balsamifera L.). ? These results indicate that maintaining 1 500 stems ha?1 of residual aspen will not effectively control the re-sprouting of aspen; however, the vulnerability of aspen regeneration to spring frost and other stressors can nearly eradicate the re-growth of aspen. 相似文献
18.
? Alterations in plant chemistry underpin a suite of physiological adaptations to arid conditions. Qualitative and quantitative differences in leaf chemistry are found in the genus Eucalyptus correlating with physiological adaptation to aridity. ? Here we investigate seasonal water relations of three field grown eucalypt species grown at a common site known to differ in their ability to accumulate the cyclic sugar alcohol, quercitol. ? We show that quercitol contributes significantly to osmotic relations in field grown trees of Eucalyptus melliodora but is present only in trace amounts in E. rubida and E. obliqua. ? Measured concentrations of quercitol account for the difference in osmotic potentials between species and can be interpreted as a mechanism for adaptation to low water availability. 相似文献
19.
? The effect of climatic variables (temperature and precipitation) on radial growth of the Mediterranean Maritime pine (Pinus pinaster Ait.) was studied using dendrochronological techniques in the Iberian Peninsula. ? Ten tree-ring width chronologies, along the central distribution area of the species, were built. Chronology variability was analysed using Principal Component Analysis (PCA) for the period 1952–2005. ? The first principal component (PC1) explained 56% of tree-growth variability. Tree-growth association with climate was analysed at regional and local scales using correlation coefficient and bootstrapped response functions. ? Radial growth at both scales was positively correlated with rainfall during and prior to the growing season at all sites, and with summer rainfall before the growing season at five sites. Mean temperature effect changed according to the sampling site, from non-significant at the highest sites to significant (positive relationship in winter) at the lowest sites. Growth season temperature also had a negative effect. ? The Kalman filter was applied to estimate changing association between growth and climate over-time. Results suggested a change in association, initiated in the 80s, from non-significant to significant (*p < 0.05) at six of the sampling sites. ? Pinus pinaster is an accurate species for analysing tree-growth association with climate and for studying plant behaviour under global change conditions. 相似文献
20.
? Artificial defoliation is often used to simulate defoliation by herbivory and is usually considered a good indication of a plant’s response to a given type of damage. However, the findings of studies directly comparing the two defoliation types are inconsistent. ? Here, the short term effects of artificial and insect defoliation by larvae of Paropsisterna agricola on growth, biomass allocation and photosynthetic capacity of Eucalyptus globulus seedlings were compared in a glasshouse experiment. The artificial defoliation was carried out to closely resemble the spatial patterns observed for insect defoliation. ? Height and diameter increments were reduced as a result of insect defoliation, whereas artificial defoliation had no significant effect on height. Increased photosynthetic capacity was observed in response to both treatments, but the magnitude of this increase was larger in insect-than in artificially-defoliated seedlings. Significant reductions in foliar carbohydrate content and total biomass were noticeable in artificially-defoliated seedlings. Although the foliar carbohydrate levels also decreased across the crown zones following insect defoliation treatment, seedlings allocated a large amount of their biomass in the branches of the damaged zone. ? Despite our best endeavours to simulate insect defoliation in the artificial treatment, the latter may not reflect accurately the full strength of the effects. However, artificial and insect defoliation were similar in their direction of the responses they caused in E. globulus seedlings. 相似文献
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