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1.
Hiroyuki Sugimoto Sakiko Kawabuchi Masatoshi Sugimori Joseph Gril 《Journal of Wood Science》2018,64(6):738-744
Transmittance and reflectance of visible light by sugi wood (Cryptomeria japonica) were investigated in the longitudinal (L) and tangential (T) directions. Transmittance was the highest in the L direction and reflectance was the highest in the T direction, suggesting that structural anisotropy influences transmittance and reflectance. Intra-ring variations observed with a microspectrometer indicated that T transmittance was higher for latewood than for earlywood, but there was no such trend in for L transmittance in which the highest levels occurred near the annual ring boundaries, on either the earlywood or latewood side, and the lowest at the transition from earlywood to latewood. Dependence of L transmittance on wavelength also showed variations according to the intra-ring position. The increasing of transmittance of earlywood at wavelengths?<?500 nm with increasing wavelength was observed, but this was not confirmed for latewood because of absorption by lignin. These observations supported a previously published finding, which was based on measurements in the radial direction, that the number of internal cell wall reflections, rather than density, determines wood lightness. Indeed, in the L direction, most of the incident light passes through lumens in earlywood and through cell walls in latewood, while it is subjected to numerous internal reflections at the interface between lumens and cell walls. This was further confirmed by the transmittance of earlywood being greatly decreased by radial compression. 相似文献
2.
Shrinkage at the tissue level has been evaluated satisfactorily using relatively simple equipment, comprising an optical microscope equipped with reflected light, a standard objective, a water immersion objective of same magnification and a digital camera connected to a computer. Shrinkage is calculated from pairs of images taken at the same magnification, one collected during immersion in water and the other in air-dry state. A novel software program has been developed to determine shrinkage based on a closed chain of reference points chosen from the anatomical markers at the external part of the zone of interest. Measurements were performed on earlywood, latewood and compression wood zones from two softwood species (Douglas fir and spruce), isolated from the rest of the annual ring with the aid of a diamond wire saw. As main results, reference should be made to the low degree of shrinkage and high anisotropy factor of earlywood, the marked and practically isotropic shrinkage in latewood and the low shrinkage (with respect to cell wall thickness) and inverse anisotropy ratio in compression wood. 相似文献
3.
对10个10年生日本落叶松无性系的木材基本密度、管胞参数进行了测定.结果表明:木材基本密度,早、晚材管胞宽度和早材长宽比无性系间差异显著;木材基本密度,早、晚材管胞长度,早晚材管胞宽度和早晚材长宽比径向变异模式相似,即从髓心向外以曲线形式不断增加,有时亦有起伏;早材从髓心向外以近似直线的形式缓慢增加,晚材从髓心向外以曲线形式增加,初期增加幅度较大,到一定年龄后趋于水平变化并略有波动;材质性状与树木年轮间的关系以对数方程、幂函数方程、指数方程拟合效果较好;除了晚材壁腔比和早材壁厚外,其它木材性质的重复力均在0.5以上,受中度或中度以上的遗传制约,按照20%的选择率,长宽比和晚材管胞长能获得较高的遗传增益. 相似文献
4.
The amount of apparently nonfreezing water per dry mass unit significantly differs between earlywood and latewood, and drying changes the nonfreezing water content of earlywood cell walls in a time-dependent manner. However, the equilibrium moisture content of spruce wood is not affected by drying and rewetting. The results indicate that different mechanisms govern these two types of phase transformations of cell wall water. The nonfreezing water content, as determined using differential scanning calorimetry, appears to be a nonequilibrium property. It is hypothesized that the measured changes in nonfreezing water content mostly reflect changes in the porous cell wall structure, on a scale well above the molecular scale, rather than the abundance of chemical adsorption sites. 相似文献
5.
Hydraulic and mechanical properties of young Norway spruce clones related to growth and wood structure 总被引：3，自引：0，他引：3
Stem segments of eight five-year-old Norway spruce (Picea abies (L.) Karst.) clones differing in growth characteristics were tested for maximum specific hydraulic conductivity (k(s100)), vulnerability to cavitation and behavior under mechanical stress. The vulnerability of the clones to cavitation was assessed by measuring the applied air pressure required to cause 12 and 50% loss of conductivity (Psi(12), Psi(50)) and the percent loss of conductivity at 4 MPa applied air pressure (PLC(4MPa)). The bending strength and stiffness and the axial compression strength and stiffness of the same stem segments were measured to characterize wood mechanical properties. Growth ring width, wood density, latewood percentage, lumen diameter, cell wall thickness, tracheid length and pit dimensions of earlywood cells, spiral grain and microfibril angles were examined to identify structure-function relationships. High k(s100) was strongly and positively related to spiral grain angle, which corresponded positively to tracheid length and pit dimensions. Spiral grain may reduce flow resistance of the bordered pits of the first earlywood tracheids, which are characterized by rounded tips and an equal distribution of pits along the entire length. Wood density was unrelated to hydraulic vulnerability parameters. Traits associated with higher hydraulic vulnerability were long tracheids, high latewood percentage and thick earlywood cell walls. The positive relationship between earlywood cell wall thickness and vulnerability to cavitation suggest that air seeding through the margo of bordered pits may occur in earlywood. There was a positive phenotypic and genotypic relationship between k(s100) and PLC(4MPa), and both parameters were positively related to tree growth rate. Variability in mechanical properties depended mostly on wood density, but also on the amount of compression wood. Accordingly, hydraulic conductivity and mechanical strength or stiffness showed no tradeoff. 相似文献
6.
We investigated morphological changes in wood tissues of sugi (Cryptomeria japonica) resulting from treatment with the ionic liquid 1-ethyl-3-methylimidazolium chloride ([C2mim][Cl]), which dissolves cellulose. Treatment with [C2mim][Cl] caused dissociation and distortion of tracheids in latewood, but not in earlywood. This difference was due to the difference in swelling behavior of the cell wall between earlywood and latewood. Many pit membranes in bordered pits were broken by treatment with [C2mim][Cl]. In addition, some chemical changes in wood components, such as cellulose and lignin, occurred before significant disruption or destruction of the cell wall. Our results show that the reaction of wood liquefaction by [C2mim][Cl] treatment is not homogeneous, both from chemical and morphological viewpoints. 相似文献
7.
人工兴安落叶松次生木质部的解剖学研究 总被引：1，自引：0，他引：1
运用木材解剖图像分析系统和显微照相的方法对人工兴安落叶松次生木质部的解剖结构进行研究,结果表明:落叶松具正常树脂道和受伤树脂道两种类型,前者常见于晚材。落叶松生长轮内的早晚材在干和枝内急变,在根内缓变。早材管胞呈六边形至多边形,胞壁常见单列具缘纹孔,偶见对列具缘纹孔;晚材管胞多呈矩形,胞壁鲜见具缘纹孔,通常为单列具缘纹孔。落叶松木射线同时具有单列木射线和纺锤形木射线两种类型,纺锤形木射线中仅含一枚纵行树脂道。纵行管胞与木射线交叉形成的纹孔场为云杉型。从根到干再到枝,管胞逐渐细化,管胞长度逐渐减小,木射线分布由密到疏。 相似文献
8.
Variation of measured cross-sectional cell dimensions and calculated water vapor permeability across a single growth ring of spruce wood 总被引：1，自引：1，他引：0
A statistical study of the cell dimensions in a growth ring of spruce along the radial and tangential directions is performed. The data are used to study the variation of the cell vapor permeability in the growth ring. Studying cell rows within one growth ring, the frequency distributions of the cell wall thickness in the radial direction and of the lumen dimension in the tangential direction are found to be both unimodal. In contrast, the frequency distributions of these dimensions in the other directions are bimodal, where the different modes can be attributed to earlywood and latewood. Analysis of the bimodal distributions results in the determination of threshold values of cell wall thickness and the lumen dimension for earlywood and latewood tracheids. The cell dimensions are used to predict cell porosity and water vapor permeability distribution within a growth ring. The bimodal frequency distributions of the tangential cell wall thickness and the radial lumen dimension provide an explanation for the observed bimodal frequency distribution of the cell water vapor permeability both in radial and in tangential directions. Contrary to measured macroscopic vapor permeability results, the tracheid geometry results in lower cell vapor permeability in radial than in tangential direction. This confirms that rays play an important role in the vapor permeability of wood, as they can be considered as pathways for vapor transport in radial direction. The dataset analyzed in this paper leads to a set of parameters characterizing the earlywood and latewood cell dimensions. Such characterization can be used, for example, for producing synthetic data for computational modeling studies. 相似文献
9.
In this study, the contact angles of four different reference liquids (including distilled water, diiodomethane, formadide
and glycerol) formed on the surfaces of wood, treated with chromated copper arsenate (CCA) and two other emerging copper-based
water-borne systems (commercial names: NW and NS) were measured with sessile drop method. Based on the contact angle data,
the surface energy was obtained from the acid-base approach. The total surface energy consisted of Lifshiz-van der Waals parameter
and acid-base parameter. Results showed that the NW and CCA treatments made the wood surface more hydrophobic while the NS
treatment had the reverse effect on the wood surface mainly owing to the increased penetration of earlywood. By using three
liquids, diiodomethane, formamide and distilled water, the total surface energy obtained for untreated earlywood, untreated
latewood, CCA-treated earlywood, CCA-treated latewood, NW-treated earlywood, NW-treated latewood, NS-treated earlywood and
NS-treated latewood were 43.1, 44.5, 43.4, 45.1, 49.4, 40.6, 46.0 and 40.9 mJ/m2, respectively. The surface energy of CCA-treated wood was almost the same as untreated wood. After NW and NS treatments,
the surface energy of both earlywood and latewood changed a little. However, the change was not so obvious as to draw any
further conclusion concerning the influence of NW and NS treatments on the surface energy of wood.
__________
Translated from Journal of Beijing Forestry University, 2006, 28(4): 1–4 [译自: 北京林业大学学报] 相似文献
10.
Kilpeläinen A Gerendiain AZ Luostarinen K Peltola H Kellomäki S 《Tree physiology》2007,27(9):1329-1338
We studied the effects of elevated temperature and carbon dioxide concentration ([CO(2)]) alone and together on wood anatomy of 20-year-old Scots pine (Pinus sylvestris L.) trees. The study was conducted in 16 closed chambers, providing a factorial combination of two temperature regimes and two CO(2) concentrations (ambient and elevated), with four trees in each treatment. The climate scenario included a doubling of [CO(2)] and a corresponding increase of 2-6 degrees C in temperature at the site depending on the season. Anatomical characteristics analyzed were annual earlywood, latewood and ring widths, intra-ring wood densities (earlywood, latewood and mean wood density), tracheid width, length, wall thickness, lumen diameter, wall thickness:lumen diameter ratio and mass per unit length (coarseness), and numbers of rays, resin canals and tracheids per xylem cross-sectional area. Elevated [CO(2)] increased ring width in four of six treatment years; earlywood width increased in the first two years and latewood width in the third year. Tracheid walls in both the earlywood and latewood tended to become thicker over the 6-year treatment period when temperature or [CO(2)] was elevated alone, whereas in the combined treatment they tended to become thinner relative to the tracheids of trees grown under ambient conditions. Latewood tracheid lumen diameters were larger in all the treatments relative to ambient conditions over the 6-year period, whereas lumen diameters in earlywood increased only in response to elevated [CO(2)] and were 3-6% smaller in the treatments with elevated temperature than in ambient conditions. Tracheid width, length and coarseness were greater in trees grown in elevated than in ambient temperature. The number of resin canals per mm(2) decreased in the elevated [CO(2)] treatment and increased in the elevated temperature treatments relative to ambient conditions. The treatments decreased the number of rays and tracheids per mm(2) of cross-sectional area, the greatest decrease occurring in the elevated [CO(2)] treatment. It seemed that xylem anatomy was affected more by elevated temperature than by elevated [CO(2)] and that the effects of temperature were confined to the earlywood. 相似文献
11.
Cornelia Krause Sergio Rossi Maxime Thibeault-Martel Pierre -Y. Plourde 《Annals of Forest Science》2010,67(4):402-402
12.
Fracture in Norway spruce wood treated with <Emphasis Type="Italic">Physisporinus vitreus</Emphasis>
Changes in the fracture behaviour of Norway spruce tonewood after fungal treatment were studied. Specimens were incubated for 6, 9 and 12 months with Physisporinus (P.) vitreus. Fracture tests were performed in a compact-tension fracture experiment set-up, and the results were compared with the morphological analysis of the degraded wood structure and transverse sections of the crack tip viewed under light and fluorescence microscopy. It was evident that both the failure load and critical stress intensity factors were reduced in wood after prolonged incubation periods. Weight losses were significantly higher in sapwood than in heartwood. With prolonged incubation periods, the frequency of unstable fracture and brittle behaviour of the wood increased. In untreated wood, cracks were initiated in the earlywood. The process involved both delamination of the cells within the middle lamellae and rupture of the cell walls, inducing a zigzag crack tip pattern. In fungally treated wood, cracks often commenced from the intersection between late- and earlywood, resulting in a straight tangential crack line. Micrographic images showed that P. vitreus was more active in the secondary walls of latewood tracheids. In this region of the wood, the cell walls were strongly degraded after 9–12 months of incubation, resulting in a reduction in tensile strength, even though the wood did not show strong features of decay at the macroscopic level. 相似文献
13.
E. A. Tyutkova S. R. Loskutov N. P. Shestakov 《Wood material science & engineering》2020,15(4):205-212
ABSTRACT This paper presents the results of FTIR spectroscopy of the 1988–1998 annual ring early and latewood of Larix gmelinii (Rupr.) Rupr.) found along the polar treeline (Taimyr Peninsula, 70°52′53″ N, 102°58′26″ E). We analyzed samples of early and latewood to identify absorption bands of groups of wood components, as well as the bands that characterized the interactions among the components. We studied bound water spectral characteristics for wood formed in different years. An analysis of the correlation between the values of spectral absorption bands and mean monthly air temperature and precipitation showed that May-August weather had the greatest influence on the synthesis of the main polymeric components of early and latewood. Air temperature had a considerable effect on polymer composition of cell walls forming in early and latewood, whereas precipitation influenced only earlywood. FTIR spectroscopy is a promising tool to develop information on the biochemical composition of the walls of early and late tracheids of annual rings and on weather and climate influences on cell wall synthesis. 相似文献
14.
Longitudinal hardness and Young's modulus of spruce tracheid secondary walls using nanoindentation technique 总被引：6，自引：2，他引：6
Dr. R. Wimmer Dr. B. N. Lucas W. C. Oliver Dr. T. Y. Tsui 《Wood Science and Technology》1997,31(2):131-141
Summary Using a mechanical properties microprobe, measurements of hardness and elastic modulus of tracheid walls in the longitudinal direction of spruce wood were obtained by continuously measuring force and displacement as a diamond indenter impressed a cell wall. Maximum mechanical properties were found at the edges of the walls of angular shaped tracheids. Both the hardness and elastic modulus of latewood cell walls were higher than cell walls in the earlywood. The high spatial resolution of this new concept of mechanical testing allows a direct comparison with ultrastructural and microchemical parameters of lignified cells which opens a wider area of applications for the understanding of intrinsic wood properties.This work was conducted while the senior author was a Visiting Scientist at the Oak Ridge National Lab, Oak Ridge, TN 37831, USA partly with joint fundings from the Austrian Science Foundation (Schrödinger scholarship J799-BIO) 相似文献
15.
Genetic parameters for various wood density traits were estimated in 29-year-old trees of 18 full-sib families of hybrid larch
(Larix gmelinii var. japonica × Larix kaempferi) F1. Intra-ring density variation (IDV) was also evaluated using a model that expresses the pattern curve from earlywood to latewood
as a power function. A high IDV indicates an abrupt change in wood density from earlywood to latewood. The ring width and
wood density traits of individual rings were determined by X-ray densitometry. Overall wood density (RD) was shown to increase
with increasing ring number, ranging from 0.42–0.59 g/cm3, whereas IDV of individual rings decreased gradually from pith outwards. Estimates of individual tree narrow-sense heritability
of RD and IDV were 0.66 and 0.67, respectively. IDV showed negative genetic and phenotypic correlations with RD (r
g = −0.99, r
p = −0.72). The predicted genetic gains in latewood proportion and IDV were higher than that of RD. These results suggest that
the intra-ring density variation is under moderate genetic control equivalent to wood density. The trend of increasing wood
density from earlywood to latewood was associated with changes in both tracheid diameter and cell wall thickness. 相似文献
16.
Trairat Neimsuwan Siqun Wang Adam M. Taylor Timothy G. Rials 《Wood Science and Technology》2008,42(6):493-506
A better understanding of the sorption behavior of different wood structures could be useful in protecting wood against wood
deterioration and fungal attack. The purpose of this study was to investigate the effect of differences among earlywood, latewood,
and tree ring location within the stem cross-section of loblolly pine (Pinus taeda) on the sorption kinetics and statics of water vapor under ambient conditions. The water vapor sorption of earlywood and
latewood in different tree rings was recorded using a dynamic contact angle analyzer under relative humidity changes from
11 to 89%, as provided by saturated salt solutions. Earlywood had higher sorption rates and diffusion coefficients than latewood,
while outer tree rings had higher sorption rates and diffusion coefficients than inner tree rings. The sorption isotherms
of earlywood, latewood, and different tree ring locations within the stem cross-section were fitted very well by a Hailwood–Horrobin
model. 相似文献
17.
Steam explosion pretreatment results in the formation of microcracks in the cell walls of wood. In the present study, steam explosion experiments were performed and structural changes in Norway spruce were identified using scanning electron microscopy. The cellular structure of the softwood spruce was simulated using the finite element method, and the effects of pressure generated during the steam explosion pretreatment on the deformation of the cells were investigated. The simulated model included earlywood, latewood, and ray cells. The effects of bordered and cross-field pits on the stresses in the cell wall were studied as well. Many similarities were observed between the microcracks in the steam-exploded wood and the high-stress regions predicted by the model. The experimental and simulation results showed that the radial cell walls in the earlywood cells experienced major deformation. The presence of the pits created stress localization and facilitated the formation of microcracks in the cell walls. 相似文献
18.
The fracture behaviour of single wood fibres is governed by geometrical constraints: in situ ESEM studies on three fibre types 总被引：1，自引：1，他引：0
In situ tensile tests were performed in an environmental scanning electron microscope (ESEM) on earlywood, transition wood
and latewood cells of Norway spruce (Picea abies [L.] Karst.). In order to examine the single wood fibres in a wet state, a specially designed tensile testing stage with
a cooling device was built. The fracture behaviour of the cell types was studied at high resolution while straining. Different
failure mechanisms were observed for the three tissue types. The thin-walled earlywood fibres showed tension buckling which
gave rise to crack initiation and resulted in low tensile strength, whereas thick-walled latewood fibres predominately failed
by transverse crack propagation without fibre folding. 相似文献
19.
Summary The variation of the lignin content within one growth increment of Abies sachalinensis was investigated with the aid of an ultraviolet microscopic image analyser. The lignin content was determined continuously in each cell within a growth increment. The direct photometric scanning method of UV image is believed to give accurate results for determination of the lignin content of the cell wall, since it contains fewer assumptions. The lignin content of the earlywood was higher than that of latewood in adult wood as same as the others. It was high, however, in the terminal zone of the latewood. The trends of the juvenile wood were quite different from those of the adult wood. The lignin content increased from earlywood to latewood.This paper was presented at the Wood Anatomy Congress of IAWA, Aug. 27, 1979, Amsterdam. The authors wish to express their gratitude to Prof. Dr. J. Bauch for helpful advice 相似文献
20.
In order to study the transverse swelling/shrinkage of wood, the microscopic swelling behavior of latewood tracheid was observed
using confocal laser scanning microscopy and the digital image correlation method. A microcrater structure was created on
the surface of the specimen by using the ion sputter etching technique to obtain a pattern-rich digital image for image analysis.
Douglas fir specimens were conditioned by two methods of absorption: rapid absorption of moisture from hot steam, and slow
absorption of moisture from the water vapor of saturated solutions. Latewood tracheid near the surface of the specimen deformed
only in the radial direction when the relative humidity of the surrounding air changed rapidly (rapid absorption of moisture
from hot steam or absorption/desorption of moisture during the observation). In addition, the diameter of the lumen decreased
upon rapid absorption of moisture, whereas it expanded upon slow absorption of moisture. These results indicate that the microscopic
swelling behavior of latewood cells is strongly influenced by the macroscopic/mesoscopic structure, for instance, the cell
arrangement or the alternation of latewood and earlywood.
Part of this report was presented at the 54th Annual Meeting of the Japan Wood Research Society, Sapporo, August 2004 相似文献