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1.
Inoculation experiments were performed in order to evaluate the virulence of Gremmeniella abietina isolates from Spain on the main pine species planted in the Iberian Peninsula, as well as the influence of seedling age on this virulence. Two different experiments were carried out with four isolates of G. abietina from Spain. The greenhouse experiments consisted of seedling inoculations. One‐ and 2‐year‐old seedlings of the following five pine species were used: Pinus halepensis, Pinus pinea, Pinus pinaster, Pinus sylvestris and Pinus uncinata; also, 1‐year‐old seedlings of P. nigra were inoculated. The relative necrosis length (RNL) caused by the pathogen after 130 days was used as a response variable. The laboratory experiments were performed on 2‐ to 6‐year‐old internodes of the above pine species excluding P. uncinata. The necrosis length after 6 weeks of incubation was measured. The results have shown that all G. abietina isolates were pathogens on seedlings of these six pine species and seedlings of P. halepensis were consistently the most susceptible ones, although it is important to take into account that all the isolates used in the present work were isolated from P. halepensis, the only pine species in Spain where G. abietina has been recovered up to now. The susceptibility of the other pine species depended on the age of the seedlings.  相似文献   

2.
Pine resistance to low- to moderate-intensity fire arises from traits (namely related to tissue insulation from heat) that enable tree survival. Predictive models of the likelihood of tree mortality after fire are quite valuable to assist decision-making after wildfire and to plan prescribed burning. Data and models pertaining to the survival of European pines following fire are reviewed. The type and quality of the current information on fire resistance of the various European species is quite variable. Data from low-intensity fire experiments or regimes is comparatively abundant for Pinus pinaster and Pinus sylvestris, while tree survival after wildfire has been modelled for Pinus pinea and Pinus halepensis. P. pinaster and P. pinea, and Pinus canariensis in special, are better equipped to survive fire, but low-intensity fire is tolerated even by species often referred to as fire-sensitive (P. halepensis and Pinus radiata). The relative fire resistance of European pine species is assessed on the basis of (i) morphological and experimental data, and (ii) mortality modelling that considers fire behaviour. Limitations of these approaches to rate fire resistance are discussed, and the current knowledge gaps are indicated.  相似文献   

3.
Forests cover almost 30% of the Mediterranean region today, yet forest management activities have influenced structure and composition of both natural and planted forests. To date no study has been conducted to evaluate the impact of forest management on saproxylic beetle assemblages, although it is known that the Mediterranean is a biodiversity hotspot with a long-lasting human pressure on natural habitats. We provide an overview of saproxylic beetle assemblages of three forest types (mature Pinus halepensis forests, mature Pinus brutia forests, young Quercus calliprinos forests) in the East Mediterranean region using a one-year sample from 12 forest plots located in the north of Israel.  相似文献   

4.
Studying species interactions in mixed forests allows us to assess their potential benefits and adapt current silvicultural tools developed in monospecific stands to multi-specific stands. We analyzed tree interactions in a Pinus halepensis Mill. and Pinus pinea L. mixed plantation using individual tree neighborhood models and competition indices that accounted for symmetric and asymmetric competition, to analyze whether the growth of each species was better explained by symmetric or asymmetric competition. We also split the competition indices into their intra- and interspecific forms, to test for competition effects on growth change based on competitor identity. Finally, we analyzed whether P. halepensis and P. pinea trees had different growth responses to competition. When calculating competition indices, we explored how spatial information and size of competitor trees contributed to the quantification of the process. Competition measurements were optimized to more precisely describe interactions. Results showed that the inclusion of competition indices generated important improvements in growth models. The main mode of competition was symmetric, which could be related to water restrictions typical of the Mediterranean climate. Considering competitor identity did not improve the growth models, while measurement without discriminating competitors by species generated more parsimonious models. P. halepensis and P. pinea trees had similar growth responses to competition, indicating that the two species cope with competition in similar ways. However, P. pinea showed lower average growth than P. halepensis in the period analyzed. Results suggest that preventing the onset of intense interspecific competition processes could help slow down the long-term replacement of P. pinea by P. halepensis and could have benefits for silvicultural management in systems with two species that share ecological niches but are capable of generating different goods and services.  相似文献   

5.
Pedraza  R.A.  Williams-Linera  G. 《New Forests》2003,26(1):83-99
Four native tree species (Liquidambar styraciflua, Juglans pyriformis, Podocarpus matudae, and Carpinus caroliniana) were evaluated for their suitability in rehabilitating degraded areas of Mexican cloud forest. Plant survival and growth in height and diameter were determined in three mixed-experimental plantations with different land use histories; their performance was compared with two on-farm plantations started by landowners for forest restoration. Nearby forest fragments were controls for soil compaction. An experimental plantation with remnant trees had the highest plant survival (82%), height, basal diameter and relative growth rate. The plantation with the steepest slope had high plant survival (63%) and growth. The plantation characterized by dominance of grasses and compacted soils had the lowest survival (22%) and growth. On-farm plantations had good establishment of planted trees (5–10 species planted), and facilitated the recruitment of 9–11 woody species. Carpinus and Liquidambar appear to be suitable species for reforestation in all these areas. Podocarpus grew relatively slowly, although it performed well in two experimental sites. Juglans had high survival (76%) under the stressful conditions of the most adverse site, and therefore may be useful for rehabilitation of degraded sites. Differences among species and sites strongly suggest that species success depends on plantation site quality.  相似文献   

6.
Reforestation is one of the most important and efficient measures of water and soil conservation. Based on field investigations in the Shangyang Soil Conservation and Reforestation Station in Huizhou, Guangdong Province, China, we studied the variation in vegetation development, vegetation succession processes and soil erosion. The regional vegetation consists mainly of monsoon evergreen broad-leaved forests (MEBF). The area was deforested and became a denuded hill area with extremely high soil erosion in the 1960s and 1970s. Then, the area was closed in order to allow recovery of the vegetation. Under natural conditions the vegetation development and succession processes were slow during which soil erosion and strong sunshine and evaporation slowed down the development of the vegetation. About 25 years later, the vegetation cover was still merely 35% or so. The dominant vegetation types were heliophilous herbage and shrubs which formed a poorly developed shrub-herbage community and erosion remained high. In contrast, reforestation with selected tree species dramatically speeded up the vegetation succession process. About 12 years after reforestation, vegetation cover of the Acacia auriculiformis plantations in the Shangyang Station was 90% and erosion was under control. After 23 years, understory vegetation, consisting of indigenous species, had developed in the plantations. The planted trees and naturally developing herbage, shrubs, bamboo, local trees and liana formed a complex vegetation community in three layers. It will take 60 years for the vegetation to succeed from bare land to a secondary growth forest under natural conditions. Reforestation may speed up the vegetation succession process. The time may be reduced to 20 years. Reforestation is the most effective measure of vegetation restoration and erosion control in this area. __________ Translated from Acta Ecologica Sinica, 2006, 26(8): 2558–2565 [译自: 生态学报]  相似文献   

7.
Knowledge of the germination characteristics of Pinus species can help in the understanding, prediction and management of the regeneration of pine forests. In the Canary Islands, several exotic pines (Pinus halepensis L. and Pinus pinea L.) were planted with Pinus canariensis Chr. Sm. Ex DC, the only native pine species, and there is now an interest in controlling these exotic species to restore the original forest. The main objective of the present study is to determine the germination response of P. canariensis, P. halepensis and P. pinea to different light regimes (darkness and light/darkness), thermal shocks and the presence of inhibitory substances from the leaves and litter of P. canariensis. P. halepensis seeds showed the highest viability and germination rate. Darkness accelerated P. halepensis germination, while exudates accelerated P. canariensis germination. Only treatments of 200 °C for 5 min and 300 °C for 5 min significantly decreased the germination of all three pine species. In the absence of strong differences in germination among species after treatments, the key for fire management or prescribed burning in this case may be the sprouting ability of P. canariensis. A possible management strategy to control the two obligate seeder exotic species could be to use one intense prescribed fire followed by a low-intensity prescribed fire after seed bank germination in the field, so in this way, only P. canariensis could resprout after fire and it would eliminate only the exotic pines in a mixed stand.  相似文献   

8.
We investigated central Mediterranean Pinus halepensis plantations under semi-arid climate in order to evaluate the combined effect of soil treatment and afforestation practices on spontaneous plant species composition,richness and evenness,and on the trend and speed of vegetation dynamics.Phytosociological relevés of three different plot typologies,i.e.(1) soil-treatment and plantation,(2) only soil-treatment,(3) no soil-treatment and no plantation,were compared by(a) multivariate analysis and(b) with reference to species richness and evenness.Moreover,in order to compare vegetation dynamics within the plantations with those ones ongoing in semi-natural garrigue communities,we compared life form and syntaxonomic spectra between phytosociological relevés taken at 8 years of distance.DCA showed that floristic species composition and similarity are influenced by the canopy cover of Pine trees as well as by soil-treatment practices.Although species richness and evenness are not clearly related to neither soil treatment nor Pine afforestation,canopy cover clearly plays a major role:in fact,the highest Ph cover rates correspond to the lowest values of understory species richness.This is true also if only species of biogeographical/conservation interest are considered.Regarding vegetation dynamics,sites with dense Pine canopy cover evolve much slower than the adjacent garrigue communities.The same factors invoqued to explain the patterns of floristic composition and similarity(i.e.allelopathy and competition for light,water and nutrients) may also explain the lowering of diversity of therophytes and the strong decline of the cover perfomed by both therophytes and hemicryptophytes underneath the canopy of dense Ph plantations.Thus,in sites where Ph cover exceeds about 80%,thinning is recommended not only in order to accelerate succession,but also to give a natural ’shape’ to afforestations.  相似文献   

9.
Drought, salinity and forest fires are adverse abiotic factors responsible for the failure of plant regeneration in the Mediterranean regions. The objective of this study was to evaluate Pinus pinea seed germination in response to heat, osmotic and salinity stress. Separate experiments were conducted, using polyethylene glycol (PEG) as an osmotic agent to reproduce drought stress, and sodium chloride (NaCl) and diluted seawater as salinizing agents to simulate salinity stress. Different temperatures were used to reproduce the effects of different fire regimes. Seed germination and the activities of the main enzymes involved in Pinus seed reserve utilization (glyoxylate cycle enzymes) decreased with increasing PEG, NaCl and seawater concentrations. Seawater appears to be the least toxic salinizing agent on Pinus pinea seed germination. The present study indicates that the germination of Pinus pinea seeds is influenced by the concentrations and even more by the nature and interactions of the ions present in the solutions. As regard heat treatments, the maximum germination percentage (80%) was obtained at 80°C and short exposure time (3 min). Increasing exposure time and temperature led to the germination percentage progressively decreasing.  相似文献   

10.
Pitch canker, caused by Fusarium circinatum, and Diplodia shoot blight, caused by Diplodia pinea, are both damaging to pines (Pinus spp.) grown in plantations throughout the world, including Spain. To assess the potential for interspecific differences in susceptibility to contribute to the management of pitch canker and Diplodia shoot blight in the Atlantic region of Spain, the present study was undertaken to characterize the susceptibility of six pine species (P. sylvestris, P. nigra, P. pinaster, P. radiata, P. halepensis and P. pinea) and Douglas‐fir (Pseudotsuga menziesii) to F. circinatum and D. pinea. Based on inoculations of 2‐year‐old trees, Ps. menziesii, P. pinea and P. nigra were the most resistant to F. circinatum, with lesion lengths ranging from 3.7 to 21.5 mm, 2.2 to 12.6 mm and 2.8 to 30.9 mm, respectively. At the other extreme, Pinus radiata was the most susceptible, sustaining lesions that ranged from 8.5 to 74.8 mm in length. Pinus sylvestris, P. pinaster and P. halepensis showed an intermediate response to F. circinatum. Broadly similar results were observed in inoculations with D. pinea, with Ps. menziesii being relatively resistant and P. radiata being highly susceptible. Consistent with these results, field surveys revealed no pitch canker in stands of Ps. menziesii and low severity of Diplodia shoot blight, whereas P. radiata was severely affected by both diseases. Our findings suggest that selection of appropriate species can greatly reduce the risk of damage from two important canker diseases affecting pine plantations in the Atlantic region of Spain. Furthermore, intraspecific variation in susceptibility implies that selection may allow for the enhancement of resistance in otherwise susceptible species.  相似文献   

11.
In some areas of the Mediterranean basin where the understory stratum represents a critical fire hazard, managing the canopy cover to control the understory shrubby vegetation is an ecological alternative to the current mechanical management techniques. In this study, we determine the relationship between the overstory basal area and the cover of the understory shrubby vegetation for different dominant canopy species (Pinaceae and Fagaceae species) along a wide altitudinal gradient in the province of Catalonia (Spain). Analyses were conducted using data from the Spanish National Forest Inventory. At the regional scale, when all stands are analysed together, a strong negative relationship between mean shrub cover and site elevation was found. Among the Pinaceae species, we found fairly good relationships between stand basal area and the maximum development of the shrub stratum for species located at intermediate elevations (Pinus nigra, Pinus sylvestris). However, at the extremes of the elevation-climatic gradient (Pinus halepensis and Pinus uncinata stands), stand basal area explained very little of the shrub cover variation probably because microsite and topographic factors override its effect. Among the Fagaceae species, a negative relationship between basal area and the maximum development of the shrub stratum was found in Quercus humilis and Fagus sylvatica dominated stands but not in Quercus ilex. This can be due to the particular canopy structure and management history of Q. ilex stands. In conclusion, our study revealed a marked effect of the tree layer composition and the environment on the relationship between the development of the understory and overstory tree structure. More fine-grained studies are needed to provide forest managers with more detailed information about the relationship between these two forest strata.  相似文献   

12.
Pinelands in a 49 ha naturally-regenerated, mature flatwoods forest in north Florida were clearcut harvested in the fall of 1978, site prepared by burning, shearing and piling, discing, and bedding, and planted to slash pine (Pinus elliottii) in 1979. Three vegetation surveys were conducted: one prior to harvest in the summers of 1977 and 1978, and two subsequent to planting in the summers of 1980 and 1981.Cover and frequency of all plant species encountered were assessed on permanent transects. Foliage biomass by species was assessed by destructive sampling of distinct subplots within permanent plots. The aim was to assess plant species cover, frequency, and biomass responses to the forest operations imposed.Pine was eliminated as a dominant genus by harvesting. Planted pines were a fast increasing, but not dominant, component of the vegetation at 2 years of age. Previously dominant shrubs were severely reduced — often by approximately two orders of magnitude. Indeed, woody species were severely reduced: woody cover from 151 to 12% of surface area at plantation age 2 years; woody biomass from 6223 to 521 kg/ha.Conversely, herbaceous species were substantially increased: herbaceous cover from 38 to 51% of surface area at 2 years; herbaceous biomass from 382 to 1439 kg/ha. Thus, a predominantly woody ecosystem was converted to a predominantly herbaceous one for 2 years following planting.There was little change in plant species richness as a result of forest operations. Plant diversity changes were mixed the 1st year but diversity substantially increased the 2nd year. Comparisons with a companion study (Conde et al., 1983) suggest that, while increases in diversity following maximum site preparation may lag increases following minimum site preparation, cover, frequency, and biomass diversity all converge to common values after 2 years despite disparity in treatments.  相似文献   

13.
In the Okavango Delta information on the riparian woodland vegetation community composition and distribution is generally lacking. Past studies in the Delta were mainly focused on the quantitative classification of seasonal floodplain herbaceous vegetation communities. The aim of this study was to determine riparian woodland vegetation communities in the Okavango Delta. Vegetation sampling was conducted in 20?m × 50?m randomly placed plots. The plots were placed along a gradient from the main water body to the drier fringe of the riparian zone. Plant species present in each plot were recorded with their estimated percentage cover using the Braun–Blanquet cover abundance scale. Hierarchical cluster analysis was used to determine vegetation communities. Indicator species analysis was used to calculate indicator values for species groups defined from the cluster analysis. Plant species and diversity were determined for each vegetation cluster. Multi-response permutation procedures (MRPP) were used to determine significance of differences between communities. The Kruskal–Wallis test was used to compare species diversity and richness between vegetation communities. Four vegetation communities were identified: Croton megalobotrys– Hyphaene petersiana, Vachellia erioloba–Diospyros lycioides, Syzygium cordatum–Phoenix reclinata and Garcinia livingstonei–Senegalia nigrescens. In this classification, the Syzygium cordatum–Phoenix reclinata and Garcinia livingstonei–Senegalia nigrescens plant communities are found in frequently flooded regions, whereas the Croton megalobotrys–Hyphaene petersiana and Vachellia erioloba–Diospyros lycioides are in occasionally flooded regions. Total number of species was highest in the Garcinia livingstonei–Senegalia nigrescens and lowest in the Syzygium cordatum–Phoenix reclinata vegetation communities. The MRPP showed that there was significant (p < 0.05) differences between vegetation communities. Species diversity and richness were significantly (p < 0.05) higher in the Garcinia livingstonei–Senegalia nigrescens vegetation community. This study provided an inventory of riparian woodland plant communities, which can be used as a vital monitoring tool of vegetation change in the Okavango Delta.  相似文献   

14.
Herbivory may be an important factor affecting seedling survival of exotic species invading new habitats. We evaluated the effect of vertebrate herbivory on the seedling survival of two widely planted and invasive tree species (Pinus radiata and Eucalyptus globulus), in a Mediterranean-type ecosystem of central Chile. An important role of herbivory on seedling survival of these two species in their introduced ranges has previously been documented. However, this has mainly been evaluated in forest plantations where habitat and vegetation conditions differ from wild habitats in which invasion occurs. We planted seedlings with and without protection against vertebrate herbivores in different aspects (a mesic south-facing slope and a xeric north-facing slope) and vegetation cover (open sites and sites with patchy tree cover). We found that regardless of aspect or vegetation cover, herbivory, in this case mainly caused by exotic vertebrates, significantly and negatively affected seedling survival of both species. However, while the effect of herbivory on P. radiata was significant in every vegetation and habitat condition, for E. globulus, the effect of herbivory was significant only for open sites in the mesic habitat. These results suggest that, as observed in forestry plantations, vertebrate herbivory may constrain seedling establishment of these two exotic trees and potentially impede the invasion. However, the importance of herbivory in controlling exotic species may vary depending on the vegetation and habitat conditions in some species such as E. globulus.  相似文献   

15.
Plantation of open grazing lands with Pinus halepensis are the most widely used practices in afforestation and reforestation in Jordan and other semi-arid areas around the Mediterranean Basin. The effect of aspect North (N), South (S), East (E) and West (W) and slope position (upper, middle, lower and valley bottom) on growth of planted Aleppo pine, nutritional status, plantation on restoring, needle, forest floor, nutrients concentration and soil properties were studied in Jubilee forest in Rakeen area, south of Jordan. Tree height was significantly higher in W than N, S and E aspects which mainly due to better moisture and nutritional conditions. All growth parameters were obtained on valley bottom were significantly higher than all aspect slope position combinations due to accumulation of run off and depositions from upper towards middle, and finally in lower slopes. In general, pH and EC were significantly reduced and soil organic matter was significantly improved by Allepo pine plantations compared to unplanted areas. West and N aspects as well as valley bottoms showed better soil physical and chemical properties.  相似文献   

16.
Tube shelters were designed to protect against browsing, but they improve seedling survival in Mediterranean dry climates. Mechanisms for this response, however, are not fully understood and this knowledge can be useful to help design optimal tube shelters for Mediterranean species and climates. Our objective in this study was to determine if the positive effect of tube shelters is due to enhanced growth during the wet season or to reduced light stress during the dry season. We performed two independent experiments. In the first, we assessed root growth during the wet season in two Mediterranean species with contrasting light tolerance (Quercus ilex L. and Pinus halepensis Mill.) growing in tubes with varying light transmissivity. In the second experiment, we studied the response of a Quercus ilex plantation to different shelter treatments. Root growth during the wet season was reduced with decreasing light transmissivity in the shade intolerant P. halepensis, but not in the shade tolerant Quercus ilex. Survival of Q. ilex shaded by a mesh shelter only during summer was higher than in unsheltered seedlings and similar to the survival in tube and mesh shelters during the whole season. This suggests that shade during the dry period was the main factor explaining survival in this species. This effect could be related to the lower leaf temperature recorded in sheltered seedlings. We conclude that Q. ilex (and perhaps other late successional, shade tolerant Mediterranean species) should be planted in tubes with the currently used light transmissivity because these shelters reduce light stress in summer without impairing root growth in the wet season. However, current tubes impair root growth in P. halepensis (and likely other pioneer, shade intolerant Mediterranean species), so higher transmissivity tubes may be necessary. Optimal transmissivity for tube shelter in Mediterranean climates is species-specific and identifiable as the point that minimizes light stress during summer without impairing root growth in the wet season.  相似文献   

17.
We lack information regarding the main factors driving growth responses to drought in tree species with different vulnerability against this stressor and considering sites with contrasting climatic conditions. In this paper, we identify the main drivers controlling growth response to a multi-scalar drought index (Standardized Precipitation Index, SPI) in eight tree species (Abies alba, Pinus halepensis, Quercus faginea, Pinus sylvestris, Quercus ilex, Pinus pinea, Pinus nigra, Juniperus thurifera). We sampled forests growing across a pronounced climatic gradient under Mediterranean conditions in north-eastern Spain. To summarize the patterns of growth responses to drought, we used principal component analysis (PCA). To determine the main factors affecting growth responses to drought, correlation and regression analyses were carried out using a set of abiotic (climate, topography, soil type) and biotic (Normalized Difference Vegetation Index, Enhanced Vegetation Index, tree-ring width, diameter at breast height) predictors and the PCs loadings as response variables. The PCA analysis detected two patterns of growth responses to drought corresponding to xeric and mesic sites, respectively. The regression analyses indicated that growth responses to drought in xeric forests were mainly driven by the annual precipitation, while in mesic sites the annual water balance was the most important driver. The management of Mediterranean forests under the forecasted warmer and drier conditions should focus on the main local factors modulating the negative impacts of drought on tree growth in xeric and mesic sites.  相似文献   

18.
The influence of woody and herbaceous plant competition, either alone or in combination, on microclimate and growth of planted eastern white pine (Pinus strobus L.) seedlings was examined over four consecutive growing seasons in a central Ontario clearcut. Treatments that manipulated the comparative abundance of these two plant functional groups significantly affected light availability, soil moisture, and air and soil temperature regimes. These microclimate alterations, coupled with the relative competitiveness of herbaceous and woody vegetation, corresponded to temporal changes in vegetation cover and dominance. The more rapid colonization and growth of the herbaceous plant community, dominated by bracken fern (Pteridium aquilinum) and ericaceous shrubs (Kalmia sp., Vaccinium sp.), resulted in this form of vegetation being a comparatively important early competitor for soil moisture. As the woody plant community, dominated by naturally regenerated trembling aspen (Populus tremuloides Michx.), grew in height and leaf area, it became a comparatively strong competitor for both light and soil moisture. For all vegetation treatments combined, white pine seedling growth responses were strongly correlated with total cover of competing vegetation and its relative influence on above- and belowground microclimatic variables. Higher total cover of competing vegetation was generally associated with lower light and soil moisture availability and cooler soil temperatures. Multiple regression analyses indicated that pine seedling relative height growth increased with soil moisture content and growing season soil heat sum, while seedling relative diameter and relative volume growth increased with light availability.  相似文献   

19.
During a survey on the distribution of Heterobasidion root rot, the North American P group of H. annosum was recorded causing root rot on Aleppo pine, Pinus halepensis, in the gardens of a historical villa in Rome. This is the first report of Heterobasidion root rot on P. halepensis in its natural distribution area, the Mediterranean Basin.  相似文献   

20.
Forested riparian buffers in California historically supported high levels of biodiversity, but human activities have degraded these ecosystems over much of their former range. This study examined plant communities, belowground biodiversity and indicators of multiple ecosystem functions of riparian areas across an agricultural landscape in the Sacramento Valley of California, USA. Plant, nematode and soil microbial communities and soil physical and chemical properties were studied along 50-m transects at 20 sites that represented the different land use, soil and vegetation types in the landscape. Riparian zones supported greater plant diversity and nearly twice as much total carbon (C) per hectare compared to adjacent land managed for agricultural uses, but had generally lower soil microbial and nematode diversity and abundance. When woody plant communities were present in the riparian zone, plant diversity and species richness were higher, and soil nitrate and plant-available phosphorus levels were lower. Belowground diversity and community structure, however, appeared to depend more on plant productivity (as inferred by vegetation cover) than plant diversity or species richness. Greater plant species richness, nematode food web structure, total microbial biomass, woody C storage and lower soil nitrate and phosphorus loading were correlated with higher visual riparian health assessment scores, offering the possibility of managing these riparian habitats to provide multiple ecosystem functions.  相似文献   

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