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1.
Improving our knowledge of pre-anthropogenic landscapes is vital for understanding landscape-scale heterogeneity and for setting goals and objectives for ecological restoration. This is especially important in highly modified landscapes that contain few remnants of pre-impact ecosystems. This study aims to develop new methodology to improve understanding of historical vegetation, using the now-degraded inhabited highlands of the Galapagos Islands as a case study. Our multidisciplinary approach innovatively combines data from interviews with residents who were familiar with the vegetation before most degradation occurred with the more traditional sources of historical aerial photography and information from early explorer and scientist reports. We reconstruct historical vegetation across the landscape by mapping it in the year 1960 and discussing this map in the historical context of anthropogenic change. Our results confirm published vegetation types but also define some other types not previously described, and suggest much greater spatial, temporal and structural heterogeneity than commonly understood. This result can be used by Galapagos land managers to better match species assemblages with sites and plan restoration actions that will maximise resilience against the ongoing and future threats of climate change and species invasions. Our methodology can be applied in extensive areas of the world where the majority of anthropogenic disturbance to natural ecosystems has been within the past 60 years.  相似文献   

2.
Effects of landscape patterns on biotic communities   总被引:4,自引:0,他引:4  
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3.
We address effects of large-scale forestry on landscape structure and the structure and composition of boreal bird communities in North Sweden. Specifically, we ask: after controlling for the effect of patch size, forest age and tree species composition, is there any residual effect attributable to the reduction in area of old forest? Pairs of landscape blocks (25 by 25 km) were selected to maximize area difference in human-induced disturbance, clear-cut as opposed to semi-natural old forest. Median distance to natural edge (wetlands, open water) from randomly selected points in forest was 250 and 200 m in high and low impact landscapes, respectively, indicating a high degree of ‘natural’ fragmentation of the pristine boreal landscape in the area. By contrast, median distance to clear-cut in uncut forest was 750 and 100 m, respectively. Clear-cuts in high impact landscapes were disproportionally more common in areas with contiguous forest land than in areas with spatially disjunct forest, implicating that forestry increases natural fragmentation of the landscape by subdividing larger forest tracts. Point counts along forestry roads showed that species richness and relative abundance of forest birds were higher in landscapes with low forestry impact. These differences can partly be explained by differences in age composition of forest and composition of tree species. After controlling for patch size, forest age and tree species composition, a significant effect of forestry impact remained for Sibirian species and the Tree pipitAnthus trivialis. Our results thus imply that this group of species and the Tree pipit may be sensitive to forest fragmentation. In contrast to previous Finnish studies, we found relatively small negative effects on relative abundance of species hypothesized to be negatively affected by large-scale clear-cutting forestry. However, our picture of the present does not contradict results from Finnish long-term population studies. Five factors may account for this: 1) clear-cut areas are not permanently transformed into other land use types, 2) planted forests are not completely inhabitable for species preferring older forest, 3) the majority of species in the regional pool are habitat generalists, 4) the region studied is still extensively covered with semi-natural forest, and 5) our study area is relatively close to contiguous boreal forest in Russia, a potential source area for taiga species.  相似文献   

4.
Landscape context can influence species richness, abundance, or probability of patch-use by birds. Little is known, however, about the effects of landscape context on birds in wetland-dominated landscapes. This lack of knowledge is alarming because many wetlands are threatened by development and other human impacts, while serving critical functions as migratory, breeding and foraging habitat. To address this lack of knowledge, we censused birds in North American Great Lakes coastal wet meadows located along the northern Lake Huron shoreline in Michigan (USA) during 1997 and 1998. Using a suite of multivariate techniques, we first accounted for effects of area and within-patch habitat characteristics before testing for effects of landscape context. Most bird variables were significantly related to landscape context, and two major patterns were apparent. First, avian species richness, abundance, and probability of patch-use by some species were higher for wet meadows located in complex contexts (adjacent to many patch types) compared to simpler contexts (adjacent to only one patch type). Second, these variables were higher for wet meadows located in wetland contexts compared to contexts that were terrestrial and road-impacted, dominated by open water habitats, or dominated by forested wetland habitats. Conservation plans for wetlands have focused on saving large wetlands and creating the vegetative habitat structure required by birds, but they should go further and explicitly consider the landscape context of wetlands as well. Specifically, wetlands located in complex and/or wetland contexts should have a higher conservation value than similar wetlands located in simpler, more terrestrial contexts. This revised version was published online in August 2006 with corrections to the Cover Date.  相似文献   

5.
A central theme in landscape ecology is that of understanding the consequences of landscape heterogeneity for ecological processes. The effects of landscape heterogeneity on parasite communities are poorly understood, although it has been shown that anthropogenic impacts may contribute to outbreaks of both parasites and pathogens. We tested for effects of landcover type, composition, configuration, and urbanisation on avian diversity and avian malaria prevalence in 26 communities of wetland-associated passerines in the Western Cape of South Africa. We predicted that avian malaria prevalence would be influenced by the pattern of farmland and urban areas in the surrounding landscapes and the sizes of the wetlands in which birds were sampled. We quantified landscape pattern using a six-class simplification of the National Landcover data set at 35 × 35 m resolution and five extents of between 1 and 20 km from each wetland. The bird community was sampled using point counts and we collected blood samples from birds at each site. We screened these for malaria using PCR and molecular techniques. Passerine species richness and infection prevalence varied significantly between different landcover types. Host richness and parasite prevalence were highest in viticultural and cropping sites respectively and lowest in urban sites. Wetlands located in indigenous vegetation had intermediate numbers of bird species and intermediate parasite prevalence. Landscape composition and habitat type surrounding wetlands emerged as useful correlates of infection prevalence. Anthropogenic landscape modification appears to have both direct and indirect effects on avian communities and their associated parasite assemblages, with attendant consequences for avian health.  相似文献   

6.
We evaluated the influence of scale on habitat use for three wetland-obligate bird species with divergent life history characteristics and possible scale-dependent criteria for nesting and foraging in South Dakota, USA. A stratified, two-stage cluster sample was used to randomly select survey wetlands within strata defined by region, wetland density, and wetland surface area. We used 18-m (0.1 ha) fixed radius circular-plots to survey birds in 412 semipermanent wetlands during the summers of 1995 and 1996. Variation in habitat use by pied-billed grebes (Podilymbus podiceps) and yellow-headed blackbirds (Xanthocephalus xanthocephalus), two sedentary species that rarely exploit resources outside the vicinity of nest wetlands, was explained solely by within-patch variation. Yellow-headed blackbirds were a cosmopolitan species that commonly nested in small wetlands, whereas pied-billed grebes were an area-sensitive species that used larger wetlands regardless of landscape pattern. Area requirements for black terns (Chlidonias niger), a vagile species that typically forages up to 4 km away from the nest wetland, fluctuated in response to landscape structure. Black tern area requirements were small (6.5 ha) in heterogeneous landscapes compared to those in homogeneous landscapes (15.4–32.6 ha). Low wetland density landscapes composed of small wetlands, where few nesting wetlands occurred and potential food sources were spread over large distances, were not widely used by black terns. Landscape-level measurements related to black tern occurrence extended past relationships between wetlands into the surrounding matrix. Black terns were more likely to occur in landscapes where grasslands had not been tilled for agricultural production. Our findings represent empirical evidence that characteristics of entire landscapes, rather than individual patches, must be quantified to assess habitat suitability for wide-ranging species that use resources over large areas.  相似文献   

7.
A meaningful interpretation of landscape metrics is possible only when the limitations of each measure are fully understood, the range of attainable values is known, and the user is aware of potential shifts in the range of values due to characteristics of landscape patches. To examine the behavior of landscape metrics, we generated artificial landscapes that mimicked fragmentation processes while controlling the size and shape of patches in the landscape and the mode of disturbance growth. We developed nine series of increasingly fragmented landscapes and used these to investigate the behavior of edge density, contagion, mean nearest neighbor distance, mean proximity index, perimeter-area fractal dimension, and mass fractal dimension. We found that most of the measures were highly correlated, especially contagion and edge density, which had a near-perfect inverse correspondence. Many of the measures were linearly-associated with increasing disturbance until the proportion of disturbance on the landscape was approximately 0.40, with non-linear associations at higher proportions. None of the measures was able to differentiate between landscape patterns characterized by dispersed versus aggregated patches. The highest attainable value of each measure was altered by either patch size or shape, and in some cases, by both attributes. We summarize our findings by discussing the utility of each metric.  相似文献   

8.
The conversion of forests and farmlands to human settlements has negative impacts on many native species, but also provides resources that some species are able to exploit. American Crows (Corvus brachyrhynchos), one such exploiter, create concern due to their impact as nest predators, disease hosts, and cultural harbingers of evil. We used various measures of crow abundance and resource use to determine crows’ response to features of anthropogenic landscapes in the Puget Sound region of the United States. We examined land cover and land use composition at three spatial scales: study sites (up to 208 ha), crow home ranges within sites (18.1 ha), and local land cover (400 m2). At the study site and within-site scales crow abundance was strongly correlated with land cover providing anthropogenic resources. In particular, crows were associated with the amount of ‘maintained forest’ cover, and were more likely to use grass and shrub cover than forest or bare soil cover. Although crows did not show a generalized response to an edge variable, they exhibited greater use of patchy habitat created by human settlements than of native forests. Radio-tagged territorial adults used resources within their home ranges relatively evenly, suggesting resource selection had occurred at a larger spatial scale. The land conversion pattern of new suburban and exurban settlements creates the mix of impervious surfaces and maintained vegetation that crows use, and in our study area crow populations are expected to continue to increase. Electronic supplementary material  The online version of this article (doi:) contains supplementary material, which is available to authorized users.  相似文献   

9.
Information on vegetation-related land cover change and the principle drivers is critical for environmental management and assessment of desertification processes in arid environments. In this study, we investigated patch-level based changes in vegetation and other major land cover types in lower Tarim River drainage area in Xinjiang, West China, and examined the impacts of environmental factors on those changes. Patterns of land cover change were analyzed for the time sequence of 1987–1999–2004 based on satellite-derived land classification maps, and their relationships with environmental factors were determined using Redundancy Analysis (RDA). Environmental variables used in the analysis included altitude, slope, aspect, patch shape index (fractal dimension), patch area, distance to water body, distance to settlements, and distance to main roads. We found that during the study period, 26% of the land experienced cover changes, much of which were the types from the natural riparian and upland vegetation to other land covers. The natural riparian and upland vegetation patches were transformed mostly to desert and some to farmlands, indicating expanding desertification processes of the region. A significant fraction of the natural riparian and upland vegetation experienced a phase of alkalinity before becoming desert, suggesting that drought is not the exclusive environmental driver of desertification in the study area. Overall, only a small proportion of the variance in vegetation-related land cover change is explainable by environmental variables included in this study, especially during 1987–1999, indicating that patch-level based vegetation change in this region is partly attributable to environmental perturbations. The apparent transformation from the natural riparian and upland vegetation to desert indicates an on-going process of desertification in the region.  相似文献   

10.
For some time, ecologists have known that spatial patterns of forest structure reflected disturbance and recovery history, disturbance severity and underlying influences of environmental gradients. In spite of this awareness, historical forest structure has been little used to expand knowledge of historical fire severity. Here, we used forest structure to predict pre-management era fire severity across three biogeoclimatic zones in eastern Washington State, USA, that contained extensive mixed conifer forests. We randomly selected 10% of the subwatersheds in each zone, delineated patch boundaries, and photo-interpreted the vegetation attributes of every patch in each subwatershed using the oldest available stereo-aerial photography. We statistically reconstructed the vegetation of any patch showing evidence of early selective harvesting, and then classified them as to their most recent fire severity. Classification used published percent canopy mortality definitions and a dichotomized procedure that considered the overstory and understory canopy cover and size class attributes of a patch, and the fire tolerance of its cover type. Mixed severity fires were most prevalent, regardless of forest type. The structure of mixed conifer patches, in particular, was formed by a mix of disturbance severities. In moist mixed conifer, stand replacement effects were more widespread in patches than surface fire effects, while in dry mixed conifer, surface fire effects were more widespread by nearly 2:1. However, evidence for low severity fires as the primary influence, or of abundant old park-like patches, was lacking in both the dry and moist mixed conifer forests. The relatively low abundance of old, park-like or similar forest patches, high abundance of young and intermediate-aged patches, and widespread evidence of partial stand and stand-replacing fire suggested that variable fire severity and non-equilibrium patch dynamics were primarily at work.  相似文献   

11.
Neutral models for the analysis of broad-scale landscape pattern   总被引:47,自引:19,他引:28  
The relationship between a landscape process and observed patterns can be rigorously tested only if the expected pattern in the absence of the process is known. We used methods derived from percolation theory to construct neutral landscape models,i.e., models lacking effects due to topography, contagion, disturbance history, and related ecological processes. This paper analyzes the patterns generated by these models, and compares the results with observed landscape patterns. The analysis shows that number, size, and shape of patches changes as a function of p, the fraction of the landscape occupied by the habitat type of interest, and m, the linear dimension of the map. The adaptation of percolation theory to finite scales provides a baseline for statistical comparison with landscape data. When USGS land use data (LUDA) maps are compared to random maps produced by percolation models, significant differences in the number, size distribution, and the area/perimeter (fractal dimension) indices of patches were found. These results make it possible to define the appropriate scales at which disturbance and landscape processes interact to affect landscape patterns.  相似文献   

12.
It is widely accepted that large protected areas are required to effectively conserve historical species composition. However, recent analyses of mammal species loss in Canadian and African national parks contradict earlier conclusions that extent of local extinctions (i.e., extirpations) is strongly inversely related to park size, suggesting that park size alone is inadequate to predict reserve designs that may sustain biodiversity. To plan protected areas that will meet conservation goals, reserve-design models that incorporate other landscape-scale factors in addition to reserve area are needed; potential factors include the types and intensity of land use and habitat change, together with land cover types, in and around parks. Additionally, human population size around parks, and visitor density in parks may affect species loss. We quantified land use, land cover, and human population in and around 24 Canadian national parks to model effects of human disturbance and changes in natural habitats on known mammal extirpations.Multiple regression models were compared using the Akaike Information Criterion (AICc). The most parsimonious model (AICc weighting w i = 0.5391) emphasized effective habitat area in and around parks and not visitor numbers nor human population size around parks. Our model suggests that parks with as little as 3140 km2 of effective habitat area inside may be large enough to conserve historical mammal species composition if they are also surrounded by at least 18 000 km2 of effective habitat within 50 km of park boundaries.  相似文献   

13.
We formulated and tested models of relationships among determinants of vegetation cover in two agroforested landscapes of eastern North America (Haut Saint-Laurent, Quebec, Canada) that differed by the spatial arrangement of their geomorphic features and intensity of agricultural activities. Our landscape model compared the woody plots of each landscape in terms of the relative influence of environmental attributes, land use history (1958 – 1997), and spatial context (i.e., proximity of similar or contrasting land cover). Our vegetation model evaluated the relative contribution of the same sets of variables to the distributions of herbs, trees, and shrubs. Relationships were assessed using partial Mantel tests and path analyses. Significant environmental and contextual differences were found between the vegetation plots of the two landscapes, but disturbance history was similar. Our vegetation model confirms the dominant effect of historical factors on vegetation patterns. Whereas land-use history overrides environmental and contextual control for trees, herbaceous and shrub species are more sensitive to environmental conditions. Context is determinant only for understory species in older, less-disturbed plots. Results are discussed in relevance to vegetation dynamics in a landscape perspective that integrates interactions between environmental and human influences.  相似文献   

14.
Mapping and analyzing landscape patterns   总被引:5,自引:0,他引:5  
Landscapes were mapped as clusters of 2 or 3 land cover** types, based on their pattern within the clusters and tendency for a single type to dominate. These landscapes, called Landscape Pattern Types (LPTs), were combined with other earth surface feature data in a Geographic Information System (GIS) to test their utility as analysis units. Road segment density increased significantly as residential and urbanized land cover components increased from absent, to present as patch, to present as matrix (i.e., the dominant land cover type). Stream segment density was significantly lower in LPTs with an urbanized or residential matrix than in LPTs with either a forest or agriculture matrix, suggesting an inverse relationship between stream network density and the prevalence of human development other than agriculture in the landscape. The ratio of average forest patch size to total forest in the LPT unit decreased as agriculture replaced forest, then increased as residential and urban components dominated. Wetland fractal dimension increased as agriculture and residential land cover components of LPTs increased. Comparison of LPT and LUDA land cover area statistics in ecoregions suggested that land cover data alone does not provide information as to its spatial arrangement.  相似文献   

15.
Rainfall in drylands is erratic. Topographic features of landscapes can dampen or amplify temporal variability by spatially influencing patterns of water loss and accumulation. The extent to which portions of a landscape may differentially capture or retain scarce water and nutrient resources is an important determinant of vegetation patterns, particularly with respect to the distribution of woody plants. We therefore hypothesized that historic changes in woody cover on landscapes experiencing similar climate and disturbance regimes would vary with catena-to-catena (hillslope-to-hillslope) variation in topography-based hydrologic features. We tested this hypothesis by comparing topographic wetness index (TWI) values on replicate landscapes where woody plant abundance has increased over the past 100 yr. These landscapes are characterized by savanna parklands on coarse-textured upland portions of catenas that grade (1–3% slopes) into closed-canopy woodlands on fine-textured (lowland) portions of catenas. TWI values for woody and herbaceous communities were comparable within uplands, suggesting factors unrelated to surface/subsurface hydrology determine patterns of woody cover in these catena locations. TWI values for upland savanna parklands were significantly lower than those of closed-canopy woodlands occupying catena footslopes. Furthermore, uplands adjoining historically static woodland boundaries had lower TWI values than those where woodland boundaries had moved upslope 2.1 m yr−1 from 1976 to 1995. Results suggest runoff–runon relationships influence patterns of woody plant cover and change at the catena scale and may override constraints imposed by soil texture. As a result, changes in woody cover potentially accompanying changes in disturbance regimes, climate or atmospheric chemistry are likely to be constrained by topoedaphic settings. Models of vegetation dynamics may therefore need to explicitly account for rainfall–topography–soil texture relationships and associated scale-dependent mechanisms to accurately predict rates and patterns of change in woody and herbaceous plant abundance.  相似文献   

16.
Farina  Almo 《Landscape Ecology》1997,12(6):365-378
Richness, abundance and distribution of birds were investigated in the Aulella watershed,a mountainous area of 300 km2, located in the extreme northwestern corner of Tuscany, Italy in spring and summer, 1995. The study area encompasses five vegetation types (from Mediterranean maqui to upland beech forest) and three main land use categories (woodlands, mixed cultivated + urban areas, montane prairies). The recent history of land abandonment in the study area has produced a rapid expansion of shrubland and woodland, reducing cultivated areas to small patches interspersed in a woodland matrix. Richness, abundance and distribution of birds recorded at 414 points, randomly selected along secondary roads, and located using a Global Positioning System (GPS), were compared with topography, vegetation type and land use in a Geographic Information Systems (GIS) with a grid cell resolution of 200 × 200 m. Bird richness (55 species in all) and abundance are correlated: (a) negatively with the increasing altitude and increasing distance from cultivated areas; (b) positively with the increasing distance from woodlands and mountain prairies. Slope orientation appears to have a negligible effect on bird assemblages. Bird richness and abundance are significantly correlated with vegetation type. Cultivated areas support the highest bird richness and abundance that increase with patch size of the cultivated areas. Local extinction and/or reduction in within-species abundance of birds are expected to continue if the process of land abandonment continues. This revised version was published online in July 2006 with corrections to the Cover Date.  相似文献   

17.
Herbivores influence the structure of plant communities in arctic-alpine ecosystems. However, little is known of the effect of herbivores on plant colonisation following disturbance, and on its variability depending on the identity of herbivores and the characteristics of the habitats. To quantify the role of large and small vertebrate herbivores, we established exclosures of two different mesh sizes around disturbed subplots in forest and nearby tundra habitats in four contrasting locations in the forest-tundra ecotone in northernmost Sweden and Norway. The study revealed that herbivores influenced the abundance but not the species composition of regenerating vegetation. Gaps were colonised by the dominant species in the surrounding vegetation. The only exception to this expectation was Empetrum nigrum, which failed to colonise gaps even though it dominated undisturbed vegetation. Significant effects of herbivory were only detected when both small and large herbivores were excluded. Herbivores decreased the abundance of three of the most common species Vaccinium myrtillus, Vaccinium vitis idaea, and Deschampsia flexuosa. The effect of herbivory on the abundance of these three species did not differ between habitats and locations. However, the composition of the regenerating vegetation differed between habitats and locations. The disturbance treatment increased the species richness on the scale of plots, habitats, and sites. However, on the scale of whole locations, all species found in disturbed areas were also found in undisturbed areas, suggesting that the natural disturbance regime in arctic landscapes is high enough to sustain colonising species.  相似文献   

18.
为研究外来植物刺萼龙葵对入侵地植被多样性的影响,调查了河北省张家口市宣化区刺萼龙葵的3个不同生境(人工国槐林、河滩、荒坡)的植被,分析物种组成、植被多样性、植被重要值等指标。结果表明,无论哪种生境,刺萼龙葵的多度、频度、盖度与重要值均远远高于其他植物,刺萼龙葵在其入侵区内已经成为优势种;与对照样地相比,刺萼龙葵的生长引起30.00%以上的本土植物在入侵区消失,改变了植被组成和多样性;而原本优势的本土植物,相对多度与盖度也明显降低,其中狗尾草的降幅最大,相对多度和相对盖度分别降低了2.5倍和2.7倍。本研究为全面理解刺萼龙葵的入侵生态风险提供了依据。  相似文献   

19.
Large disturbances such as fires and floods are landscape processes that may alter the structure of landscapes in nature reserves. Landscape structure may in turn influence the viability of species and the functioning of ecosystems. Past reserve design and management strategies have been focussed on species and ecosystems rather than on landscape-scale processes, such as disturbance.An essential feature of a natural disturbance regime is the variation in disturbance attributes (e.g., size, timing, intensity, spatial location). Although some past reserve management policies have included natural disturbances, perpetuating disturbance variation has not been the explicit goal of either reserve design or management.To design a reserve to perpetuate the natural disturbance process requires consideration of: (1) the size of the reserve in relation to maximum expected disturbance size, (2) the location of the reserve in relation to favored disturbance initiation and export zones and in relation to spatial variation in the disturbance regime, and (3) the feasibility of disturbance control at reserve boundaries, or in reserve buffers.Disturbance management possibilities are constrained by the design of the reserve and the reserve goals. Where a natural disturbance regime is not feasible, then it is important that the managed disturbance regime mimic historical variation in disturbance sizes and other attributes as well as possible. Manipulating structure on the landscape scale to restore landscapes thought to have been altered by historical disturbance control is premature given our understanding of spatial disturbance processes in landscapes.  相似文献   

20.
张波 《中国园艺文摘》2013,29(1):128-131
广州万亩果园湿地是罕见的城中次生湿地和农耕湿地,在该区域建设湿地公园将有助于加强对湿地资源的保护和合理利用,提升周边环境质量,开发旅游资源.结合该地域的条件和发展历史,探索通过最少的干扰和合理的工程手段,将该地域转化成水网纵横交错、果基密布、百鸟翔集、野趣盎然,具有岭南园林特色和亚热带自然风光的未来城市中心湿地,同时为日后万亩果林保护建设、开发管理提供参考.  相似文献   

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