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1.
Lianas are often overlooked in temperate ecological studies even though they are important components of forest communities. While lianas have been shown to damage tropical canopy trees and reduce the growth of juvenile trees, the impact of lianas on canopy tree growth in temperate systems is largely unknown. Growth of trees ≥8 cm dbh was examined over a 9-year period within 50-year old post-agricultural secondary forests in the Piedmont region of New Jersey, USA. Five lianas, Celastrus orbiculatus, Lonicera japonica, Parthenocissus quinquefolia, Toxicodendron radicans, and Vitis species, occurred throughout the forest. Total liana basal area, number of stems, and percent cover within host trees were evaluated to assess liana burdens on 606 previously censused trees. These data were related to tree growth to assess liana impacts. Forest trees were separated based on their dominance in the canopy to determine whether lianas had the potential to influence forest composition. In general, lianas in the forests were fairly abundant, with 68% of the trees having at least one liana present. On average, each tree supported 9.7 cm2 of liana basal area and 23% of the canopy was covered by lianas. Most of the variation in tree growth was related to the dominance of trees within the canopy, with canopy dominant and co-dominant trees growing 2.5× more than suppressed trees. Liana basal area and number of lianas stems were not related to tree growth, but liana canopy cover decreased tree growth. However, not all trees were equally affected as canopy cover of lianas only reduced growth in dominant and co-dominant trees. Lianas were most influential on host tree growth in unsuppressed trees when occupying a majority of the canopy, only a minority of forest trees. This suppression was not related to differential liana colonization of canopy trees as all canopy classes supported equivalent liana burdens. Though lianas impacted only a minority of the trees in this system, some liana species, C. orbiculatus and Vitis spp., are still increasing and may pose future risks to forest growth and development.  相似文献   

2.
  • ? Understanding the effects of tree species diversity on biomass and production of forests is fundamental for carbon sequestration strategies, particularly in the perspective of the current climate change. However, the diversity-productivity relationship in old-growth forests is not well understood.
  • ? We quantified biomass and above-ground production in nine forest stands with increasing tree species diversity from monocultures of beech to stands consisting of up to five deciduous tree species (Fagus sylvatica, Fraxinus excelsior, Tilia spp., Carpinus betulus, Acer spp.) to examine (a) if mixed stands are more productive than monospecific stands, (b) how tree species differ in the productivity of stem wood, leaves and fruits, and (c) if beech productivity increases with tree diversity due to lower intraspecific competition and complementary resource use.
  • ? Total above-ground biomass and wood production decreased with increasing tree species diversity. In Fagus and Fraxinus, the basal area-related wood productivity exceeded those of the co-occurring tree species, while Tilia had the highest leaf productivity. Fagus trees showed no elevated production per basal area in the mixed stands.
  • ? We found no evidence of complementary resource use associated with biomass production. We conclude that above-ground productivity of old-growth temperate deciduous forests depend more on tree species-specific traits than on tree diversity itself.
    •   相似文献   

    3.
    Liana habitat and host preferences in northern temperate forests   总被引：1，自引：0，他引：1  
    Lianas and other climbers are important ecological and structural components of forest communities. Like other plants, their abundance in a given habitat depends on a variety of factors, such as light, soil moisture and nutrients. However, since lianas require external support, host tree characteristics also influence their distribution. Lianas are conspicuous life forms in tropical regions, but in temperate areas, where they are less prominent, little is known about factors that control their distributions in these forests. We surveyed the climbing plant species in 20 mature (100 years and greater) forested habitats in the Midwest USA at a variety of levels from simple presence/absence, to ground layer abundances, to those species that had ascended trees. We also examined attributes of the tree species with climbers attached to them. Using cluster analysis, we distinguished five different tree communities in our survey locations. We determined that 25% of the trees we surveyed had one or more lianas attached to it, with Parthenocissus quinquefolia (Virginia creeper) the most common climbing species encountered. Canopy cover and soil attributes both influenced climber species presence/absence and ground layer climber abundance. The proportion of liana species of a given climbing type (roots, stem twiner, tendril climber) was significantly related to the DBH of the host tree, with more root climbers and fewer stem and tendril climbers on large trees. In general, the DBH of climbing lianas had a significant positive relationship to the DBH of the host tree; however this varied by the identity of the liana and the tree species. The greater the DBH of the host tree, the higher the probability that it was colonized by one or more lianas, with tree species such as Pinus banksiana (jack pine) and Quercus alba (white oak) being more susceptible to liana colonization than others. Finally, some liana species such as Celastrus scandens (American bittersweet) showed a preference for certain tree species (i.e., P. banksiana) as hosts. The information obtained about the relationship between the tree and climber community in this study provides insight into some of the factors that influence liana distributions in understudied temperate forest habitats and how lianas contribute to the structure of these mature forests. In addition, these data can provide a point of comparison to other liana communities in both temperate and tropical regions.  相似文献   

    4.
    栎属树种生长模型研究进展   总被引：1，自引：0，他引：1  
    栎属树种是亚热带常绿阔叶林和温带落叶阔叶林的主要建群树种之一,分布范围极为广泛,在国内外被广泛应用于城市园林绿化、水源涵养林、水土保持林,也是重要的珍贵用材树种,同时其果实、栓皮等具有重要的工业和药用价值.研究栎林的生长过程,对其经营抚育决策具有重要的参考意义.文中对近年来国内外关于栎属树种的生长过程,特别是栎属树种生长模型的研究进行了综述,包括栎属树种全林分生长模型、单木生长模型、径阶分布模型等;阐述了栎属树种生长模型的研究现状及发展趋势,以期为栎林的经营抚育决策提供参考.  相似文献   

    5.
    分别在哀牢山湿性常绿阔叶林沟谷和坡面调查了10个20m×50m的样地,研究哀牢山湿性常绿阔叶林木质藤本植物的物种多样性及其与支柱木的关系。结果表明:研究区共记录到DBH≥1cm的木质藤本植物402株(隶属于23种21属16科)和DBH≥10cm的林木1522株(隶属于47种30属15科);与其他亚热带森林比较,该森林中木质藤本植物物种较为丰富但多度较低;藤本植物的物种丰富度、密度和基面积在沟谷显著高于坡面,而林木的差异性不显著;木质藤本植物在支柱木上呈集群分布,并且不同种支柱木被藤本植物攀援的百分比间存在显著的差异(P0.001),说明藤本植物的攀援对支柱木具有选择性;云南越桔、薄叶马银花和景东冬青等树种因其树皮光滑而不易被藤本植物攀援,而腾冲栲、七裂槭、山矾和多花山矾等则易于被藤本植物攀援;大径级支柱木被藤本植物攀援的比率高于小径级支柱木;茎缠绕和钩刺攀援藤本的胸径与支柱木胸径极显著相关(P0.001),根攀援和卷须攀援藤本的胸径与支柱木胸径相关性不显著(P0.05)。  相似文献   

    6.
    How to quantify forest management intensity in Central European forests   总被引：1，自引：0，他引：1  
    Existing approaches for the assessment of forest management intensity lack a widely accepted, purely quantitative measure for ranking a set of forest stands along a gradient of management intensity. We have developed a silvicultural management intensity indicator (SMI) which combines three main characteristics of a given stand: tree species, stand age and aboveground, living and dead wooden biomass. Data on these three factors are used as input to represent the risk of stand loss, which is a function of tree species and stand age, and stand density, which is a function of the silvicultural regime, stand age and tree species. Consequently, the indicator consists of a risk component (SMIr) and a density component (SMId). We used SMI to rank traditional management of the main Central European tree species: Norway spruce (Picea abies [Karst.] L.), European beech (Fagus sylvatica L.), Scots pine (Pinus sylvestris L.), and oak (Quercus robur L. and Quercus petraea L.). By analysing SMI over their whole rotation period, we found the following ranking of management intensity: oak<beech<pine?spruce. Additionally, we quantified the SMI of actual research plots of the German Biodiversity exploratories, which represent unmanaged and managed forest stands including conifer forests cultivated outside their natural range. SMI not only successfully separate managed from unmanaged forests, but also reflected the variability of forest management and stand properties across the entire sample and within the different management groups. We suggest using SMI to quantify silvicultual management intensity of stands differing in species composition, age, silvicultural system (even-aged vs. uneven-aged), thinning grade and stages of stand conversion from one stand type into another. Using SMI may facilitate the assessment of the impact of forest management intensity on biodiversity in temperate forests.  相似文献   

    7.
    Root disease pathogens, including Armillaria, are a leading cause of growth loss and tree mortality in forest ecosystems of North America. Armillaria spp. have a wide host range and can cause significant reductions in tree growth that may lead to mortality. DNA sequence comparisons and phylogenetic studies have allowed a better understanding of Armillaria spp. taxonomic diversity. Genetic sequencing has facilitated the mapping of species distributions and host associations, providing insights into Armillaria ecology. These studies can help to inform forest management and are essential in the development of disease risk maps, leading to more effective management strategies for Armillaria root disease. Armillaria surveys were conducted on publicly owned lands in North Dakota, South Dakota, and Nebraska, U.S.A. Surveyed stands consisted of riparian forests ≥0.4 hectares in area. Armillaria was found at 78 of 101 sites. A total of 57 Armillaria isolates—associated with 12 host tree species—were used for DNA sequencing of the translation elongation factor‐1 alpha (tef1) gene. Armillaria gallica was the only species identified within the study sites. Results suggest that A. gallica is a common root pathogen of hardwood trees in riparian forests of the northern Great Plains with a wider host range and geographic distribution than previously recognized.  相似文献   

    8.

    This study investigated the stand structure in pine, spruce and deciduous forests in the border district of Finland and Russia. A total of 46 mature forest stands was selected as pairs, the members of each pair being as similar as possible with respect to their forest site type, age, moisture and topography. The stands were then compared between the two countries by means of basal areas and number of stems. The proportions of dominating tree species were 2-12% lower, and correspondingly the proportions of secondary tree species higher, in Russian forests. The density of the forest stock was also higher in each forest type in Russia. The forests in the two countries differed most radically in terms of the abundance of dead trees. The amount was two to four times higher in Russian deciduous and spruce forests, and in pine forests the difference was 10-fold. The stand structures indicated that Russian coniferous stands, in particular, were more heterogeneous than intensively managed pine and spruce stands in Finland.  相似文献   

    9.
    Bees and wasps provide important ecosystem services such as pollination and biocontrol in crop-dominated landscapes, but surprisingly little information is available on hymenopteran communities in temperate forest ecosystems. Species richness and abundance of bees and wasps can be hypothesised to increase with plant diversity, structural complexity, and availability of food and nesting resources. By experimentally exposing standardised nesting sites, we examined abundance and species richness of cavity-nesting bees (pollinators), wasps (predators) and their associated parasitoids across a tree diversity gradient in a temperate deciduous forest habitat. In addition, spatial distribution of individuals and species across forest strata (canopy vs. understory) was tested. Abundance and species richness was high for predatory wasps, but generally low for pollinators. Species-rich forest stands supported increased abundance, but not species richness, of pollinators and predatory wasps, and also increased abundance and species richness of natural enemies. In addition, the forests showed a distinct spatial stratification in that abundance of bees, wasps and parasitoids as well as parasitism rates were higher in the canopy than understory. We conclude that particularly the canopy in temperate forest stands can serve as an important habitat for predatory wasp species and natural enemies, but not bee pollinators. Enhanced tree diversity was related to increased hymenopteran abundance, which is likely to be linked to an increase in nesting and food resources in mixed forest stands.  相似文献   

    10.
    We aimed to study tree effects on the chemical properties of forest soils. We compared soil features of three types of forest ecosystems, each with four stands (replicates): beech forests (Fagus sylvatica), oak forests (dominated by Quercus pyrenaica) and pine plantations (Pinus sylvestris). Five samples from the top 10 cm of soil were taken per stand, from which pH, organic matter content (O.M.), total nitrogen (N) and available calcium (Ca2+), magnesium (Mg2+), potassium (K+) and sodium (Na+) were determined. Litter layer depth was measured at each soil sampling point. We also measured tree density and crown diameters at each stand. Our results indicated that soil samples from the four pine plantation stands were more similar while oak and beech stands were characterised by great variability in terms of soil properties and leaf litter depth. Although the identity of the dominant tree species significantly influenced several topsoil chemical properties (increase in pH and available cations in oak forests and higher organic matter and total nitrogen in beech and pine ecosystems), there were other important factors affecting soil features that may be taken under consideration. Differences between soil properties of the three types of forest ecosystems were mainly related to the characteristics of the litter layer and less related to the tree layer structure. Finally, the establishment of pine plantations in naturally deciduous tree areas made the topsoil features more homogeneous.  相似文献   

    11.
    The relationships between the structural complexity of coniferous forests and the epiphytic lichen communities that inhabit them were examined in 51 conifer-dominated stands in southwestern Nova Scotia. One hundred and fifteen lichen species were studied in stands in the age range of 50–300 years. Environmental variables shaping the structural complexity of each forest stand were measured and their relationship with lichen species were assessed using a canonical correspondence analysis (CCA). The CCA revealed that the considerable variation in lichen community composition can be explained by several environmental variables associated with forest structure. The stand orientation on the first axis of the CCA found the most important variables for lichen richness to be stand age, tree stem density and snag stem density. The stand orientation on the second axis is strongly correlated with deciduous stem density and abundance including specific deciduous tree species such as Acer rubrum abundance. The analysis indicates that the greater the structural complexity in the forest, and thus the more microhabitats available, the greater the lichen species richness. These results should provide forest managers with a better understanding of the environmental variables that influence lichen diversity, and contribute to the development of more sustainable forest management strategies.  相似文献   

    12.
    A field study was conducted in the moist deciduous forests of the Western Ghats (India) to test the following three hypotheses: (1) Litter production in tropical forests is a function of the floristic composition, density, basal area and disturbance intensity; (2) Decay rate constants of tropical species is an inverse function of the initial lignin/nitrogen ratio; (3) Decomposition rates in tropical forests are faster than temperate forests.

    Litter fall was estimated by installing 63 litter traps in the moist deciduous forests of Thrissur Forest Division in the Western Ghats at three sites. Litter fall followed a monomodal distribution pattern with a distinct peak during the dry period from November–December to March–April.Dillenia pentagyna, Grewia tiliaefolia, Macrosolen spp.,Xylia xylocarpa, Terminalia spp.,Lagerstroemia lanceolata, Cleistanthus collinus, Bridelia retusa, andHelicteres isora were the principal litter producing species at these sites. The annual litter fall ranged from 12.18 to 14.43 t ha−1. Structural attributes of vegetation such as floristic composition, basal area, density and disturbance intensity did not directly influence litter fall rates.

    Leaf litter decay rates for six dominant tree species were assessed following the standard litter bag technique. One hundred and eight litter bags per species containing 20 g samples were installed in the forest floor litter layer at the same three sites selected for the litter fall quantification exercise. The residual litter mass decreased linearly with time for all species. In general, less disturbed sites and species adapted to higher nitrogen availabilities exhibited relatively higher decay rate coefficients (k). The rapid organic matter turnover observed in comparison with published temperate forest litter decay rates confirms that tropical moist deciduous forest species are characterised by faster decomposition rates.

    Mean concentrations of N, P and K in the litter were profoundly variable amongst the dominant species. Initial nitrogen content of the leaf litter varied from 0.65 to 1.6%, phosphorus from 0.034 to 0.077% and potassium from 0.25 to 0.62%.C. collinus, an understorey shrub consistently recorded the lowest litter concentrations for all nutrients. The overriding pattern is one of higher nutrient levels in the overstorey leaf litter and lower concentrations in the understorey litter. Furthermore, as decomposition proceeded, the nitrogen concentration of the residual biomass increased.  相似文献   


    13.
    How can we accommodate the diversity in tree species and sizes in mixed-species/size/age stands in the sustainable management of natural forests and woodlands in Africa for diverse timber and non-timber forest products and services, and during rehabilitation of degraded forests? The evergreen moist tropical to warm-temperate forests, from the equator to 34°S, generally function through relative shade tolerance. The tropical, strongly seasonal, drier deciduous woodlands generally function through adaptation to fire and/or grazing/browsing. Silvicultural systems, when implemented, are often not aligned with the ecological characteristics of the particular forest systems or the specific targeted species. This paper presents the concept of using the basic disturbance–recovery processes, with recovery development via stand development stages, as the basis for the development of silvicultural systems suitable for maintenance of forest complexity. Grain analysis and stem diameter distributions, analysed from resource inventories, are used to determine the specific shade- or fire-tolerance characteristics of key economical and ecological tree species. The gained knowledge of the ecosystem and species characteristics (including modes of regeneration, i.e. from seed or vegetative regrowth) and processes is used to simulate the ecological disturbance–recovery processes through the development of mixed silvicultural systems, such as a single-tree selection system, a group felling system and a coppice management system in the same forest. Very similar concepts are used to develop rehabilitation strategies to recover the processes towards regrowth stands of diverse species and structure: through stands of introduced plantations and invader tree species; and through early regrowth stages in deciduous woodlands and evergreen forests, after fire, slash-and-burn traditional agriculture, charcoal production or open-cast mining.  相似文献   

    14.
    The effect of forest management on biodiversity is a crucial issue for sustainable forestry and nature conservation. However, the ways in which management affects macrofungal and plant communities and diversity of mountain temperate forests still remain poorly understood. We performed a random sampling stratified by stand age and stand type on the sites of temperate montane fir–beech forests. Diversity of macrofungi and the vascular plant understorey in beech- and spruce-dominated managed stands was investigated and compared to primeval forests located in the Po?ana Biosphere Reserve, Western Carpathians. Both the vascular plant and the macrofungal communities were altered by management, and the response of the macrofungal species (especially wood-inhabiting fungi) was more pronounced in terms of species composition change. Species turnover evaluation seems to be an important tool of forest natural status assessment, because alpha diversity did not change as much as species composition. Certain species of Carpathian primeval forests were confirmed as good indicators for natural forest change; others were proposed. Species pool and mean number of species per plot were the highest in unmanaged fir–beech forests, and species diversity significantly decreased in spruce plantations. The number of species decreased significantly due to the change of canopy tree species composition only in the macrofungal communities. As an outcome for forest management, we recommend keeping mixed forests involving all natural tree species and providing at least a minimal amount of dead wood necessary for wood-inhabiting organisms and leaving some area of unmanaged natural forests within complexes of managed stands.  相似文献   

    15.
    To document the spatial and temporal variation of environmental signals inducing seed germination in temperate forests, we measured temporal patterns of environmental signals and seed germination of six pioneer tree species in unthinned and thinned stands of conifer forests (Cryptomeria japonica plantations) and in the understory and gaps of hardwood forests in Japan. We also conducted germination experiment in laboratory for the six pioneer species to test the effects of red:far-red (R:FR) light ratio and temperature fluctuations on the seed germination. In conifer forests, the photosynthetic photon flux density (PPFD), the R:FR ratio, and the amplitude of temperature fluctuations in thinned stands were 2, 1.5, and 3 times higher, respectively, than those of unthinned stands. The PPFD and R:FR ratios just above forest floor also increased after the removal of thick litter accumulation. As a result, higher seed germination was observed in thinned compared to unthinned stands for three photoblastic species, whereas little differences were observed for three non-photoblastic species. These findings suggest that thinning, which frequently reduces litter accumulation, can substantially affect the regeneration of pioneer species and the resultant species diversity in conifer plantations. None of the measured environmental signals changed seasonally in unthinned stands of conifer forests, but they all changed remarkably in the understory of the hardwood forests. In this system, all signals were high and nearly identical to those in the gaps in early spring prior to canopy closure. Thus, the percent germination of the three photoblastic species was enhanced by high R:FR ratios and/or large temperature fluctuations even beneath the canopy and was nearly equal to that in the thinned conifer stands where the environmental conditions were nearly identical to those in the gaps. However, all of the environmental signals decreased with the expansion of canopy leaves and reached minimums at canopy closure. Even in the thinned stands and the gaps, the PPFD and magnitude of temperature fluctuations decreased over time due to shading by growing herbs and/or emerging canopy leaves. In these temporally changing environments, the germination of all photoblastic species ceased simultaneously. This study clearly demonstrated that the environmental signals inducing seed germination of photoblastic pioneer species spatially and temporally change in temperate forests, particularly in deciduous hardwood forests. Furthermore, these signals, PPFD, R:FR ratio, and the amplitude of temperature fluctuations, appear to play a very important role in tree regeneration and subsequent species diversity.  相似文献   

    16.
    Large cavity-nesting birds depend on large-diameter trees for suitable nest sites. The increased spatial extent of commercial timber harvesting is modifying forest structure across the land base and may thus compromise the availability of large trees at the landscape scale. In this study, our objectives were to (1) characterize the availability of large living and dead trees in old-growth stands dominated by different tree species and surficial deposits that encompass the range of natural cover types of eastern Québec's boreal forest; (2) analyze the distribution of trees among decay-classes; and (3) compare the availability of large trees in unharvested, remnant, and harvested stands for the entire range of decay-classes. A total of 116 line transects were distributed across unharvested forests, remnant linear forests, and cutblocks in cutover areas. Unharvested forest stands (black spruce [Picea mariana], balsam fir [Abies balsamea]–black spruce, balsam fir–white spruce [Picea glauca] and balsam fir) reflected a gradient of balsam fir dominance. The remnant forests selected were isolated for 5–15 years. Analyses were performed at two diameter cut-off values. Trees with DBH ≥20 cm were considered for availability of total trees whereas trees with DBH ≥30 cm were considered for availability of large trees. Forest stands comprised high proportions of standing dead trees (33% of all stems, 8% were large dead stems). Availability of total and large standing trees increased with the dominance of balsam fir in stands. Forest stands located on thick surficial deposits showed higher densities of large dead trees for every stand type suggesting a higher productivity on those sites. Availability of stems according to decay-classes showed a dome-shaped distribution with higher densities of snags in intermediate decay stages. However, for large stems, black spruce stands showed a significantly lower availability that was consistent across all decay-classes. In linear remnant forests, pure balsam fir stands were absent. Remnant stands thus showed a much lower availability in large trees when compared with unharvested balsam fir stands. Clearcuts had the lowest densities of dead trees across sampled stands. Current even-aged management practices clearly affect availability and recruitment of large trees, therefore forest-dwelling wildlife relying on these structures for breeding is likely to be affected by large-scale harvesting in coniferous boreal forests.  相似文献   

    17.
    Within the eastern deciduous forest region, forest composition varies, with some areas dominated by a mix of oaks (Quercus spp.) and other areas dominated by a mix of sugar maple (Acer saccharum) and other tree species. Prescribed fire is being used on an experimental basis to assess its effectiveness in restoring and maintaining oak-dominated forests. Maple-dominated forests are susceptible to invasion by non-native earthworms, such as Lumbricus terrestris, given the palatability of leaf litter and suitable soil conditions, especially in northern parts of the region. What are the implications of this variation on leaf litter availability and habitat for ground-nesting songbirds? We investigated this question by comparing forest composition, leaf litter, and songbird communities in maple-dominated forests in west-central Indiana and oak-dominated forests, recently burned and unburned, in southeastern Ohio. We also assessed abundance of earthworms and decomposition rates of different types of leaves in the maple-dominated forests in Indiana. Leaf litter and ground-nesting bird species were abundant in unburned oak-dominated forests, but were absent or nearly absent in recently burned oak-dominated forests and in maple-dominated forests. The lack of leaf litter and absence of ground-nesting bird species in maple-dominated forests may be due to the combination of abundant non-native earthworms, alkaline and calcium-rich soils, palatable leaves, and rapid leaf litter decomposition rates. Effects of burning on leaf litter and ground-nesting bird species in oak-dominated forests are probably temporary, as long as prescribed fires are not applied on a frequent or widespread basis. Our study is the first one to show a correlation between forest composition, leaf litter availability, earthworm abundance, and songbird populations. Many researchers are investigating effects of non-native earthworm invasions on ecosystem properties in eastern deciduous forests. We recommend that researchers should also monitor songbird populations to assess whether declines in ground-nesting bird populations are occurring in response to these changes.  相似文献   

    18.
    Lianas (woody vines) are an important component of tropical forests, with a strong impact on forest dynamics, but their responses during forest succession have received relatively little attention. Here, we present an analysis of the changes in stem density, biomass, and species richness of lianas and self-supporting plants during tropical forest succession. We surveyed lianas ≥0.5 cm diameter at breast height (dbh) and self-supporting plants ≥2.5 cm dbh in 0.1 ha inventory plots in a chronosequence of 30 sites in northeastern Costa Rica, 23 sites on abandoned pastures 10–44 years of age, and seven sites in old-growth forest. Stem density of self-supporting plants showed no predictable chronosequence trend, but liana stem density declined significantly with forest age. Aboveground biomass of self-supporting vegetation increased rapidly during succession, with forests 31–44 years exhibiting higher levels of biomass than old-growth forests. Liana biomass accumulated more slowly, with the highest levels in old-growth sites. Species richness of self-supporting vegetation increased significantly during succession, but species richness of lianas showed no change or a slight decline with forest age, depending on the method of assessment. The differences between tree and liana responses during succession stem from the unique physiology and life history traits of lianas.  相似文献   

    19.
    Abstract

    In this study, I defined a gap as a small opening formed in a forest canopy (area < 0.1 ha) and tried to synthesize gap-disturbance regimes of primary mature stands in different forest types of Japan, such as warm temperate evergreen broad-leaved (4 stands in 3 localities), cool temperate deciduous broad-leaved (10 in 5) and subalpine evergreen coniferous (3 in 1) forests. Mean percentage of the total gap area within the total forested area (percentage gap area) in each forest type was 17.0% in warm temperate (number of surveyed gaps was 161), 13.8% in cool temperate (278 gaps), and 8.0% in subalpine (100 gaps) forests. Mean gap density (ha-1) and mean gap size (m2) were 19.5 and 77.1 in warm temperate, 16.4 and 92.0 in cool temperate, and 19.1 and 41.9 in subalpine forests, respectively. These figures indicate that gap density is not substantially different among the forest types, but the mean gap size of subalpine forests is smaller than the other two, resulting in lower percentage gap area of this forest type. The gap size distributions were similar among the forest types; smaller gaps were much more frequent than larger ones, and gaps > 400 rrr were rare in warm temperate and cool temperate forests. In subalpine forests gaps > 200 m2 were rare. Gaps formed by multiple gapmakers comprised 19.9% of all gaps in warm temperate, 9.9% in cool temperate, and 44.9% in subalpine forests, which implies that gap formation by simultaneous tree fall or gap enlargement is more frequent in subalpine forests. Canopy trees died less often by uprooting in every forest type; dominant mode for the death of canopy trees was by leaving standing-dead or with broken trunks in every forest type. Since typhoons are obviously a chief agent of forest disturbance in Japan, frequency or magnitude of typhoon disturbance may influence these differences in the gap-disturbance regimes. In addition to the disturbance, tree architectures seem to affect some of these differences; narrower crown size of conifers compared with broad-leaved trees is considered one major cause for smaller gap size of the conifer forest.  相似文献   

    20.
    Where natural production capacity permits, modern silvicultural management in Central Europe frequently aims at the development of mixed broadleaved stands, instead of pure European beech (Fagus sylvatica) stands. It is crucial to study the effects of these tree-layer diversity variations on herb-layer vegetation, since herb-layer vegetation contributes significantly to ecosystem functioning in forests. In Hainich National Park (Thuringia, Germany), we conducted observational research in deciduous stands to investigate whether herb-layer diversity was related to canopy-layer diversity, and to ascertain possible causal mechanisms. We found that herb-layer vegetation of deciduous forest stands rich in canopy species appeared to be more diverse than herb-layer vegetation of beech-dominated stands. We surmise that herbaceous understorey diversity was indirectly influenced by canopy tree species through the medium of the altered environmental factors soil pH and litter layer thickness. Apparently, lower beech proportion had a more profound effect than the number of secondary tree species. There were no correlations between herb-layer diversity and light transmissibility of the canopy layer, indicating that the light factor was not crucial for herb-layer diversity. At least for the Hainich research sites, our results indicated that small-scale light and soil heterogeneity is insignificant for herb-layer diversity. We found several herb-layer species whose occurrence was particularly correlated with tree-layer diversity and environmental factors. Remarkably, all species positively correlated with soil pH were important for the phytosociological classification of the research sites. Beech-dominated research sites showed high tree-layer volumes, whereas research sites with high tree-layer diversity tended to feature lower tree-layer volumes. These findings could be the result of differing former silvicultural systems and varying soil clay contents affecting tree species composition. In contrast, herb-layer biomass was positively correlated with tree-layer diversity. Herb-layer productivity might be promoted in more diverse research sites by increased nutrient supply and base saturation. It is also possible that greater beech proportion interfered with herb-layer productivity. However, herb-layer biomass was also positively correlated with herb-layer diversity. Hence, our study hints that positive diversity-functioning relationships might occur in the herb-layer of the deciduous forest under investigation.  相似文献   

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