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1.
Competition between established Dactylis glomerata or Festuca pratensis / Taraxacum officinale swards and seedlings of Trifolium pratense was studied under field conditions in Norway. Root competition from the Festuca pratensis / Taraxacum officinale sward was significantly greater than that from a Dactylis glomerata sward. The balance in root and shoot competition varied between experimental years because of variation in environmental conditions. In the first experimental year, root competition had a greater effect on seedling dry weight than shoot competition, whereas the opposite was found in the second year. However, both root and shoot competition (full competition) in comparison with no competition gave proportionally the same effects in both years, indicating that an alleviation in root competition will be followed by intensified shoot competition. An increasing effect of shoot competition with time on plant dry weight was found. In a second experiment, seedlings of Lolium perenne and Trifolium pratense produced the highest plant dry weights in competition with an established Phleum pratense sward, followed by seedlings of Poa pratensis , Phleum pratense and Trifolium repens in decreasing order. However, no interactions between seedling species and competition treatments were found, indicating no differences in sensitivity between seedling species to competition from the established grass sward. The results are discussed in the context of renovating grassland by direct drilling.  相似文献   

2.
Seeds of white clover ( Trifolium repens ) were sown alone or in fluid, or pre-germinated in fluid on to the soil surface of slots cut in an old Agrostis/Festuca sward. Seeds were left uncovered or covered with a thin layer of soil and irrigated from the outset or after 14 days delay.
With irrigation from the outset, seedlings established soonest from pre-germinated seeds but final percentage establishment ranged only from 50 to 63% for the three states of seed. With delayed Irrigation, however, few pre-germinated s1 survived and although germination of dry seed was delayed, final percentage establishment was unaffected. Soil covering increased establishment slightly. The number of leaves per seedling was greatest with pre-germination and early irrigation and least with pre-germination and delayed irrigation. Delayed irrigation also decreased the number of leaves per seedling for seedlings grown from dry seed.
The effects of early irrigation persisted so that at 84 days it significantly increased total yield of clover from all three states of seed, on average threefold, but much more from pre-germinated than from dry seed. Soil covering greatly increased yield, especially where irrigation had been applied from the outset. The interaction of early irrigation and soil covering was even more pronounced for stolon weights, and stolon growth of Individual seedlings appeared to benefit more from soil covering than from early irrigation.
The results showed the importance of early seedling emergence, but no advantages of fluid sowing per se. They also indicated large overall beneficial effects of early irrigation and its additive interaction with soil covering.  相似文献   

3.
The effects of delayed emergence of white clover (cv. Grasslands Huia) seedlings, following slot-seeding during spring and early summer 1983, were simulated by sowing seeds at intervals into slots cut in turves of permanent grassland in soil-filled tanks or in the field. The resulting spread and growth of the species were assessed during the following 15–18 months. The effects of propyzamide, a grass-suppressing herbicide (at 0·2 kg ha-1), were also investigated in the field.
A delay of 10 or 18 days in the sowing of seeds in turves in the tanks reduced clover dry weight yield from 23 to 11 during 1983 and from 118 to 96 g (0·12 m)-2 during 1984. Spread of stolons from the slots was also greatly delayed and effects persisted for at least a year. Increased clover growth on the earliest sown treatment more than compensated for a slightly smaller yield of grass.
In the field, yields of clover from late summer 1983 until spring 1984 closely reflected the order of emergence. Afterwards, differences were less clear-cut but the cumulative yield of clover until August 1984 of the earliest-sown treatment was twice that of a treatment sown 20 d later. Effects of delayed sowing in summer 1983 were more pronounced on stolons than on foliage growth when measured in September 1984, with significant reductions in many stolon attributes even with 8 d delay in sowing. Application of propyzamide in autumn 1983 increased clover yield during 1984 from 1·7 to 3·0 and total herbage yield from 7·1 to 8·6 t ha-1.
The results demonstrated some of the benefits of rapid seedling emergence and of the use of a grass-suppressing herbicide, but did not indicate any interaction between the two factors.  相似文献   

4.
Establishing Kura clover (Trifolium ambiguum) in mixtures with grass species is challenging, because slow growth of clover seedlings results in low competitive ability. This study examined establishment success by altering time of seeding of the grass component to reduce competition with Kura clover seedlings. Two trials, one of Kura clover–meadow bromegrass (Bromus biebersteinii) and the other Kura clover–orchardgrass (Dactylis glomerata) mixtures were planted in Edmonton, Alberta. Grasses were seeded at the same time as the clover, or introduced when the clover reached one true leaf or three true leaves, in the autumn of the planting year or the following spring. Species composition varied significantly between treatments. When sown at the same time, Kura clover contributed 31 and 14% of yield in the establishment year when sown with meadow bromegrass and orchard grass, respectively. Delaying grass sowing until Kura clover had one or three leaves gave a higher percentage of Kura clover compared with planting at the same time. Autumn and spring grass sowing resulted in stands of 78 and 80% clover with meadow bromegrass, and 74 and 67% clover with orchardgrass. Altering the competitive advantage of the grass species to produce a more balanced mixture was successfully achieved by delaying seeding of the grass relative to Kura clover. A long interval before introducing the grass (autumn or following spring), was not successful as established Kura clover seedlings have an increased competitive ability.  相似文献   

5.
The productivity of a mixed sward, comprising perennial ryegrass cvs Barlano and Bastion and white clover cvs Donna and Aran, was measured under sixteen fertilizer N treatments. These involved 0.25, 50 and 75 kg N ha-1 in spring only, in autumn only and in all combinations of spring N and autumn N. A simulated grazing regime of six cuts annually at 3- to 6-week intervals was imposed. Increasing rates of total N application increased total herbage DM regardless of application pattern. Yield response was greater with N applied in the spring, and total herbage DM was higher with high spring N-low autumn N than the reverse. Mean yield responses at the first harvest to 25, 50 and 75 kg ha-1 N in spring were 13.6, 10.8 and 11.6 kg DM per kg N. Corresponding responses at the final harvest to N rates in the autumn were 7.2, 5.8 and 6.8 kg DM per kg N. Responses were similar at these times for treatments receiving combined spring and autumn N. Over all treatments, mean annual production of total herbage was between 7.08 t ha-1 DM with no N and 8.19 t ha-1 with 75 kg ha-1 N in both spring and autumn. Owing to drought, mean production in year 2 fell by 32% compared with year 1. White clover production fell progressively with increasing N application. Treatments with spring-applied N gave the most marked decrease. White clover was more markedly depressed than the associated grass by the drought in the second year. The mean reductions in white clover content were 0.17, 0.07 and 0.12 percentage units per kg applied N for spring N, autumn N and combinations. Autumn N use depressed white clover less than spring N but the yield response of grass was less. It is concluded that any applied N adversely affects white clover performance to some degree. Where management factors are unfavourable to white clover even strategic N use may not be wise. Instead, it is suggested that a ‘dual-sward’ approach be adopted in practice, namely, grass/white clover swards with no N. and complementary grass swards receiving optimum applied N to give better production at times when grass/white clover swards are relatively less productive.  相似文献   

6.
Growth of grass/clover mixtures during winter   总被引:2,自引:0,他引:2  
The percentage of clover in a mixture of Huia white clover and Melle perennial ryegrass decreased during each of three winters. In the most severe winter, both grass and clover suffered a net loss of weight, clover losing two–thirds of its maximum weight while grass lost about one–third. In the mildest winter, grass continued to gain weight throughout, while clover lost weight. Such losses, when combined with conditions such as the use of nitrogen fertilizer, which prevent a recovery of clover percentage during summer, will lead to a long–term decrease in the clover content of a sward.
The losses from the clover were contributed to by all plant parts especially leaf, which lost between 60% and 95% of lamina weight. The change from autumn to winter caused a greater decrease in the number of live leaves per shoot. and in the area and weight of individual leaves, in clover than in grass. Other work has shown that there is a greater decrease in photosynthesis in clover than in grass, probably owing to changes in canopy structure; this is one cause of clover's poor performance, though differences between the two species in the effect of winter on the distribution of the carbon fixed in photosynthesis and in losses of weight in respiration and tissue death may also be involved.  相似文献   

7.
In three successive years, sward height was maintained at 3, 5, 7 or 9 cm on grass swards receiving a total of 300 kg N ha?1 in six equal monthly dressings from April, and on grass/clover swards receiving 50 kg N ha?1 as a single dressing in early spring. From turnout in April until weaning in July, 64 ewes and their lambs (mean litter size 1·5) were continuously grazed at the four sward surface heights on the two sward types. White clover content of grass/clover swards remained low throughout the experiment ranging from 0·2 to 7·4% of the herbage mass. During the first two years, lamb gains averaged over sward types were 204, 260, 285 and 308 g d?1 up to weaning, while in the third year gains were 238, 296, 296 and 260 g d?1 on 3, 5, 7 and 9 cm swards respectively. Ewes lost live weight on 3 cm swards but apart from this sward height had little effect on performance. During the autumn, weaned lambs gained — 27, 87, 147 and 167 g d?1 on 3, 5, 7 and 9 cm swards respectively. Sward type had only a small effect on the performance of lambs up to weaning but in the autumn, mean gains of weaned lambs were lower on grass/N swards (73 g d?1) than on grass/clover swards (115 g d?1). Relative to 3 cm swards, carrying capacities of 5, 7 and 9 cm swards were 0·76, 0·57 and 0·52 respectively from turnout to weaning and 0·66, 0·52 and 0·44 respectively during autumn. Grass/clover swards carried 0·67 of the ewes carried by grass/N swards from turnout to weaning and 0·51 of the live weight carried by grass/N swards during autumn. The reaction of the two sward types to sward height did not appear to differ but in the third year there was evidence of a reduction in white clover content when swards were grazed at 9 cm. The data suggest that lamb growth rates will increase as sward height increases up to 9 cm and the evidence for this was stronger with weaned lambs in autumn than with suckling lambs in spring.  相似文献   

8.
The annual yield of tall fescue was higher than that of Italian ryegrass in the third year after sowing, but the total yield of herbage from grass plus clover swards was similar.
In both the second and third year after sowing, the yield of herbage in the spring grazing was higher when fescue was used as the sown grass. The method of establishment of both tall fescue and Italian ryegrass affected the total and seasonal yield in the second and third year after sowing, but the magnitude of these effects was not nearly as marked as it was earlier in the life of the leys. In the second year after sowing, swards of both Italian ryegrass and tall fescue had a higher yield of total herbage and of white clover, and a lower ingress of unsown species, when established without a cover crop and grazed frequently in the year of sowing.
The inclusion of red clover did not increase total yield of DM in the second and third year after sowing, and it slightly decreased the yield of the tall fescue mixture in the third year following sowing when N was applied. S170 tall fescue was readily grazed by sheep in spring and autumn.
The apparent recovery of applied N varied with the mixture sown, and the management given during establishment.  相似文献   

9.
Small plots of a Festuca-Agrostis upland sward on a peaty gley podsol were strip-seeded during late June 1986 with white clover cvs Aberystwyth S184 or Menna at 4 kg ha−1 and defoliated early (20 August) or late (3 September) and then frequently or infrequently (every 2 weeks or 4 weeks) until the end of September. All plots were defoliated in early November, at 3-weekly intervals during the growing season in 1987 and then grazed rotationally during 1988.
Satisfactory seedling establishment, representing 46% emergence, was achieved 5 weeks after sowing. The differential defoliation regimes had no persistent significant effects on clover development. S184 soon produced more leaves per seedling than Menna and a smaller proportion of its leaf number and weight were removed at each defoliation. Following large losses of leaves over the 1986–87 winter, SI84 had significantly more leaves per stolon than Menna; subsequently it also colonized the sward at a quicker rate. During 1987 amounts of herbage harvested (6.1 t ha−1) were similar with the two clover cultivars, with S184 contributing 47% and Menna 44% of this respectively. SI84 made a larger contribution to yield during May and June but Menna was more productive during September and October. During 1988 clover populations were maintained with rotational grazing without additional fertilizer inputs.
The results show that, despite initial soil and climatic contraints, both small and medium-leaved clovers can be strip-seeded into upland swards with large subsequent benefits to yield and herbage quality. However, they also indicate the need for further experiments to determine the influence of sward morphology and defoliation regime on stolon branching rates and accumulation of growing points which, in turn, govern sward colonization.  相似文献   

10.
The aim was to study the effects of white clover cultivar and combinations with perennial ryegrass cultivars on seedling establishment in autumn‐sown swards and on winter survival of seedlings. Large‐leaved white clover cv. Alice and small‐leaved white clover cv. Gwenda, and an erect and a prostrate perennial ryegrass cultivar were sown in autumn in pure stands and as four binary grass‐clover mixtures. Mixtures of white clover cv. Huia and Aberherald with perennial ryegrass were also sown. Companion grasses had no significant impact on the establishment of white clover. The number of seedlings of white clover cv. Alice in mixtures (335 m?2) was higher than cv. Gwenda (183 m?2) and pure swards had similar white clover population densities as mixed swards. White clover cv. Huia tended to have more seedlings than Aberherald (355 and 205 m?2 respectively). No stolons were produced prior to a severe winter, because of the late sowing date. Winter survival of clover seedlings was 0·56 in mixtures and 0·69 in pure stands, irrespective of white clover or companion grass cultivar. Stolon development of white clover in autumn is often considered essential for overwintering survival and spring growth. In this study, there was considerable survival of the non‐stoloniferous tap‐rooted seedlings of all four clover cultivars despite a severe winter.  相似文献   

11.
The effect of 4 or 5 forms of nitrogenous fertilizer on the yield and botanical composition of a perennial ryegrass/white clover sward was examined in 3 different years. The uptake of N by the grass was also estimated. Ammonium nitrate (as Nitro-Chalk), ammonium sulphate and urea gave similar yield increases, although in one year, when high rainfall followed the application of fertilizer, urea was slightly inferior. Gas liquor gave variable responses in the 3 years. Nitroform, a urea-formaldehyde, was generally less effective than the other forms of fertilizer. The uptake of N from Nitro-Chalk, sulphate of ammonia and urea did not vary greatly from year to year although the uptake of soil N by the untreated control did.
Response to N in the yield of dry-matter varied from year to year. This variation was due partly to differences in the amount of clover present and partly to differences in the extent to which N taken up by the grass was used for the synthesis of dry-matter.
The white clover content of the sward was not greatly reduced in spring by the application of the fertilizer, and there were no consistent differences between the effects on clover of the forms of the fertilizer. A reduction was apparent in the summer but not in the autumn.
It may be necessary to increase replication above the level of 3 to 6 which is frequently employed in field experiments if more precise quantitative estimates of response and of the relative efficiency of various forms of nitrogenous fertilizer are required.  相似文献   

12.
The effect of applying 100kg P ha-1 per year in the form of triple superphosphate to mixed swards of perennial ryegrass ( Lolium perenne ) and white clover ( Trifolium repens ) was examined on a soil of low P status.
The dry matter yield of total herbage was increased by 10'/i by added P. In the year of sowing the ryegrass benefited more than the clover from added P: in the subsequent four years the two species benefited equally. Both species responded 10 added P to a similar extent in terms of leaf size; the clover responded less well than the ryegrass in terms of rate of leaf emergence. However, clover responded positively to added P in terms of stolon internode length, length of stolon per m2 and number of growing points per m2. It is suggested that the application of P may promote the spread of white clover within an open sward, but that its application may not enhance the competitive power of white clover when growing with vigorous grasses.  相似文献   

13.
Effects of sowing date, fungicide seed treatments, and the head smut fungus Ustilago bullata Berk, on establishment, herbage, and seed production of prairie grass cv. Grasslands Matua ( Bromus willdenowii Kunth ) were measured in autumn- and spring-sown field trials in New Zealand.
Autumn seedling establishment from thiram-treated seed relative to untreated seed was increased by 27% from the earliest sowing (early March), and by 65% from the latest sowing 8 weeks later. Infestation of seed with ustilospores of U. bullata reduced seedling establishment at all but the latest autumn sowing. Fungicide seed treatments had little effect on spring seedling establishment, but V. bullata infestation of seed reduced establishment compared with healthy seed, particularly after late sowings (early November).
Herbage production was greater from early than from late sowings in both autumn and spring, and was less in plants grown from U. bullata infested seed than in plants from healthy seed. Enhanced autumn establishment from thiram-treated seed led to improved herbage production when plants were young, but this effect disappeared about 18 weeks after late autumn sowing. Overall herbage production from spring sowing was more than twice that from autumn sowing.
A 6-week delay in autumn sowing led to a 58% reduction in seed production the following summer.
Early autumn and spring sowings, together with adequate control of seedling diseases, are likely to give optimum herbage and seed production from prairie grass.  相似文献   

14.
Populations of white- and red-clover were examined after they had been grown for several years in grass/clover swards. The method of sowing, nitrogen application, management and season affected the ratio of very similar clover cultivars differing specifically in the leaf mark when sown in the same mixture. The companion grass also infiuenced the proportion of legume cultivars in the sward. In addition, the mean date of emergence of a single cultivar of broad-red clover was affected by management.  相似文献   

15.
Experiments are described in which a study was made of the top growth, root weight, nodule number, and type of nodules of white clover growing in a grass sward in the field. Soil cores were taken and the clover root-system examined after removal of soil by washing.
The most important factor governing the total number of nodules and the number of large (2 mm. or more in length) nodules under the sward was the amount of clover root-material present.
Nitrogenous fertilisers and the return of dung and urine by grazing sheep reduced the amount of clover root-weight and consequently the number of nodules found under the pasture. 36 1b. of nitrogen applied in the early spring of 1956 had a beneficial effect on both clover top-growth and nodulation. 80–100 1b. of nitrogen per acre had no effect on the number of nodules per gram of root. However, the application of 200 1b. of nitrogen per acre caused a significant decrease in the number of large nodules per gram of root.
The numbers of large nodules present reached a midsummer peak where nitrogen was not applied.
During the autumn many nodules showed part green and part pink colouration or were completely green or brown, an indication of a decline in nitrogen-fixing ability. Decaying nodules increased considerably in number when the legume was being suppressed by nitrogen application and top growth was poor. The practical application of these results is discussed and methods of maximising the benefit from clover and fertiliser nitrogen are suggested.  相似文献   

16.
This study assesses near-infrared reflectance spectroscopy (NIRS) as a rapid and reliable method for estimating the clover content of clover/grass mixtures. NIRS calibrations were derived both for red clover ( Trifolium pratense ) and white clover ( T. repens ) mixtures. To maximize variability, the set of samples for analysis was selected from field plots receiving a wide range of nitrogen fertilizer application and harvested over 2 years at weekly intervals throughout the growing season. The samples were scanned using a NIR-Systems model 5000 monochromator. A total of 183 white clover/grass and 282 red clover/grass samples were used to calibrate and cross-validate the equations derived. The NIRS calibrations obtained from this study have the potential to be used in research on clover/grass mixtures as well as for advisory work.  相似文献   

17.
Under mowing conditions in the absence of the animal, ultra-simple swards based either on ryegrass or cocksfoot gave large increases in yield of dry matter and nitrogen from the inclusion of white clover in the seeds mixture. Raising the clover seed rate from 1 to 3 lb./acre gave a further response in yield.
As the swards aged, the continuing effect of increased clover seed rate was dependent on applied nitrogen.
The rate of transference of nitrogen from clover to the soil depended upon the species of the associated grass, the clover seed rate, the age of the sward and the degree of reduction of the clover in the sward.
Suppression of clover led to an increase in soil nitrogen, particularly with high seed rates of clover.
Dry matter response to applied nitrogen (69 lb./acre/annum) was greater in cocksfoot than in ryegrass. Fertilizer nitrogen had little effect on the yield of nitrogen from the sward except where the high seed rate of clover had been used.  相似文献   

18.
White clover varieties, potentially suitable for inclusion in seed mixtures for mixed stock-rearing farm systems, were evaluated when growing with S23 perennial ryegrass under rotational sheep grazing with a silage cut in late May or early June, as practiced on farms. Monoculture grass swards were also included to enable the direct and indirect contribution of white clover to total sward production to be evaluated.
White clover increased total sward production during all three years of the trial by an average of 50%. Although clover content of swards were similar, large differences occurred in their grass content, especially in the third year, when difference in total yields of swards based on large–leaved clover varieties was 2 t ha −1 while difference in clover yield was only 0-6 to ha–1 The indirect contribution of white clover, namely the extra grass resulting from N transfer, was greater in the spring than in the autumn. It was also greater for Nesta than for other varieties. and exceptional for this variety in that the increase in grass yield above that of grass monoculture was maintained over three harvest years.  相似文献   

19.
Plots of white clover ( Trifolium repens ) and perennial ryegrass ( Lolium perenne ) white clover were sown with and without fungicide seed treatment, seed treatment with Cytozyme (a proprietary growth-promoting agent) and pesticide granules (carbofuran or carbosulfan) mixed with the seed. Sowings were made in April/May at Hurley (1986 and 1987), Dorchester (1987) and Bronydd Mawr (1987). Only treatments containing pesticide significantly improved seedling emergence or herbage yield in the 1986 sowing. In 1987, pesticide greatly increased seedling emergence of ryegrass and clover at Hurley and Dorchester, but had no effect at Bronydd Mawr. At Hurley, herbage yield and the proportion of clover were more than doubled for up to a year, by chemical treatment after sowing in 1986.
Populations of plant parasitic nematodes were recorded at all sites and it is suggested that at Hurley and Dorchester seedlings with nematode damage were unable to survive the dry soil conditions that prevailed. In a subsequent pot experiment, pesticide granules controlled root endoparasitic nematodes and significantly increased seedling emergence of white clover in dry soil but had no effect in moist soil.  相似文献   

20.
Plants of a white clover population, Ac 3785, were grown in binary mixtures with perennial ryegrass cv. Talbot and the 'coexisting' population, Ba 10281, in boxes. Treatments imposed were 'no root competition' and 'full competition', which prevented or allowed the root systems of the competing species to intermingle respectively. Shoot competition was permitted in both treatments. Measurements of dry-matter yield and leaf area of the clover and grasses were made on three occasions following sowing.
The dry-matter yield of both grasses was higher than that of clover at all three harvests and under both treatments, and this yield advantage tended to increase with time. Both grasses yielded more under full competition than where root competition was prevented. Talbot was higher yielding than the coexisting population under both treatments. Clover dry weight and leaf area index (LAI) were considerably higher under the 'no root competition' treatment. There was no effect of companion grass on clover dry weight, but clover LAI was significantly higher with its coexisting population than with Talbot in the 'no root competition' treatment.
The LAI and dry weight data indicate that clover leaf morphology changed in response to the companion grass where root competition was prevented. Thus, the superior aerial compatibility of Ac 3785 with its coexisting grass was limited by below-ground factors. Possible ecological reasons for this effect are discussed.  相似文献   

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