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1.
Early ocean survival of Chinook salmon, Oncorhynchus tshawytscha, varies greatly inter‐annually and may be the period during which later spawning abundance and fishery recruitment are set. Therefore, identifying environmental drivers related to early survival may inform better models for management and sustainability of salmon in a variable environment. With this in mind, our main objectives were to (a) identify regions of high temporal variability in growth potential over a 23‐year time series, (b) determine whether the spatial distribution of growth potential was correlated with observed oceanographic conditions, and (c) determine whether these spatial patterns in growth potential could be used to estimate juvenile salmon survival. We applied this method to the fall run of the Central Valley Chinook salmon population, focusing on the spring and summer period after emigration into central California coastal waters. For the period from 1988 to 2010, juvenile salmon growth potential on the central California continental shelf was described by three spatial patterns. These three patterns were most correlated with upwelling, detrended sea level anomalies, and the strength of onshore/offshore currents, respectively. Using the annual strength of these three patterns, as well as the overall growth potential throughout central California coastal waters, in a generalized linear model we explained 82% of the variation in juvenile salmon survival estimates. We attributed the relationship between growth potential and survival to variability in environmental conditions experienced by juvenile salmon during their first year at sea, as well as potential shifts in predation pressure following out‐migration into coastal waters.  相似文献   

2.
Variation in prey quantity and quality can influence growth and survival of marine predators, including anadromous fish that migrate from freshwater systems. The objective of this study was to examine the energy dynamics of subyearling Chinook salmon (Oncorhynchus tshawytscha) following freshwater emigration. To address this objective, a population of Chinook salmon and their marine prey were repeatedly sampled from June to September over 2 years in coastal waters off Oregon and Washington. Subyearlings from the same population were also reared under laboratory conditions. Using a bioenergetics model evaluated in the laboratory, we found that growth rate variability in the field was associated more with differences in northern anchovy (Engraulis mordax) consumption and less with variation in diet energy density or ocean temperature. Highest growth rates (2.43–3.22% body weight/day) occurred in months when anchovy biomass peaked, and the timing of peak anchovy biomass varied by year. Our results support a general pattern among subyearling Chinook salmon occurring from Alaska to California that feeding rates contribute most to growth rate variability during early marine residence, although dominant prey types can differ seasonally, annually, or by ecosystem. In the northern California Current, faster growth appears to be associated with the availability of age‐0 anchovy. Identifying factors that influence the seasonal development of the prey field and regulate prey quantity and quality will improve understanding of salmon growth and survival during early marine residence.  相似文献   

3.
We determined the habitat usage and habitat connectivity of juvenile Chinook (Oncorhynchus tshawytscha) and coho (Oncorhynchus kisutch) salmon in continental shelf waters off Washington and Oregon, based on samples collected every June for 9 yr (1998–2006). Habitat usage and connectivity were evaluated using SeaWiFS satellite‐derived chlorophyll a data and water depth. Logistic regression models were developed for both species, and habitats were first classified using a threshold value estimated from a receiver operating characteristic curve. A Bernoulli random process using catch probabilities from observed data, i.e. the frequency of occurrence of a fish divided by the number of times a station was surveyed, was applied to reclassify stations. Zero‐catch probabilities of yearling Chinook and yearling coho salmon decreased with increases in chlorophyll a concentration, and with decreases in water depth. From 1998 to 2006, ～ 47% of stations surveyed were classified as unfavorable habitat for yearling Chinook salmon and ～ 53% for yearling coho salmon. Potentially favorable habitat varied among years and ranged from 9 856 to 15 120 km2 (Chinook) and from 14 800 to 16 736 km2 (coho). For both species, the smallest habitat area occurred in 1998, an El Niño year. Favorable habitats for yearling Chinook salmon were more isolated in 1998 and 2005 than in other years. Both species had larger and more continuous favorable habitat areas along the Washington coast than along the Oregon coast. The favorable habitats were also larger and more continuous nearshore than offshore for both species. Further investigations on large‐scale transport, mesoscale physical features, and prey and predator availability in the study area are necessary to explain the spatial arrangement of juvenile salmon habitats in continental shelf waters.  相似文献   

4.
Unusually large returns of several stocks of fall Chinook salmon (Oncorhynchus tshawytscha) from the U.S. Northwest commonly occurred during the late 1980s. These synchronous events seem to have been due to ocean rather than freshwater conditions because natal rivers of these stocks were geographically disconnected. We examined year‐to‐year variability in cohort strength of one of these stocks, Upriver Bright (URB) fall Chinook salmon from the Columbia River Hanford Reach for brood years 1976–99 (recovery years 1979–2002). We used the ocean recovery rate of coded‐wire‐tag (CWT) fish as an index of cohort strength. To analyse year‐to‐year variability in the ocean recovery rate, we applied a log‐linear model whose candidate explanatory variables were ocean condition variables, fishing effort, age of recovered fish, and fish rearing type (hatchery versus wild). Explanatory variables in the best model included fishing effort, and the quadratic term of winter sea surface temperature (SST) measured from coastal waters of British Columbia, Canada during the fish's first ocean year. The coefficient of the quadratic term of SST was significantly negative, so the model shape was convex. Our findings can be used to infer year‐to‐year variability in cohort strength of other fall Chinook salmon whose life history and ocean distributions are similar to the URB fish.  相似文献   

5.
Extreme variability in abundance of California salmon populations is often ascribed to ocean conditions, yet relatively little is known about their marine life history. To investigate which ocean conditions influence their distribution and abundance, we surveyed juvenile Chinook salmon (Oncorhynchus tshawytscha) within the California Current (central California [37°30′N) to Newport, Oregon (44°00′N]) for a 2‐week period over three summers (2010–2012). At each station, we measured chlorophyll‐a as an indicator of primary productivity, acoustic‐based metrics of zooplankton density as an indicator of potential prey availability and physical characteristics such as bottom depth, temperature and salinity. We also measured fork lengths and collected genetic samples from each salmon that was caught. Genetic stock identification revealed that the majority of juvenile salmon were from the Central Valley and the Klamath Basin (91–98%). We constructed generalized logistic‐linear negative binomial hurdle models and chose the best model(s) using Akaike's Information Criterion (AIC) to determine which covariates influenced the salmon presence and, at locations where salmon were present, determined the variables that influenced their abundance. The probability of salmon presence was highest in shallower waters with a high chlorophyll‐a concentration and close to an individual's natal river. Catch abundance was primarily influenced by year, mean fork length and proximity to natal rivers. At the scale of sampling stations, presence and abundance were not related to acoustic indices of zooplankton density. In the weeks to months after ocean entry, California's juvenile Chinook salmon population appears to be primarily constrained to coastal waters near natal river outlets.  相似文献   

6.
Chinook salmon (Oncorhynchus tshawytscha) is one of several economically‐important species of salmon found in the Northeast Pacific Ocean. The first months at sea are believed to be the most critical for salmon survival, with the highest rate of mortality occurring during this period. In the present study, we examined interannual diet composition and body condition trends for late‐summer subyearling Chinook salmon caught off Oregon and Washington from 1998 to 2012. Interannual variability was observed in juvenile salmon diet composition by weight of prey consumed. Juvenile subyearling Chinook salmon were mainly piscivorous, with northern anchovy (Engraulis mordax) being especially important, making up half the diet by weight in some years. Annual diets clustered into two groups, primarily defined by their proportion of invertebrate prey (14% versus 39% on average). Diet composition was found to influence adult returns, with salmon from high‐invertebrate years returning in significantly larger numbers 2–3 yrs later. However, years that had high adult returns had overall lower stomach fullness and poorer body condition as juveniles, a counterintuitive result potentially driven by the enhanced survival of less fit individuals in better ocean conditions (top‐down effect). Ocean conditions in years with a higher percentage of invertebrates in salmon diets were significantly cooler from May to August, and bottom‐up processes may have led to a fall plankton community with a larger proportion of invertebrates. Our results suggest that the plankton community assemblage during this first fall may be critical in predicting adult returns of Chinook salmon in the Pacific Northwest.  相似文献   

7.
Salmon from different locations in a watershed can have different life histories. It is often unclear to what extent this variation is a response to the current environmental conditions an individual experiences as opposed to local‐scale genetic adaptation or the environment experienced early in development. We used a mark–recapture transplant experiment in the Shasta River, CA, to test whether life‐history traits of juvenile Chinook salmon Oncorhynchus tshawytscha varied among locations, and whether individuals could adopt a new life history upon encountering new habitat type. The Shasta River, a Klamath River tributary, has two Chinook salmon spawning and juvenile rearing areas, a lower basin canyon (river km 0–12) and upper basin spring complex (river km 40–56), characterised by dramatically different in‐stream habitats. In 2012 and 2013, we created three experimental groups: (i) fish caught, tagged and released in the upper basin; (ii) fish caught at the river mouth (confluence with the Klamath River, river km 0), tagged and released in the upper basin; and (iii) fish caught at the river mouth, tagged and released in the lower basin. Fish released in the upper basin outmigrated later and at a larger size than those released in the lower basin. The traits of fish transplanted to the upper basin were similar to fish originating in the upper basin. Chinook salmon juvenile life‐history traits reflected habitat conditions fish experienced rather than those where they originated, indicating that habitat modification or transportation to new habitats can rapidly alter the life‐history composition of populations.  相似文献   

8.
To examine the efficacy of juvenile salmon research as a tool for forecasting adult returns, the results from a study on the early marine life stage of juvenile chum salmon, conducted in the Nemuro Strait during 1999–2002 (i.e., 1998–2001 brood years), were compared with the return rates of adult salmon. Among the four brood years, the 2000 brood year (i.e., salmon migrating to the sea in 2001) was previously reported as showing higher abundance, higher growth rate and better somatic condition during the coastal residency period. Consequently, we expected it to have the highest return rate, under a hypothesis that juvenile survival in coastal residency regulates brood-year strength. Contrary to this expectation, the 2000 brood year had almost the lowest return rate. Alternatively, a statistical model in which sea surface temperature during the first year of marine life and size at release were utilized as explanatory variables reconstructed the actual variability in return rates more accurately than that based on the early marine life stage. Possible reasons for the discrepancy between the results of the juvenile salmon research and adult returns are discussed, and we suggest improvements for future research on juvenile salmon.  相似文献   

9.
We examined 1454 juvenile Chinook salmon, Oncorhynchus tshawytscha (Walbaum), captured in nearshore waters off the coasts of Washington and Oregon (USA) from 1999 to 2004 for infection by Renibacterium salmoninarum, Nanophyetus salmincola Chapin and skin metacercariae. The prevalence and intensities for each of these infections were established for both yearling and subyearling Chinook salmon. Two metrics of salmon growth, weight residuals and plasma levels of insulin-like growth factor-1, were determined for salmon infected with these pathogens/parasites, both individually and in combination, with uninfected fish used for comparison. Yearling Chinook salmon infected with R. salmoninarum had significantly reduced weight residuals. Chinook salmon infected with skin metacercariae alone did not have significantly reduced growth metrics. Dual infections were not associated with significantly more severe effects on the growth metrics than single infections; the number of triple infections was very low and precluded statistical comparison. Overall, these data suggest that infections by these organisms can be associated with reduced juvenile Chinook salmon growth. Because growth in the first year at sea has been linked to survival for some stocks of Chinook salmon, the infections may therefore play a role in regulating these populations in the Northeast Pacific Ocean.  相似文献   

10.
Chinook salmon (Oncorhynchus tschawytscha) populations within the highly modified San Francisco Estuary, California, have seen precipitous declines in recent years. To better understand this decline, a decade of coded‐wire tag release and recovery data for juvenile salmon was combined with physicochemical data to construct models that represented alternative hypotheses of estuarine conditions that influence tag recovery rate in the ocean. An information theoretic approach was used to evaluate the weight of evidence for each hypothesis and model averaging was performed to determine the level of support for variables that represented individual hypotheses. A single best model was identified for salmon released into the Sacramento River side of the estuary, whereas two competitive models were selected for salmon released into the San Joaquin River side of the estuary. Model averaging found that recovery rates were greatest for San Joaquin River releases when estuary water temperatures were lower, and salmon were released at larger sizes. Recovery rate of Sacramento releases was greatest during years with better water quality. There was little evidence that large‐scale water exports or inflows influenced recovery rates in the ocean during this time period. These results suggest that conceptual models of salmon ecology in estuaries should be quantitatively evaluated prior to implementation of recovery actions to maximise the effectiveness of management and facilitate the recovery of depressed Chinook populations.  相似文献   

11.
Juvenile salmonids display highly variable spatial and temporal patterns of early dispersal that are influenced by density‐dependent and density‐independent factors. Although juvenile coho salmon (Oncorhynchus kisutch) movement patterns in streams and their relationship with body mass and growth have been examined in previous studies, most observations were limited to one season or one stream section. In this study, we monitored the movement of juvenile coho salmon throughout their period of residence in a coastal basin to identify prevalent dispersal strategies and their relationships with body mass, growth rates and survival. Our results revealed seasonally and spatially variable movement patterns. Juvenile coho salmon that dispersed to tidally affected reaches soon after emergence remained more mobile and expressed lower site fidelity than those individuals that remained in upper riverine reaches. We did not detect significantly different growth rates between sedentary and mobile individuals. Although a greater proportion of sedentary than mobile fish survived winter to emigrate from the creek in the spring, reach of residence at the onset of winter influenced these survival estimates. Hence, apparent summer‐to‐smolt survival for mobile individuals was greater than for sedentary fish in tidally influenced reaches, whereas in riverine reaches the sedentary strategy seemed to be favoured. Our research identified complex movement patterns that reflect phenotypic and life history variation, and underscores the importance of maintaining diverse freshwater and estuarine habitats that support juvenile coho salmon before marine migration.  相似文献   

12.
Abstract Many habitat enhancement techniques aimed at restoring salmonid populations have not been comprehensively assessed. The growth and diet of juvenile Chinook salmon, Oncorhynchus tshawytscha (Walbaum), rearing in a reach designed to enhance spawning were evaluated to determine how a non‐target life stage fared in the engineered habitat. Prior work demonstrated differences in food web structure between restored and unenhanced reaches of the Merced River, thus juvenile salmon feeding dynamics were also hypothesised to vary. Dependent variables were compared among fish collected from within and near the upper boundary of the restored reach and in an unenhanced habitat upstream. Diets, otolith‐derived growth and stable isotope‐inferred trophic positions were compared. Baetidae mayflies were particularly important prey in the restored reach, while elsewhere individuals exhibited heterogeneous diets. Salmon residing at the bottom of the restored reach exhibited slightly faster growth rates relative to fish collected elsewhere, although stable isotope and diet analyses suggested that they fed at a relatively low trophic position. Specialised Baetis predation and/or abundant interstitial refugia potentially improved rearing conditions in the restored reach. Data suggest that gravel enhancement and channel realignment designed to augment adult spawning habitat may simultaneously support juvenile Chinook salmon despite low invertebrate food resources.  相似文献   

13.
Population diversity is a mechanism for resilience and has been identified as a critical issue for fisheries management, but restoration ecologists lack evidence for specific habitat features or processes that promote phenotypic diversity. Since habitat complexity may affect population diversity, it is important to understand how population diversity is partitioned across landscapes and among populations. In this study, we examined life history diversity based on size distributions of juvenile Central Valley Chinook salmon (Oncorhynchus tshawytscha) within the Yolo Bypass, a remnant transitional habitat from floodplain to tidal sloughs in the upper San Francisco Estuary (SFE). We used a generalized least squares model with an autoregressive (AR1) correlation structure to describe the distribution of variation in fish size from 1998 to 2014, and tested the effect of two possible drivers of the observed variation: (i) environmental/seasonal drivers within the Yolo Bypass, and (ii) the juvenile Chinook source population within the Sacramento River and northern SFE. We found that the duration of floodplain inundation, water temperature variation, season, and sampling effort influenced the observed time‐specific size distribution of juvenile Chinook salmon in the Yolo Bypass. Given the lack of seasonally inundated habitat and low thermal heterogeneity in the adjacent Sacramento River, these drivers of juvenile size diversification are primarily available to salmon utilizing the Yolo Bypass. Therefore, enhancement of river floodplain‐tidal slough complexes and inundation regimes may support the resilience of imperiled Central Valley Chinook salmon.  相似文献   

14.
Abstract – High‐head dams in Oregon’s Willamette River basin inhibit seaward migration and present significant mortality risks to ESA‐listed juvenile Chinook salmon (Oncorhynchus tshawytscha). Over 7 years, we passively collected 13,365 downstream‐migrating juvenile salmon in rivers above and below Willamette dams. Most salmon emigrated from upstream sites in February–June, but passed dams in November–February when reservoirs were drawn down near annual lows, and access to deep‐water passage routes improved. Samples collected above reservoirs were dominated by subyearlings, whereas below‐dam samples were a phenotypically diverse mix of subyearling, yearling and older salmon. The life history data indicated that Willamette reservoirs seasonally entrap many salmon and some sea‐ready smolts probably residualise. Annual dam‐passage mortality estimates were 8–59% (mean = 26%). Individual salmon mortality risk increased significantly with body length and varied with reservoir elevation and discharge. Operational changes that allow timely volitional emigration and development of less hazardous passage routes would benefit this threatened population.  相似文献   

15.
Macneale KH, Sanderson BL, Courbois J-YP, Kiffney PM. Effects of non-native brook trout ( Salvelinus fontinalis ) on threatened juvenile Chinook salmon ( Oncorhynchus tshawytscha ) in an Idaho stream.
Ecology of Freshwater Fish 2010: 19: 139–152. © 2009 John Wiley & Sons A/S
Abstract –  Non-native fishes have been implicated in the decline of native species, yet the mechanisms responsible are rarely apparent. To examine how non-native brook trout may affect threatened juvenile Chinook salmon, we compared feeding behaviours and aggressive encounters between these sympatric fish in Summit Creek, Idaho. Snorkelers observed 278 focal fish and examined diets from 27 fish in late summer 2003. Differences in feeding behaviours and diets suggest that there was minimal current competition for prey, although individual Chinook feeding activity declined as their encounter rate with other fish increased. While difference in size between fish generally determined the outcome of encounters (larger fish 'winning'), it was surprising that in some interspecific encounters aggressive Chinook displaced slightly larger brook trout (≤20 mm longer). We suggest that in late summer, frequent intraspecific interactions may be more important than interspecific interactions in potentially limiting Chinook growth in Summit Creek and perhaps in other oligotrophic streams where they co-occur.  相似文献   

16.
Yearling Chinook (Oncorhynchus tshawytscha) and coho salmon (Oncorhynchus kisutch) were sampled concurrently with physical variables (temperature, salinity, depth) and biological variables (chlorophyll a concentration and copepod abundance) along the Washington and Oregon coast in June 1998–2008. Copepod species were divided into four different groups based on their water‐type affinities: cold neritic, subarctic oceanic, warm neritic, and warm oceanic. Generalized linear mixed models were used to quantify the relationship between the abundance of these four different copepod groups and the abundance of juvenile salmon. The relationships between juvenile salmon and different copepod groups were further validated using regression analysis of annual mean juvenile salmon abundance versus the mean abundance of the copepod groups. Yearling Chinook salmon abundance was negatively correlated with warm oceanic copepods, warm neritic copepods, and bottom depth, and positively correlated with cold neritic copepods, subarctic copepods, and chlorophyll a concentration. The selected habitat variables explained 67% of the variation in yearling Chinook abundance. Yearling coho salmon abundance was negatively correlated with warm oceanic copepods, warm neritic copepods, and bottom depth, and positively correlated with temperature. The selected habitat variables explained 40% of the variation in yearling coho abundance. Results suggest that copepod communities can be used to characterize spatio‐temporal patterns of abundance of juvenile salmon, i.e., large‐scale interannual variations in ocean conditions (warm versus cold years) and inshore‐offshore (cross‐shelf) gradients in the abundance of juvenile salmon can be characterized by differences in the abundance of copepod species with various water mass affinities.  相似文献   

17.
Abstract – Understanding population regulation in juvenile salmonids requires distinguishing the effects of intrinsic (size, behaviour) and extrinsic (food, competition) factors. To examine the relative influence of these variables on juvenile Chinook salmon (Oncorhynchus tshawytscha) in the Salmon River drainage (ID, USA), we examined diel differences in foraging microhabitats, behaviour and consumption in two areas with consistent differences in parr‐to‐smolt survival. In lower Big Creek (LBC, high‐survival area) and upper Big Creek (UBC, low‐survival area), we observed fish by snorkelling, recording length, behaviour (foraging rate and aggression) and physical (depth, velocity, cover, temperature) and biotic (density, size and species of neighbouring fish) microhabitat features. Stomach contents were extracted to estimate consumption. Depth and temperature were greater in LBC, where Chinook salmon were significantly larger and present at lower densities. Fish in LBC exhibited higher foraging activity during the day than night, but there were no size differences between day and night foragers. In UBC, a higher density area, foraging behaviour did not change between day and night, although the smallest size classes did not forage nocturnally. Regression models that integrated physical and biotic variables suggested that physical factors influenced foraging in both areas, but competition also affected foraging in UBC. Our results demonstrate that fish from low‐ and high‐survival populations in Big Creek are exposed to different physical and biological influences during their first growth season, which are reflected in different diel foraging behaviours.  相似文献   

18.
Habitat quality often varies substantially across space and time, producing a shifting mosaic of growth and mortality trade‐offs across watersheds. Traditional studies of juvenile habitat use have emphasised the evolution of single optimal strategies that maximise recruitment to adulthood and eventual fitness. However, linking the distribution of individual behaviours that contribute to recruitment at the population level has been elusive, particularly for highly fecund aquatic organisms. We examined juvenile habitat use within a population of sockeye salmon (Oncorhynchus nerka) that spawn in a watershed consisting of two interconnected lakes and a marine lagoon. Otolith microchemical analysis revealed that the productive headwater lake accounted for about half of juvenile growth for those individuals surviving to spawn in a single river in the upper watershed. However, 47% of adults had achieved more than half of their juvenile growth in the downstream less productive lake, and 3% of individuals migrated to the estuarine environment during their first summer and returned to freshwater to overwinter before migrating back to sea. These results describe a diversity of viable habitat‐use strategies by juvenile sockeye salmon that may buffer the population against poor conditions in any single rearing environment, reduce density‐dependent mortality and have implications for the designation of critical habitat for conservation purposes. A network of accessible alternative habitats providing trade‐offs in growth and survival may be important for long‐term viability of populations.  相似文献   

19.
Linnansaari T, Keskinen A, Romakkaniemi A, Erkinaro J, Orell P. Deep habitats are important for juvenile Atlantic salmon Salmo salar L. in large rivers. Ecology of Freshwater Fish 2010: 19: 618–626. © 2010 John Wiley & Sons A/S Abstract – Juvenile Atlantic salmon were studied by underwater video surveillance and self contained underwater breathing apparatus‐diving in deep (i.e. >1.0 m), fast flowing areas of two large river systems (River Teno, River Tornionjoki) in northern Finland. Both video and diving data indicated that young‐of‐the‐year (YOY) salmon (0+) and salmon parr (>0+) readily utilised habitats deeper than 1 m. Young‐of‐the‐year salmon and parr were observed through a range of 0.5–1.9 m and 0.4–2.2 m, respectively. A negative linear relationship between the mean abundance of YOY salmon and mean depth was noted from the diving transects. Salmon parr were similarly abundant throughout the range of depths studied. Video data showed that deep habitats were used throughout the summer (June–August). It was concluded that deep, fast‐flowing areas in large rivers may constitute a significant habitat resource for juvenile salmon that has not been traditionally accounted for when estimating salmon production.  相似文献   

20.
We used the average fork length of age‐3 returning coho (Oncorhynchus kisutch) and age‐3 ocean‐type and age‐4 stream‐type Chinook (Oncorhynchus tshawytscha) salmon along the northeast Pacific coast to assess the covariability between established oceanic environmental indices and growth. These indices included the Multivariate El Niño‐Southern Oscillation Index (MEI), Pacific Decadal Oscillation (PDO), Northern Oscillation Index, and Aleutian Low Pressure Index. Washington, Oregon, and California (WOC) salmon sizes were negatively correlated with the MEI values indicating that ultimate fish size was affected negatively by El Niño‐like events. Further, we show that the growth trajectory of WOC salmon was set following the first ocean winter. Returning ocean‐type British Columbia‐Puget Sound Chinook salmon average fork length was positively correlated with the MEI values during the summer and autumn of return year, which was possibly a result of a shallower mixed layer and improved food‐web productivity of subarctic Pacific waters. Size variation of coho salmon stocks south of Alaska was synchronous and negatively correlated with warm conditions (positive PDO) and weak North Pacific high pressure during ocean residence.  相似文献   

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