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1.
A dentary of the oldest known monotreme, the Early Cretaceous Teinolophos trusleri, has an internal mandibular trough, which in outgroups to mammals houses accessory jaw bones, and probable contact facets for angular, coronoid, and splenial bones. Certain of these accessory bones were detached from the mandible to become middle ear bones in mammals. Evidence that the angular (homologous with the mammalian ectotympanic) and the articular and prearticular (homologous with the mammalian malleus) bones retained attachment to the lower jaw in a basal monotreme indicates that the definitive mammalian middle ear evolved independently in living monotremes and therians (marsupials and placentals).  相似文献   

2.
The separation of the middle ear bones from the mandible is considered a defining feature of all modern mammals. But did this event happen once in a primitive mammalian ancestor or independently in the monotreme lineage and therian (marsupial and placental) lineage? As Martin and Luo discuss in their Perspective, a new fossil-the dentary bone of an ancient toothed monotreme-suggests that the middle ear bones formed independently in these two mammalian lineages, providing a remarkable example of homoplastic evolution.  相似文献   

3.
Wang Y  Hu Y  Meng J  Li C 《Science (New York, N.Y.)》2001,294(5541):357-361
An ossified Meckel's cartilage has been recovered from two early Cretaceous mammals from China. This element is similar to Meckel's cartilage in prenatal and some postnatal extant mammals and indicates the relationship of Meckel's cartilage with the middle ear in early mammals. The evidence shows that brain expansion may not be the initial factor that caused the separation of postdentary bones from the dentary as middle ear ossicles during mammalian evolution. The failure of the dentary to seize reduced postdentary elements during ontogeny of early mammals is postulated as an alternative mechanism for the separation. Modifications of both feeding and hearing apparatuses in early mammals may have led to the development of the definitive mammalian middle ear.  相似文献   

4.
Ji Q  Luo ZX  Yuan CX  Tabrum AR 《Science (New York, N.Y.)》2006,311(5764):1123-1127
A docodontan mammaliaform from the Middle Jurassic of China possesses swimming and burrowing skeletal adaptations and some dental features for aquatic feeding. It is the most primitive taxon in the mammalian lineage known to have fur and has a broad, flattened, partly scaly tail analogous to that of modern beavers. We infer that docodontans were semiaquatic, convergent to the modern platypus and many Cenozoic placentals. This fossil demonstrates that some mammaliaforms, or proximal relatives to modern mammals, developed diverse locomotory and feeding adaptations and were ecomorphologically different from the majority of generalized small terrestrial Mesozoic mammalian insectivores.  相似文献   

5.
Many hypotheses have been postulated regarding the early evolution of the mammalian brain. Here, x-ray tomography of the Early Jurassic mammaliaforms Morganucodon and Hadrocodium sheds light on this history. We found that relative brain size expanded to mammalian levels, with enlarged olfactory bulbs, neocortex, olfactory (pyriform) cortex, and cerebellum, in two evolutionary pulses. The initial pulse was probably driven by increased resolution in olfaction and improvements in tactile sensitivity (from body hair) and neuromuscular coordination. A second pulse of olfactory enhancement then enlarged the brain to mammalian levels. The origin of crown Mammalia saw a third pulse of olfactory enhancement, with ossified ethmoid turbinals supporting an expansive olfactory epithelium in the nasal cavity, allowing full expression of a huge odorant receptor genome.  相似文献   

6.
A full understanding of primate morphological and genomic evolution requires the identification of their closest living relative. In order to resolve the ancestral relationships among primates and their closest relatives, we searched multispecies genome alignments for phylogenetically informative rare genomic changes within the superordinal group Euarchonta, which includes the orders Primates, Dermoptera (colugos), and Scandentia (treeshrews). We also constructed phylogenetic trees from 14 kilobases of nuclear genes for representatives from most major primate lineages, both extant colugos, and multiple treeshrews, including the pentail treeshrew, Ptilocercus lowii, the only living member of the family Ptilocercidae. A relaxed molecular clock analysis including Ptilocercus suggests that treeshrews arose approximately 63 million years ago. Our data show that colugos are the closest living relatives of primates and indicate that their divergence occurred in the Cretaceous.  相似文献   

7.
Phoberomys is reported to be the largest rodent that ever existed, although it has been known only from isolated teeth and fragmentary postcranial bones. An exceptionally complete skeleton of Phoberomys pattersoni was discovered in a rich locality of fossil vertebrates in the Upper Miocene of Venezuela. Reliable body mass estimates yield approximately 700 kilograms, more than 10 times the mass of the largest living rodent, the capybara. With Phoberomys, Rodentia becomes one of the mammalian orders with the largest size range, second only to diprotodontian marsupials. Several postcranial features support an evolutionary relationship of Phoberomys with pakaranas from the South American rodent radiation. The associated fossil fauna is diverse and suggests that Phoberomys lived in marginal lagoons and wetlands.  相似文献   

8.
Jepsen GL 《Science (New York, N.Y.)》1966,154(3754):1333-1339
A fossil skeleton of an early Eocene bat, the oldest known flying mammal, was found in southwest Wyoming. The bat is assigned to the new species Icaronycteris index of the suborder Microchiroptera. It was apparently of a young male whose body was buried in varved marls of the Green River Formation, on the bottom of Fossil Lake, about 50 million years ago. The bones, some as slender as a human hair, show a few "primitive" characteristics such as a clawed index finger and a complete phalangeal formula, but the bat was fully developed -an anatomically precocious contemporary of the dog-sized polydactylous horse.  相似文献   

9.
Partial skeletons of four species of extinct carnivores have been found in their dens excavated in the floodplain of an early Miocene ephemeral braided stream at Agate Fossil Beds National Monument, Nebraska. Bear dogs (Carnivora: Amphicyonidae) were the principal occupants; their den dimensions and body size are similar to those of living wolves and hyenas. Discovery of this predator community extends the record of denning behavior of large mammalian carnivores to the early Miocene, 20 million years ago.  相似文献   

10.
Deep-sea drilling in the Southern Ocean south of Australia and New Zealand shows that the Circum-Antarctic Current developed about 30 million years ago in the middle to late Oligocene when final separation occurred between Antarctica and the continental South Tasman Rise. Australia had commenced drifting northward from Antarctica 20 million years before this.  相似文献   

11.
Rhenium and osmium concentrations and the osmium isotopic compositions of iron meteorites were determined by negative thermal ionization mass spectrometry. Data for the IIA iron meteorites define an isochron with an uncertainty of approximately +/-31 million years for meteorites approximately 4500 million years old. Although an absolute rheniumosmium closure age for this iron group cannot be as precisely constrained because of uncertainty in the decay constant of (187)Re, an age of 4460 million years ago is the minimum permitted by combined uncertainties. These age constraints imply that the parent body of the IIAB magmatic irons melted and subsequently cooled within 100 million years after the formation of the oldest portions of chondrites. Other iron meteorites plot above the IIA isocbron, indicating that the planetary bodies represented by these iron groups may have cooled significantly later than the parent body of the IIA irons.  相似文献   

12.
Seventeen of eighteen orders of living placental mammals are not known before 65 million years ago. The monophyly of each order is well established, but interrelations have been less certain. A superordinal grouping of up to seven extant orders plus a variety of extinct orders, all included within Ungulata ("hoofed" mammals), can be linked to Late Cretaceous mammals from the 85-million-year-old Bissekty Formation, Uzbekistan (and, less certainly, North America and Europe), thus pushing the origin of this major clade back by 20 million years. Ungulatomorphs are not closely related to primates, rodents, or rabbits.  相似文献   

13.
The emergence of the seed habit in the Middle Paleozoic was a decisive evolutionary breakthrough. Today, seed plants are the most successful plant lineage, with more than 250,000 living species. We have identified a middle Givetian (385 million years ago) seed precursor from Belgium predating the earliest seeds by about 20 million years. Runcaria is a small, radially symmetrical, integumented megasporangium surrounded by a cupule. The megasporangium bears an unopened distal extension protruding above the multilobed integument. This extension is assumed to be involved in anemophilous pollination. Runcaria sheds new light on the sequence of character acquisition leading to the seed.  相似文献   

14.
Pliocene and Pleistocene fossil localities near the western shoreline of Lake Turkana, ranging in age between 1 million and 3.5 million years in age, have produced important new hominid specimens including most of a Homo erectus skeleton and a relatively complete early robust australopithecine cranium. The lacustrine, fluviatile, and terrestrial strata are designated the Nachukui Formation, which is subdivided into eight members. The distribution of sedimentary facies within the Nachukui Formation suggests that, as today, the Labur and Murua Rith ranges formed the western margin of the basin and were drained by eastward-flowing rivers that fed into the forerunner of the present lake or a major river system. There is also stratigraphic evidence for tectonic movement during the deposition of these sediments. Twenty-three of the tuffs observed in the succession occur also in the Koobi Fora Formation east of the lake and in the Shungura Formation of the lower Omo Valley and permit precise correlation among these three localities. Fortyseven fossiliferous sites from West Turkana have yielded more than 1000 specimens of 93 mammalian species. The mammalian fossils represent nine sequential assemblages that augment information about faunal and environmental change from elsewhere in the basin.  相似文献   

15.
Gartner S 《Science (New York, N.Y.)》1970,169(3950):1077-1079
Evidence from leg 2 of the Deep Sea Drilling Project suggests a constant spreading rate for the floor of the North Atlantic over the past 80 million years; a major lowering of the carbonate dissolution level during the early Pliocene; and an early to middle Eocene age for horizon A.  相似文献   

16.
Laterally extensive black shales were deposited on the S?o Francisco craton in southeastern Brazil during low-latitude Neoproterozoic glaciation approximately 740 to 700 million years ago. These rocks contain up to 3.0 weight % organic carbon, which we interpret as representing the preserved record of abundant marine primary productivity from glacial times. Extractable biomarkers reflect a complex and productive microbial ecosystem, including both phototrophic bacteria and eukaryotes, living in a stratified ocean with thin or absent sea ice, oxic surface waters, and euxinic conditions within the photic zone. Such an environment provides important constraints for parts of the "Snowball Earth" hypothesis.  相似文献   

17.
It is not possible at present to demonstrate hominid occupation of southern Africa prior to the middle or late Pliocene, perhaps 3 million years ago. It may be the case that much, if not most, of the subcontinent was in fact uninhabited before that. The earliest hominid known to have lived in southern Africa is Australopithecus africanus. It was apparently replaced by Homo (?evolved into Homo) by 2 million years ago, at approximately the same time as A. robustus is first recorded locally. Homo and A. robustus then coexisted until perhaps 1 million years ago, after which Homo survived alone. There is no solid evidence that either of the southern African australopithecines made tools or accumulated bones. In fact, at the known sites, it now seems more likely that the bones, including those of the australopithecines themselves, were accumulated by carnivores. The known archeological record of southern Africa begins 2 million to 1.5 million years ago and the oldest stone tools may belong to the Oldowan Industry. Far better documentation exists for the succeeding Acheulean Industrial Complex, which was present in southern Africa almost certainly before 1 million years ago and persisted with modifications probably until sometime between 300,000 and 130,000 years ago. Although it is known that Acheulean peoples made handaxes, cleavers, and other stone tools, very little else is known about the activities of Acheuleans in southern Africa. Far more is known about their Middle and Later Stone Age successors. Southern African MSA peoples were perhaps among the earliest anywhere to take systematic advantage of aquatic resources for their subsistence, although they apparently did so far less effectively than did the LSA peoples who followed them. There are also contrasts between the ways in which MSA and LSA peoples dealt with terrestrial prey and between the contents of MSA and LSA artifact assemblages. The LSA peoples, for example, seem to have made much more extensive use of bone as a raw material, and they were the first to manufacture articles that are clearly interpretable as ornaments or art objects. From an evolutionary perspective, the LSA may represent a quantum advance over the MSA, perhaps correlated with the replacement of an archaic human physical type by the modem one. However, this must remain only a working hypothesis until much more is learned about the earliest LSA, dating to 35,000 to 40,000 years ago or more, and until there are adequate samples of well-provenienced MSA and early LSA physical remains. The later LSA, postdating 20,000 to 18,000 years ago, is reasonably well known. Later LSA peoples were probably at least partly responsible for the extinction of several large mammals in southern Africa about 10,000 years ago. By that date or shortly thereafter, at least some LSA peoples established basic hunting-gathering adaptations, which continued until the introduction and spread of agriculture and pastoralism, beginning roughly 2000 years ago. Thereafter, hunters and gatherers became progressively restricted in numbers and distribution, such that today only a very few exist, restricted to some of the most marginal environments of the subcontinent. It remains a major goal of southern African archeology to shed more light on the evolution and operation of hunting-gathering cultures during the vast time span when they covered all of southern Africa.  相似文献   

18.
Excavations have demonstrated that stone artifacts occur stratified within beds of Lower Pleistocene (or end Pliocene) age. At one site a low-density scatter of worked stone objects occurs together with small but significant quantities of broken-up bones. Potassium-argon dates indicate an age greater than 2 million years; thus, this may be the oldest known hominid occupation site. More than 20 hominid fossils have been recovered from various sedimentary formations in the area.  相似文献   

19.
It has been suggested that variation in brain structure correlates with the sizes of individuals' social networks. Whether variation in social network size causes variation in brain structure, however, is unknown. To address this question, we neuroimaged 23 monkeys that had been living in social groups set to different sizes. Subject comparison revealed that living in larger groups caused increases in gray matter in mid-superior temporal sulcus and rostral prefrontal cortex and increased coupling of activity in frontal and temporal cortex. Social network size, therefore, contributes to changes both in brain structure and function. The changes have potential implications for an animal's success in a social context; gray matter differences in similar areas were also correlated with each animal's dominance within its social network.  相似文献   

20.
Microtektites have been recovered from three horizons in eight middle Eocene to middle Oligocene marine sediment sequences. Five of these occurrences are coeval and of latest Eocene age (37.5 to 38.0 million years ago); three are coeval and of early late Eocene age (38.5 to 39.5 million years ago); and three are of middle Oligocene age (31 to 32 million years ago). In addition, rare probable microtektites have been found in sediments with ages of about 36.0 to 36.5 million years. The microtektite horizon at 37.5 to 38.0 million years can be correlated with the North American tektite-strewn field, which has a fission track age (minimum) of 34 to 35 million years and a paleomagnetic age of 37.5 to 38.0 million years. There is no evidence for mass faunal extinctions at any of the microtektite horizons. Many of the distinct faunal changes that occurred in the middle Eocene to middle Oligocene can be related to the formation of the Antarctic ice sheet and the associated cooling phenomena and intensification of bottom currents that led to large-scale dissolution of calcium carbonate and erosion, which created areally extensive hiatuses in the deep-sea sediment records. The occurrence of microtektite horizons of several ages and the lack of evidence for faunal extinctions suggest that the effects of extraterrestrial bolide impacts may be unimportant in the biologic realm during middle Eocene to middle Oligocene time.  相似文献   

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