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1.
The intake of forage grasses by grazing ruminants is closely related to the mechanical fracture properties of grasses. The relationship between the tensile fracture properties of grasses and foraging behaviour is of particular importance in tropical reproductive swards composed of both stems and leaves. This study (i) quantified and compared the tensile fracture properties of stems and leaves of seven tropical grass species and (ii) provided insight into the underlying plant traits that explain differences in fracture properties between species. Fracture force, tensile strength, fracture energy and toughness of stems (in various phenological stages) and leaves were measured and compared among five introduced tropical grasses (Cenchrus ciliaris, Chloris gayana, Digitaria milanjiana, Megathyrsus maximus (syn. Panicum maximum), Setaria sphacelata) and two native tropical grasses (Setaria surgens and Dichanthium sericeum). Species differed significantly in fracture force and fracture energy, with stems and leaves of C. ciliaris and S. surgens requiring less force and energy to fracture and stems and leaves of M. maximus and S. sphacelata requiring more force and energy to fracture in comparison with the other species. Differences in tensile strength and toughness were less pronounced. The differences among species in fracture force and energy mainly resulted from differences in cross‐sectional area of plant parts rather than from differences in tensile strength and toughness.  相似文献   

2.
The leaves of tropical grasses are eaten in larger quantities than stems of similar DM digestibility. To determine whether the same fact applied to a temperate grass, studies were made with leaf and stem fractions of Lolium perenne. The leaves of L. perenne had a 20% higher voluntary intake than the stems, white the DM digestibilities were only slightly higher (67·3%ν. 64·8%). The leaves also had a larger surface area/g, a higher initial (12 h) fermentation rate in vitro, larger content of pepsin-soluble DM, and a smaller energy require ment to grind I g DM in a laboratory mill compared with the stems. The difference in voluntary intake between the leaves and stems of L. perenne was smaller than that for a number of tropical grasses.  相似文献   

3.
The concentration of water‐soluble carbohydrates (WSC) of cocksfoot is lower than that of other temperate grasses. Increasing the WSC concentration in cocksfoot is important in increasing its digestibility and preference by ruminants. The genetic variation in mono‐ and disaccharides, fructans and total WSC concentrations and their inter‐relationships with crude protein (CP) and fibre concentrations of cocksfoot (Dactylis glomerata L.) at the vegetative growth stage in half‐sib cross populations were assessed in two experiments conducted under spaced planting and sward conditions. There was a wide range in the means for concentrations of fructan, WSC and neutral‐detergent fibre (NDF) in parents and progeny whereas there was a narrow range of the means for concentrations of mono‐ and disaccharides, CP and acid‐detergent fibre (ADF). Mean concentrations of mono‐ and disaccharides showed the smallest range. Mean squares of entries in parents and progenies were significant for mono‐ and disaccharides, fructan and total WSC concentrations in all populations. The entry × year interactions for fructan and total WSC concentrations were significant in the parents and the half‐sib progeny. However, the entry × year interactions for mono‐ and disaccharide concentrations in progeny were not significant. Concentration of WSC was under genetic control as mean squares of the concentration values were significant and variance components for all traits were significantly larger than zero. There were negative genetic correlations between WSC and ADF and NDF concentrations. Narrow‐sense heritabilities (hn) estimated from variance components of progeny for total WSC concentrations were 0·59 and 0·53 in sward and spaced planting conditions respectively. The hn of fructan and WSC concentrations were similar in both sward and spaced planting conditions, whereas that of mono‐ and disaccharide concentrations varied from 0·20 to 0·69. It was found that the genetic variation of the total WSC concentration in cocksfoot depended mainly on genetic variation in fructan concentration. These results suggested that the forage quality of cocksfoot at the vegetative growth stage was influenced by an additive gene effect and could be improved genetically by recurrent selection.  相似文献   

4.
To optimize yields of long‐chain fructans extracted from herbage, fructan concentrations and chromatographic profiles were compared in four cool‐season grasses extracted by different methods. In a preliminary study, extraction temperatures, pH prior to extract concentration, numbers of extractions, and tissue coarseness were varied to evaluate their effects on chromatographic profiles and quantities of total long‐chain fructan (degree of polymerization (DP) of ~7 or higher). Fructans were separated by anion‐exchange high‐performance liquid chromatography (HPLC) with pulsed amperometric detection (PAD). The identity of the putative fructan peaks was confirmed by acid hydrolysis of fructans to fructose. Tall fescue and bluegrass fructans were below the detection limits used in this study. Pooled timothy and cocksfoot data indicated that two or three extractions at the same temperature yielded similar amounts of fructans and that adjusting pH from 6 to 7 did not affect recovery. Replicated extractions of cocksfoot by four methods chosen from the preliminary study demonstrated that the least fructan was extracted from chopped cocksfoot at ambient temperature (P = 0·019). One C18 solid‐phase extraction cleanup step sufficed for analysis. The extraction method recommended is two boiling water extractions of either chopped or ground material. However, ground material extracted at ambient temperature is comparable in total fructan concentration and maximum DP.  相似文献   

5.
Grazing lands in warm‐temperate and subtropical North America have become less diverse. Pastures are typically grass monocultures, while rangelands are generally managed for the grass components. Overstocking, selective herbicides, fire exclusion and heavy rates of nitrogen fertilizer have contributed to near exclusion of native, warm‐season legumes. The simplicity of managing grass monocultures, pasture production responses to nitrogen fertilizer and profitability of grass‐only systems have limited interest in legume‐based approaches. Changing economics and ecological concerns with ecosystem accumulation of industrial inputs contribute to an increasing interest in legumes. Unlike the development of temperate pasture legumes and recent research in the tropics, legumes tolerant of both freezing temperatures and hot weather have received less attention. Poor establishment, limited persistence and potential invasiveness limit currently available introduced species. Native, herbaceous, warm‐season legume species occur throughout warm‐temperate North America, but little attention has been directed to these plants as potential forage species. Some success with a few native legume species, primarily in the genus Desmanthus, suggests potential for expanded assessment of forage value of the many species available. Current assessments of native legumes, primarily for conservation purposes, provide an opportunity to expand evaluations of these species for pasture and rangeland potential while economics of livestock production and public interest in ecosystem health are supportive. Experiences with legumes of warm‐temperate origin in North America, along with results with temperate and tropical pasture legumes globally, provide a starting point for future efforts at incorporating greater legume diversity in pastures and rangelands of subtropical and warm‐temperate regions around the world.  相似文献   

6.
Tall fescue (Festuca arundinacea) is a Eurasian forage grass extensively planted in the United States. However, an endophytic fungus in tall fescue, Epichloë coenophiala, causes health problems in cattle. We predicted that cattle prefer to graze alternative forages when available. We also predicted that cattle use tall fescue more intensively in recently burned areas, as fire can increase forage quality. We tested these predictions in four diverse‐forage pastures in Iowa, comparing use by cattle of tall fescue and four alternative forages (non‐fescue cool‐season grasses, native warm‐season grasses, non‐leguminous forbs and legumes) to their availabilities at the pasture scale. We also examined how tall fescue influences the distribution of grazing at a fine scale (0.1‐m2 quadrats). Tall fescue was the most abundant forage (46% of plants), but composed only 26% of grazed vegetation. In contrast, legumes composed 12% of available forage but 25% of grazed vegetation. Other forages were used in proportion to availability. At a fine scale, total grazing frequency (proportion of plants grazed) was lower in quadrats containing abundant tall fescue, and higher in quadrats with abundant warm‐season grasses. Grazing frequency of tall fescue and other cool‐season grasses was greatest in recently burned quadrats, but total grazing frequency did not increase after burning. Our results show that although cattle graze tall fescue, particularly following burns, they limit their use of this grass. Given that tall fescue is underused, creates health risks for cattle, and degrades wildlife habitat quality, it may be advisable to reduce tall fescue in pastures.  相似文献   

7.
8.
Mediterranean forage systems suffer from limited availability of fresh forage because of water deficits and extreme temperatures. Consequently, fresh forage is unavailable for at least 6–7 months a year, and farmers must buy feed to support livestock production. With the aim of overcoming these limitations, a 2‐year trial was conducted on three distinct sites in Sicily (at 10, 600 and 1200 m elevation) with thirty‐four varieties of forage species belonging to nine biennial/perennial and thirteen annual species. Results showed that by integrating grasses and legumes, species from environments with different climatic conditions enable the season of forage production to be extended from mid‐April to mid‐November. Quality traits of forage in different areas varied in relation to species and varieties. In general, the sown‐forage quality was better than in pastures and fallows in the same areas commonly used to feed animals. This also leads to a reduction in the use of supplementary feeds. Among the tested species, Lolium multiflorum and Medicago sativa emerged as the most promising for filling the forage‐deficit periods, and Trifolium spp. and Vicia sativa were found to be superior for increasing forage quality. The results are discussed in the context of adapting Mediterranean forage supplies for ensuring greater sustainability of livestock production in mountain, hill and plain areas. The proposed forage chain arrangement represents part of local potential adaptation to climate limitations and climate change.  相似文献   

9.
Cereal-based products, such as bread, are staple foods in the western diet. Due to the nature of their basic ingredients and the diversity of recipes, the amount of fermentable short-chain carbohydrates (FODMAPs) in those products may be high. This study characterized the FODMAP-profiles of a broad range of cereal-product ingredients, serving as a basis for low FODMAP product development. Different cereals, pseudo-cereals, gluten-free flours, pulses, pulse protein ingredients, commercial sprouts, and other cereal-product ingredients were analyzed, using anion-exchange chromatography with electrochemical detection. Wheat and related cereals were high in fructans. Pulses, such as peas contained high galactooligosaccharides (GOS) amounts. Whereas GOS levels in pulse protein ingredients varied, depending on their production. Gluten-free flours, for instance, rice-flour, showed low FODMAP-profiles. Amongst those, buckwheat, which does not contain any of the FODMAPs investigated, contained high amounts of other soluble non-digestible carbohydrates, namely fagopyritols; these may have a similar effect on a sensitive gut as GOS. Finally, ingredients contained mainly high levels of fructans and GOS. Yet, the analysis of commonly consumed commercial cereal products, including bread, pasta, crackers and biscuits, highlighted the relevance of lactose, fructose in excess of glucose and polyols. These products serve as benchmarks for further product development.  相似文献   

10.
The digestible crude protein percentage (DCP) of 218 different feeds derived from 16 tropical grasses and 8 tropical legumes containing 2·2 to 25·3% crude protein (CP) was Treasured with sheep. DCP was closely related to CP (r =· 0·979) and could be predicted from the equation DCP=O·899 CP-3·25 (SE estimate · 0·84). Predicted values for tropical grasses and legumes were similar to, but more variable than, those calculated from a similar equation derived from temperate pasture species. It was suggested that CP per se is a useful criterion for selecting tropical pasture plants when direct in vivo measurements of DCP cannot be obtained.  相似文献   

11.
Seed mass and shape of grasses were analysed in a temperate flora containing 178 and 26 species belonging to C3 and C4 photosynthetic types respectively. The weedy character and the annual or perennial status were also considered. On the basis of the seed traits studied, three groups were distinguished: C4 grasses, annual C3 grasses of weedy character and perennial non‐weedy C3 grasses. The C4 group had more isodiametric (same diameter in all directions) seed shape and lower average seed mass than the C3 group. To our knowledge, this has not yet been described for temperate C4 grasses and is certainly associated with their preference for open habitats where competition for light is small. Weedy annual C3 grasses had heavier and less isodiametric seeds than C4 grasses did. These species are mostly specialized to establish in the dense cover of perennial vegetation, and this ability distinguishes them from the C4 group. Non‐weedy perennial C3 species possessed less isodiametric seeds than did C4 grasses, but did not differ from weedy annual C3 grasses. As most alien C4 grasses naturalized in Hungary are annuals with small, isodiametric seeds, these traits are good candidates to be included in screening for potential future invasives in open habitats.  相似文献   

12.
Provision of an adequate mineral supply in the diets of ruminants fed mainly on grassland herbage can present a challenge if mineral concentrations are suboptimal for animal nutrition. Forage herbs may be included in grassland seed mixtures to improve herbage mineral content, although there is limited information about mineral concentrations in forage herbs. To determine whether herbs have greater macro‐ and micromineral concentrations than forage legumes and grasses, we conducted a 2‐year experiment on a loamy‐sand site in Denmark sown with a multi‐species mixture comprised of three functional groups (grasses, legumes and herbs). Herb species included chicory (Cichorium intybus L.), plantain (Plantago lanceolata L.), caraway (Carum carvi L.) and salad burnet (Sanguisorba minor L.). We also investigated the effect of slurry application on the macro‐ and micromineral concentration of grasses, legumes and herbs. In general, herbs had greater concentrations of the macrominerals P, Mg, K and S and the microminerals Zn and B than grasses and legumes. Slurry application indirectly decreased Ca, S, Cu and B concentrations of total herbage because of an increase in the proportion of mineral‐poor grasses. Our study indicates that including herbs in forage mixtures is an effective way of increasing mineral concentrations in herbage.  相似文献   

13.
The objective was to compare alternative models for fitting to data on degradation of dry matter (DM), organic matter (OM) and crude protein (CP) in the rumen of two harvests of four temperate grasses, made into hay in Belgium, and one harvest of three tropical grasses commonly used in Burundi. Fifteen non-linear models were used and these included generalized Mitscherlich, simple Mitscherlich (or exponential), generalized Michaelis–Menten, Gompertz, logistic, simple Michaelis–Menten, segmented Van Milgen and von Bertalanffy models. Degradation profiles of DM, OM and CP obtained in sacco from the rumen of three mature sheep were fitted to all the models. The convergence success rate along with acceptable parameters was used as a first tool to eliminate models. Comparisons between the remaining twelve models were then made using the run test of sign, the root mean square prediction error, the root mean square error, the lack of fit test and the variance ratio test criteria. On the basis of an overall assessment using these tests, the Gompertz lagged model was the best suited to fit the degradation data of both temperate and tropical grasses. This study confirmed the superiority of sigmoid type functions over diminishing return-type models.  相似文献   

14.
Enhancing pasture persistence is crucial to achieve more sustainable grass‐based animal production systems. Although it is known that persistence of perennial ryegrass is based on a high turnover of tillers during late spring and summer, little is known about other forage species, particularly in subtropical climates. To address this question, this study evaluated survival of grazed tall fescue tillers growing in a subtropical climate. We hypothesized that hard tactical grazing during winter to remove reproductive stems (designated as ‘flowering control’), and nitrogen fertilization in spring, would both improve tiller survival over summer, and thus enhance tiller density. This was assessed in two experiments. In both experiments, few tillers appeared during late spring and summer and so tiller density depended on the dynamics of vegetative tillers present in the sward in spring. In Experiment 2, flowering control and nitrogen fertilization both enhanced the survival of that critical tiller cohort, but the effects were not additive. Responses were similar but not statistically significant in Experiment 1, which had a warmer, drier summer and lower overall survival rates. Unlike grasses in temperate environments, persistence of tall fescue in this subtropical site appeared to follow a ‘vegetative pathway’; i.e., new tillers were produced largely in autumn, from vegetative tillers that survived the summer.  相似文献   

15.
In the moist mid‐latitudes of eastern Australia, soil water dynamics, herbage production and water use efficiency (WUE) were monitored during 2006–2008, for five perennial pastures: digit grass (Digitaria eriantha), Rhodes grass (Chloris gayana), forest bluegrass (Bothriochloa bladhii), native grass (Bothriochloa macra and Rytidosperma bipartita dominant), lucerne (Medicago sativa); and two forage crops: oat (Avena fatua) and sorghum (Sorghum bicolor). Ground cover formed more quickly in Rhodes grass and lucerne (>70% ground cover in 120 and 175 days after sowing [DAS] respectively) than in forest bluegrass and digit grass (245 and 365 DAS respectively). Values of maximum extractable water (MEW) for Rhodes grass and lucerne were similar (180–242 mm), while values for digit grass and forest bluegrass (129–175 mm) were equal to or greater than those for native grass, and two annual forage crops (77–144 mm). Lucerne expressed the maximum root depth (1.46 m), while values for the tropical grasses (0.96–1.39 m) were greater than native grasses and forage crops (0.87–0.96 m). Native grasses (6.5–12 t DM/ha) had the lowest herbage production, which resulted in values of WUE that were significantly less than most other treatments (16–21 vs. 23–43 kg DM ha?1 mm?1). Digit grass (33–34 kg DM ha?1 mm?1) had higher WUE compared with the other tropical grasses (20–27 kg DM ha?1 mm?1). The data collected here suggest that a forage system comprising digit grass, lucerne and forage oat would provide high production and WUE in this environment.  相似文献   

16.
Changes in livestock production systems have led to land‐use changes and abandonment, especially of semi‐natural grassland in agriculturally less favoured regions. The generation of energy from biomass of extensive, high‐diversity grasslands can be an alternative to their abandonment, and anaerobic digestion is one possible method for converting grassland biomass into energy. However, little is known about the effects of species richness (SR) and functional groups on chemical constituents relevant for anaerobic digestion and the resulting energy potential. In this study, changes in the herbage chemical constituents that are relevant for forage quality were studied along a well‐defined diversity gradient (one to sixty species) and across different combinations of functional groups (legumes, small herbs, tall herbs and grasses). Substrate‐specific methane yield (CH4 sub) was estimated through the concentrations of forage‐quality parameters such as crude fibre (CF), crude protein (CP), crude lipid, nitrogen‐free extract and their documented digestibility values, as well as the respective methane yields. Results show that with increasing SR, the CF increased and CP decreased, even though these effects could not be fully disentangled from the presence of grasses. These trends led to a negative effect of SR on CH4 sub, while the area‐specific methane yield (CH4 area = CH4 sub × biomass yield) increased due to a strong increase in biomass with increasing SR. The CH4 sub was increased when legumes were present, and it declined with the presence of grasses. Generally, CH4 sub and CH4 area varied between functional‐group monocultures and all functional‐group mixtures.  相似文献   

17.
Protein degradability in forage legumes is of global importance because utilization efficiency of forage has economic and environmental consequences. However, there are no published studies on the effect of legume stand structure on differences in crude protein (CP) fractions. The main objective of the present research was therefore to investigate differences in CP fractions in leaves and stems of lucerne (Medicago sativa L.) during the growing season. Stand traits were measured over 2 years, and forage was sampled at the early bud and early flower stages in the first, second and third cuts. Stems had significantly higher concentrations (in g kg?1 CP) of non‐protein (fraction A: 430 g kg?1 CP) and indigestible nitrogen (fraction C: 92 g kg?1 CP) than leaves and had lower relative content of true protein (fraction B: 478 g kg?1 CP). In the total forage (stems and leaves combined), about 80% of the variation in CP fractions was explained by year, cut and maturity. Year was the most important factor, particularly for the B fractions. Cut was the second‐most important factor; its main effect was that the relative abundance of fraction A declined from 394 g kg?1 CP in the first cut to 293 in g kg?1 CP the third cut. Maturity increased the amounts of indigestible fraction C and protein fractions B1 and B3. This was associated with the leaf weight ratio, which had an inverse relationship with maximal stem length and dry matter yield. Variation partitioning showed that 75% of CP fraction variability associated with cut, maturity and year could be explained by the evaluated stand traits. This research has highlighted the need to consider plant morphological traits when legume CP fractions are evaluated.  相似文献   

18.
Atmospheric carbon dioxide (CO2), global mean temperature and interannual variability in temperature and rainfall are expected to increase significantly by the end of the 21st century. To review the effects of these factors on forage quality, we carried out a meta-analysis of climate manipulation experiments. The first notable result was a lack of effect of elevated CO2 on structural carbohydrates and digestibility. Elevated CO2 increased the total non-structural carbohydrates of forage tissues by an average of 25% and decreased forage nitrogen (N) content by 8%. Increased legume abundance in multispecies swards can, however, maintain N concentration in the harvested biomass. There were no consistent effects of warming on contents of N, water-soluble and structural carbohydrates, or on digestibility. We highlight the continuum in the effect of water availability, from drought to irrigation, with a curvilinear increase of forage N as water availability decreased. Digestibility increased, on average, by 7% with drought, but with strong experimental variations. The review places special emphasis on discussion of the specificities of mountain and Mediterranean grasslands, the former being limited by low temperature, the latter by drought and heat. Elevated CO2 decreased forage N content in mountain areas and in temperate plains alike. It increased N content by an average of 3% in Mediterranean areas; this could be due to shifts in vegetation communities under elevated CO2 or to a greater concentration of N in plant tissues under drought conditions. Further experiments are needed to investigate the effects of combined factors, including extreme climatic events.  相似文献   

19.
This is the first study describing the fine structure of the main, individual fructan oligosaccharides present in wheat grains. Wheat grain fructan structure was investigated in developing wheat grains and in different tissues of mature grains with liquid chromatography-mass spectrometry. Fructan oligosaccharides with a low degree of polymerization (<5) were mainly of the graminan- and inulin-type in developing wheat grains during the first week after anthesis. Starting from 14 days after anthesis, neo-type fructans, fructans with an internal glucose, were observed for the first time. Several neo-type fructan structures were identified and their portion in the total fructan pool gradually increased during grain development. In the mature kernel, almost no differences were noted between the fructan distributions of wheat flour and two wheat bran fractions enriched in either pericarp or aleurone tissue. Results are related to wheat fructan metabolizing enzymes and the nutritional implications are discussed.  相似文献   

20.
Development of a standard evaluation protocol has been a pressing problem for the selection of drought‐resistant genotypes of tall fescue (Festuca arundinacea). This study was conducted to evaluate the association of forage yield with specific phenological and morphological traits to find a proper model for indirect selection under irrigated (normal) and drought‐stress conditions in tall fescue. A random sample of seventy‐five genotypes were clonally propagated and evaluated in normal and drought‐stress environments in the field during 2009 and 2010. Results showed that water stress had a negative effect on forage yield and most of the morphological traits measured and reduced genotypic variation for most of them. Forage yield had the highest genotypic variation, whereas days to pollination had the lowest variation. Low broad‐sense heritability estimates were obtained for dry‐matter yield, but heritability for the traits of number of stems per plant, plant height and crown diameter was moderately high. These traits were identified as the main components of forage yield. The importance of these components and their direct and indirect effects on forage yield was different in normal and drought‐stress conditions. This suggests that indirect selection for developing high‐yielding, drought‐tolerant varieties should be performed under drought‐stress conditions with a specific model.  相似文献   

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