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1.
Management of subterranean clover (Trifolium subterraneum L.) understory grown with loblolly pine (Pinus taeda L.) and slash pine (Pinus elliottii Engelm.) resulted in differences in both clover yield and pine diameter. Loblolly and slash pines were planted into a subterranean clover pasture in 1984. Suppression of warm season herbaceous vegetation by applying herbicides or disking the site in late summer resulted in significantly greater subterranean clover production than on the control site where no additional understory management was applied. Pine species had no effects on clover yield. The N content of the pine foliage on the blocks managed with herbicide application or disking generally was significantly greater than the control from the 5th through 7th growing seasons (1988–1990). The dbh of the pine trees in the disked and herbicide treatments was greater than the control after the 7th growing season (1990).  相似文献   

2.
When combining pines and cattle on tame pasture, grazing is often delayed for several years until trees are large enough to resist injury. As an alternative approach to delayed or deferred grazing during the early years, this study in central Louisiana, USA, examined the effects of cattle grazing in subterranean clover (Trifolium subterraneum L.) pastures on slash (Pinus elliottii Engelm.) and loblolly pine (P. taeda L.) seeding survival and growth during the first 3 years of tree establishment. Pines were planted at about 1200 trees/ha in 3 rows on 0.4-ha subclover units with 1.3 m spacing within rows. Three grazing treatments included: (1) ungrazed pines, (2) limited grazing with a single-wire electric fence above the planted pines, and (3) grazed pines. Thirty Brahman crossbred cows with calves and a bull grazed the tame pasture on a controlled grazing, rotational basis from December through May each year, during the subterranean clover growing season. Pine trampling injury during the year was 8% on the grazed seedlings while essentially none occured under limited grazing. During the first 2 growing seasons, survival and height of the pines were significantly less on the grazed seedlings than on either the limited grazing or ungrazed seedlings. Pine heights from the limited grazing and ungrazed treatments were not different during the 3-year study; loblolly pine heights from the limited grazing treatment continued to be taller than the grazed treatment through the third year while the slash pine heights were similar for all treatments by the third year. Seedling mortality became more acute as severity of grazing injury increased; the greatest mortality occured when the terminal bud and needles were both browsed off.  相似文献   

3.
Successful establishment and growth of tree seedlings in an agroforestry practice is dependent on the control of competing herbaceous vegetation. Conventional weed control methods (i.e., chemical, mechanical, and physical suppression) are effective but can be costly in terms of time, damage to non-target vegetation, or increased soil erosion. Alternatively, some living mulches can exclude undesirable vegetation, protect the soil, compete minimally with associated trees, and supplement soil nitrogen. In this study, small and large white clover (Trifolium repens L.), red clover (T. pratense L.), kura clover (T. ambiguum Bieb.), strawberry clover (T. fragiferum L.), birdsfoot trefoil (Lotus corniculatus L.), hairy vetch and ’AU Early‘ hairy vetch (Vicia villosa Roth) were selected for evaluation as living mulches. These species were established in 9 m ×9 m replicated plots along with two controls [tall fescue (Festuca arundinacea Schreb.) and bare soil kept vegetation free using glyphosate]. The controls represent ’high competition‘ and ’no competition‘, respectively. Eastern black walnut (Juglans nigra L.), honeylocust (Gleditsia triacanthos L.), and hybrid pitch × loblolly pine (Pinus rigida L. × P. taeda L.) seedlings were planted in association with each living mulch. Tree seedling heights and diameters were measured during the first two years and forage performance evaluated. All seedlings grew better in the absence of competition. Tall fescue greatly reduced the growth of hardwood seedlings. Red clover and ladino clover showed the greatest promise as living mulches for use with the hardwood and pine seedlings, respectively. This revised version was published online in June 2006 with corrections to the Cover Date.  相似文献   

4.
Although gravitropism of maize (Zea mays L.) roots is dependent on light, it is not known if light affects the gravitropism of pine radicles. Therefore, seeds of slash pine (Pinus elliottii Englm.) and loblolly pine (Pinus taeda L.) were germinated under constant light or constant darkness. A gravitropic set-point angle (GSA) of 90° indicates the radicle is pointed horizontally while a GSA of 0° indicates the radicle is growing toward the center of the earth. After 20 days from sowing, the GSA was less than 10° in both light and darkness. Therefore, gravitropism of both pine species occurred in the dark. The presence of light initially stimulated gravitropism but this effect was ephemeral. After 25 days from sowing, there was no significant difference in radicle growth among light treatments, suggesting that darkness did not affect pine radicle gravitropism. However, light did increase germination of both species. Radicle length was slightly increased when seeds germinated in the dark.  相似文献   

5.
Abstract

The effects of fencing and site preparation on performance of birch (Betula pendula Roth), oak (Quercus robur L.), pine (Pinus sylvestris L.) and spruce [Picea abies (L.) Karst.] seedlings were studied over four growing seasons in southern Sweden. The experiment was composed of four browsing treatments, from which roe deer (Capreolus capreolus L.) were excluded for 0, 12, 24 or 42 months, and four soil treatments. Natural browsing was combined with artificial browsing (clipping). It was demonstrated that browsing by roe deer could be isolated from browsing by moose (Alces alces L.) by selective fencing. Except for birch, browsing had little effect on seedling survival. Browsing reduced height growth by more than 100% for oak and pine, and more than 60% of pine seedling developed multiple stems. Except for oak, site preparation increased seedling survival. Inverting site preparation, in combination with or without fertilization, produced positive growth responses in seedlings, whereas patch scarification did not. There was no interaction between browsing and soil treatments on seedling performance. It was concluded that long-term protection against browsing by roe deer is needed for oak and pine, whereas short-term fences might be used for birch, and that spruce is relatively unaffected by browsing.  相似文献   

6.
A study was initiated in 1994 to evaluate the effects of bedding timing and frequency and pre-plant herbicide application, with and without post-plant herbaceous weed control, on growth of three slash pine (Pinus elliottii Engelm.) plantations and one loblolly pine (Pinus taeda L.) plantation on flatwoods sites. Site preparation treatments included early bedding alone, late bedding alone, double bedding, early bedding plus banded pre-plant herbicide, and early bedding plus broadcast pre-plant herbicide. Each site preparation treatment was tested with and without a first-year post-plant herbicide application for herbaceous weed control (HC). Results ranging from age 5 to 11 are reported, depending on installation. Pine growth following late bedding alone was similar or greater than that with early bedding alone. Double bedding did not consistently increase pine growth as compared with single bedding alone. Early bedding plus either broadcast or banded pre-plant herbicide application generally exhibited greater pine growth than did bedding alone treatments or bedding plus post-plant herbaceous weed control treatments. Broadcast pre-plant herbicide application resulted in similar or greater growth than banded pre-plant herbicide application. Post-plant herbaceous weed control improved loblolly pine growth across all site preparation regimes on the one test location. For slash pine plantations evaluated, post-plant herbaceous weed control resulted in significant growth increases on some but not all sites. The differences in growth response to bedding and pre- and post-plant herbicide applications are discussed in relation to treatment efficacy in controlling woody shrub and herbaceous competing vegetation. Effective bedding and chemical site preparation reduces the need for post-plant herbicide applications for slash pine on certain sites. Good site preparation, including the application of herbicides for competition control, is essential for fast growing slash and loblolly pine plantations in the flatwoods.  相似文献   

7.
This study reports 14th-year response of a boreal mixedwood stand to different harvest intensities (uncut, 50% partial cut with and without removal of residuals after 3 years, and clearcut), spot site preparation treatments (none and scalped), and chemical weeding frequencies (none, single, and multiple) in northeastern Ontario. The response variables include the survival and growth of planted white spruce (Picea glauca [Moench] Voss) and jack pine (Pinus banksiana Lamb.), height and density of natural regeneration and shrubs, and cover of shrubs and non-woody vegetation. Harvesting and weeding generally improved survival and growth of planted trees, although white spruce survival did not significantly differ among the three weeding frequencies. Harvesting tended to increase heights of hardwood (mostly trembling aspen (Populus tremuloides Michx.)) and conifer (largely balsam fir (Abies balsamea (L.) Mill.).) natural regeneration, cover and density of shrubs, and cover of herbs, lichens, and ferns. Chemical weeding reduced height, density and cover of shrubs, height and density of hardwood regeneration, and fern cover, but increased moss and lichen cover. Spot scalping did not significantly affect planted seedling, natural regeneration, or the vegetation.Maximum survival and growth of planted white spruce and jack pine were achieved using a combination of clearcutting and multiple weeding. However, partial cutting followed by a single weeding produced acceptable survival and reasonable growth of planted trees, particularly for white spruce. Partial canopy removal alone substantially reduced the amount of hardwood regeneration, relative to clearcutting, but did not adequately suppress understory shrubs. Significant improvement in seedling growth following multiple weedings was evident primarily in the complete canopy removal treatments: 50% partial cut with removal of residuals after 3 years and clearcut. While the effects of harvesting and weeding on planted crop trees found in the 5th-year assessments generally persisted at year 14, survival decreased, likely due to light competition from developing hardwood and shrubs.  相似文献   

8.
Pedunculate oak (Quercus robur L.) and Scots pine (Pinus sylvestris L.) seedlings were lifted on several occasions during autumn 1997 to determine the relationships between storability and frost hardiness. On each lifting date their physiological status was determined by assessment of shoot and root electrolyte leakage and frost hardiness, assessed as freeze-induced electrolyte leakage. Additional seedlings were simultaneously cold-stored for field planting and assessment of preplanting root growth potential in April 1998. First year field performance was determined the following winter. Storability and cold acclimation patterns differed between the two species. Both were negatively affected by early lifting, but oak was less sensitive with respect to survival, and pine attained tolerance to cold storage more rapidly and earlier with respect to growth increment. The correlations between shoot frost hardiness and performance suggest that freeze-induced shoot electrolyte leakage (SELdiff?20) below a threshold of 5% is a good storability predictor for Scots pine in Denmark. A completely reliable criterion for pedunculate oak could not be established.  相似文献   

9.
Two annual crops of container-grown loblolly pine (Pinus taeda L.) seedlings raised outdoors differed from those grown inside a covered chamber with a clear top. After a 90-day growth period, outside-grown container seedlings had shorter heights, smaller root collar diameters, and lower root, stem, foliage, shoot, and total dry weights than chamber-grown seedlings. Fall-planted, outside-grown seedlings raised the first year had up to 10% higher field survival than those from the chamber. However, chamber-grown container seedlings that survived retained their initial height and diameter advantages following one complete growing season. Fall-planted seedlings raised the second year showed no differences due to growth treatments when favorable weather for seedling growth occurred after field-planting. Container-grown seedlings raised outdoors during the summer months are well suited for fall-outplanting.  相似文献   

10.
Black walnut (Juglans nigra L.) produces both valuable lumber and a nut crop. Because of this, it is an important tree for agroforestry plantings in the Midwest USA. However, during processing of the nut crop, the outer tissue of the nut (husks) accumulates in great quantities. Applying this material to pastures is a possible method of disposal. However, black walnut has been implicated in allelopathic interactions with numerous plant species and may inhibit the growth of pasture species. Greenhouse and field studies and a chemical analysis of the husks were conducted to determine the effects of applying walnut husks to orchardgrass (Dactylis glomerata L.) or red clover (Trifolium pratense L.) pastures. Applying husks at rates up to 68 Mg ha−1 to pot-grown orchardgrass resulted in root and shoot yields that were greater than in pots receiving no husks. Field studies indicated that orchardgrass and red clover benefited from the application of husks. For orchardgrass, significant increases in growth occurred when husks were applied at a rate of 34 Mg ha−1 while red clover responded to rates up to 68 Mg ha−1. A chemical analysis of husks indicated that significant quantities of N and K were present in walnut husks. A whole effluent toxicity test (WETT) indicates that runoff from walnut husk-treated pastures, at the rates used in this study, would not adversely affect aquatic organisms.  相似文献   

11.
Soil moisture content from 0 to 2 m depth was monitored under 2–6 year old radiata pine (Pinus radiata) with three understoreys of bare ground, lucerne (Medicago sativa) and ryegrass/clover (Lolium perenne/Trifolium spp.) and under adjacent open-grown lucerne and ryegrass/clover pastures. By the fifth year soil moisture depletion/recharge pattern under the trees alone was similar to that under open pasture and under trees with pasture understoreys. Maximum plant available moisture storage was 207–223 mm in the top meter of this Templeton silt loam soil but only 69–104 mm at 1–2 m depth where coarse textures often predominated. Lucerne reduced soil moisture content (SMC) to lower levels during drier summers and extracted more water from 1 to 2 m depth than ryegrass/clover. Evapotranspiration (ET) during early summer when soil moisture was high was close to the Penman potential evapotranspiration (E p ), but the difference increased when SMC in the top meter dropped below 200 mm. The silvopasture treatments had higher ET in winter than pasture alone but this was still less than E p . Soil moisture deficits (SMD) at the end of each summer were sufficiently large to require slightly higher than normal winter rainfall and ET < E p to recharge the soil to field capacity before the next summer. The soil moisture results, taken together with root and growth data, suggest that trees and understorey pastures are complementary in the first three or four growing seasons but this balance subsequently declines in favor of the pine trees. Management options, to extend the period that understorey pastures are productive, include reducing tree stockings, more vigorous pruning, using competitive understoreys and changing from pines to deciduous trees. Research on new silvopastoral combinations is suggested.  相似文献   

12.
Correlations between root growth capacity (RGC), at the time of planting, and field performance were studied for Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) seedlings. Before planting a gradient in seedling viability was generated through exposure to low root temperatures and different winter storage regimes. The hypothesis that high RGC values would improve field performance was to some extent verified for pine seedlings while no correlations could be registered for spruce. Reasons for these results are discussed.  相似文献   

13.
Abstract

Tree vitality and height of lodgepole pine (Pinus contorta var. latifolia) originating from the six Swedish seed orchards and native Scots pine (Pinus sylvestris L.) were estimated in three 8-yr-old test plantations established in the Komi Republic (north-west Russia). A randomized row-plot design with 9–13 replicates of each entry was used. In general, the Scots pine tree vitality was better than that of lodgepole pine but these differences were significant only for seed sources of southern origins Larslund, Rumhult, and Österby. On the contrary, the lodgepole pine height growth was faster (6–21%) except for seed sources of southern origins Rumhult and Österby. In comparison with the native Scots pine, lower mean heights (4–10%) were recorded for these seed sources. Among the lodgepole pine seed sources the northern origins had better vitality, while the effect of latitude on the tree height was insignificant.  相似文献   

14.
The objective of this study was to evaluate how competition would affect the physiology, and thus productivity of American beautyberry (Callicarpa americana L.), wax myrtle [Morella cerifera (L.) Small] and inkberry [Ilex glabra (L.) A. Gray] when intercropped in a longleaf pine (Pinus palustris Mill.) plantation in the southeastern United States. The effect of competition was assessed via comparisons of mortality, biomass, light transmittance, gas exchange and soil moisture between intercropping and monoculture (treeless) treatments. Overall, shrubs in the intercropping treatment performed worse than those in the monoculture, with higher mortality, and reductions in biomass of 75.5, 50.6, and 68.7% for C. americana, M. cerifera and I. glabra, respectively. Root–shoot ratios for all species were significantly higher and soil moisture during dry periods was significantly lower in the intercropping treatment. Light transmittance below the pine canopy was high (57.7%) and I. glabra was the only species that exhibited reduced A max when belowground resources were not limiting. These results suggest that the effect of shading is minimal and belowground competition is likely the most important determinant of productivity in this system.  相似文献   

15.
The patterns of current‐year shoot, needle and terminal bud elongation in seedlings of three Scots pine (Pinus sylvestris L.) and three lodgepole pine (Pinus contorta Dougl. var. latifolia Engelm.) provenances were compared during the third and fourth growing seasons after planting. Lodgepole pine produced longer shoots and buds than did Scots pine, mainly because lodgepole pine formed more stem units and elongated at a faster rate. Stem unit length and the duration of shoot and bud elongation differed relatively little between species and provenances. Lammas or polycyclic growth occurred in some lodgepole pine provenances, but not in any Scots pine provenance, and was associated with enhanced shoot elongation. Needle elongation commenced earlier, proceeded at a faster rate, and was greater in lodgepole pine than in Scots pine, but ceased about the same time in all species and provenances. The heat sum required to attain 50% of final length was lower for shoots and needles in lodgepole pine than in Scots pine, and for shoots in northern provenances than in southern ones. Mitotic activity in the apical meristem of the terminal bud, which occurred less than one week after the seedlings were free from snow, started and ceased about the same time in each species, but was higher in lodgepole pine than in Scots pine early in the shoot elongation period.  相似文献   

16.
The success of various grass-legume mixtures in controlling competing vegetation, and their effect on subsequent survival and growth of Sitka spruce seedlings was studied on a coastal alluvial site in northwestern British Columbia. Mechanically scarified (bladed) strips were hand seeded to pure and combined mixtures of legumes, bunchgrasses, and sodforming grasses. An unseeded control (bladed but not seeded) was also established.Alsike clover (Trifolium hybridum L.) was the most successful legume species. Orchardgrass (Dactylis glomerata L.), big bluegrass (Poa ampla Merr.), and creeping red fescue (Festuca rubra L.) were the most successful grass species. Red alder (Alnus rubra Bong.) density and height were lower in legume-seeded treatments. Red elderberry (Sambucus racemosa L.) and salmonberry (Rubus spectabilis Pursh.) were effectively reduced by blading and have reestablished slowly. A combination of blading and early establishment of either sodforming grasses or bunchgrasses effectively decreased reinvasion by thimbleberry (Rubus parviflorus Nutt.).Growth of Sitka spruce was best in the unseeded control treatment or in the treatments with legumes but no sodforming grasses. Presence of sodforming grasses decreased both diameter and height growth. Sitka spruce diameter decreased with increasing red alder density. After 5 growing seasons, spruce has outgrown all competitors except red alder.  相似文献   

17.
The chilling requirement of rest completion and the high temperature requirement of growth initiation were determined in three origins of silver birch (Betula pendula Roth.) seedlings and five origins of Scots pine (Pinus sylvestris L.) seedlings. In both the pine and birch the chilling requirement was highest in maritime Scottish origins and lowest in the most continental Finnish and Russian origins. The requirement for southern mountainous Spanish and Bulgarian pine origins was in between. In terms of the high temperature requirement, there were no clear differences between origins. These results suggest that owing to their high chilling requirement, which prevents the beginning of growth and the loss of frost hardiness during the frost‐exposed season, origins from a maritime climate could be the most tolerant under climatic warming.  相似文献   

18.
The integration of forage crops in an alley-cropped system was examined as a method of encouraging tree planting to increase farm income, improve soil quality, and enhance biodiversity on Midwestern U.S. farms. Crop and tree performance were evaluated in an alley-cropped system using four forage intercrops grown in tree alleyways to simulate a potential hay crop – oat (Avena sativa L.) and red clover (Trifolium pratense L.); oat, red clover, and red fescue (Festuca rubra L.); oat, red clover, and orchardgrass (Dactylis glomerata L.); and oat and hairy vetch (Vicia villosa Roth) – compared to a herbicide, mowing and control (no management) treatment. Five tree species, divided into fast-growing hardwoods of two poplar (Populus spp.) clones [Crandon (P. alba L. × P. grandidentata Michx.) and Eugenei (P. deltoids Bartr. × P. nigra L.)], and silver maple (Acer saccharinum L.) were compared with two high-value, slow-growing hardwoods planted from seed and as seedlings: red oak (Quercus rubra L.) and black walnut (Juglans nigra L.). Tree survival in the first year was greater for the fast-growing species, and by the fourth year, tree height among the four forage treatments was equivalent for all upland locations. The oat/hairy vetch ground cover was associated with the shortest trees in the bottomland site. Herbicide-treated trees were taller than the average of the four forage treatments for all four planting conditions. Tree height in the mowing and the control treatments was not significantly different for all four planting conditions. The nutritive value of the forage crop was excellent in the second year of tree establishment, with crude protein content and digestibility at 17 and 71%, respectively, in the oat/red clover/red fescue treatment, suggesting the viability of forage crops as alternatives to herbicides in alley-cropped systems.  相似文献   

19.
Two species of pine native to Oregon and three native to Mexico were reciprocally planted in southwest Oregon and northeastern Mexico. The Oregon study site has very dry, hot summers; the Mexican site has hot, dry springs, but has rainy summers. Total rainfall and native genera are similar on both sites. Site-preparation trials were carried out on north and south slopes at both locations. For all species on both sites, survival on south slopes was satisfactory only with site-preparation treatments that gave near-complete vegetation control. Overall survival was best on north slopes in Mexico, and herb control was not required. Woody cover decreased survival and growth at all locations and aspect combinations. Chemical control improved survival, but the standing dead shade did reduce growth slightly as compared to growth on the totally cleared sites. Clearing without chemical control gave poor results in Oregon, but permitted good seedling survival and growth at the location in northeastern Mexico. Overall, ponderosa pine (Pinus ponderosa Dougl. ex Laws.) was the least sensitive species to environmental stress, and Hartweg pine (Pinus hartwegii Engelm.), the most sensitive. Mexican pines appeared to require more protection from water stress than did ponderosa pine.  相似文献   

20.
Abstract

Above- and below-ground responses of juvenile loblolly pine (Pinus taedaL.) and white oak (Quercus albaL.) to tropospheric ozone (0.0, 0.06, and 0.12 μL L-1 O3), acid precipitation (pH 4.8, 4.2, and 3.6), and induced ectomycorrhizal colonization (Pisolithus tinctorius [Pers.] Coker & Couch vs. natural mycobiont), alone and in combination, were examined. After 30 weeks, the filtered air treatment (ozone 0.0) reduced foliar injury but also produced smaller stem diameters, lower total foliage and total shoot dry weights, decreased needle density, and shorter root systems in inoculated pine, growth responses not found in uninoculated pine. Height growth of inoculated oak was increased by the filtered air treatment while leaf density was reduced by 0.12 μL L-1 O3, results also not found in uninoculated oak. Ectomy-corrhizal colonization percentages in pine decreased with increasing O3 concentration regardless of mycorrhizal treatment, and the filtered air treatment produced the highest infection percentages in inoculated oak as well. Rains of pH 4.2 stimulated height growth in inoculated pine while diameter decreased in inoculated oak but increased in uninoculat-ed oak with declining pH. Also, pH 4.2 rainfall increased root weight in inoculated oak while that of pH 4.8 reduced foliar injury but also shoot weight in oak regardless of mycorrhizal treatment. Generally, the my-corrhization response to increasing rainfall acidity was reduced colonization. Independent of the other treatments, inoculation produced a substantial increase in above- and below-ground growth of both pine and oak, and also increased foliage density but reduced specific root length in each species. Overall, these results indicate that the growth responses to O3 and acid deposition of loblolly pine and white oak are strongly influenced by mycorrhization, but that the magnitude of these responses is exceeded by those to mycorrhization alone. However, the propensity revealed here for both O3 and acid precipitation to suppress mycorrhizal colonization in these two species may indicate the potential for a gradual reduction in long-term forest productivity in the southeastern United States.  相似文献   

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