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1.
Gert J van den Berg Steven D Verryn Paxie W Chirwa Francois van Deventer 《Southern Forests》2016,78(3):209-216
In South Africa, Eucalyptus urophylla is an important species due to its disease tolerance to fungal diseases such as Crysoporthe austroafricana and the Coniothyrium sp. cankers. It is mainly planted as a parental species in a hybrid combination with E. grandis. Generally, the E. grandis × E. urophylla hybrid has better disease tolerance and higher wood density than pure E. grandis. The current strategy is to maintain large breeding populations of both parental species in order to provide improved elite selections for hybrid crosses on a regular basis. With this in mind, two E. urophylla populations, consisting of five provenance/progeny trials, were established in the subtropical region of Zululand. The aims of this study were, firstly, to determine the magnitude of genotype × environment interaction of E. urophylla in Zululand; secondly, to estimate genetic parameters and correlations for diameter at breast height (DBH), height and volume; and thirdly, to identify selections to advance the current breeding population as well as to hybridise with E. grandis. Results indicated that genotype × environment interaction effects would be practically negligible for growth in Zululand and therefore a single breeding population will be appropriate. In general, all growth traits were under low to moderate genetic control, with narrow-sense heritabilities ranging between 0.14 and 0.48 for volume. The genetic correlations between growth traits were high (0.98 and 0.99 for DBH– volume). This is an indication that DBH is a sufficient growth measure to use in E. urophylla breeding programmes. Best linear unbiased prediction estimates indicated that aa selection scenario of 200 individuals will generate genetic gains of 44.7% over the population mean. The estimated gains for the top 50 individuals that could potentially be used as hybrid parents to cross with E. grandis was 59.8% over the population mean. 相似文献
2.
《Southern Forests》2013,75(2):147-153
Tree diameter under and over bark at breast height (dbh), wood density and bark thickness were assessed on samples from control-pollinated families of Eucalyptus grandis, E. urophylla, E. grandis × E. urophylla and E. urophylla × E. grandis. The material was planted in field trials in the coastal Zululand region of South Africa. At 75 months, between three and seven of the best trees per family were felled and wood samples collected. Genetic parameters for wood density, bark thickness and bark percentage (ratio of double bark thickness to overbark diameter) and the inter-trait correlations for the different species and hybrids were calculated. Genetic parameter estimates for wood density, bark thickness and bark percentage in the E. urophylla × E. grandis hybrids showed these traits to be under total additive genetic control. This was confirmed by the intermediate hybrid means for these traits relative to those of the parental species. There was a very low correlation between dbh and wood density for the E. urophylla × E. grandis hybrids (rG = –0.07 and rP = 0.064). Amongst the E. urophylla families there was a moderate positive and significant phenotypic correlation between wood density and bark thickness (rP = 0.391), and between wood density and bark percentage (rP = 0.442). 相似文献
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4.
The Forestry Commission of Zimbabwe initiated a hybrid breeding program of Eucalyptus grandis with E. tereticornis (G × T) and E. camaldulensis (G × C) in order to provide suitable genotypes for planting in areas that are marginal for E. grandis in terms of drought and frost. A total of seven hybrid trials were established at four sites, representative of low to medium rainfall. The female parents were from E. grandis preselected for superior volume production. No pure species were included in the trials to act as controls and allow for estimation of hybrid vigour. Survival, growth and stem straightness were assessed at 18 and 43 months from planting. Survival was generally above 80% for all hybrids across sites. G × C hybrids performed better than G × T hybrids, showing a 20% difference in mean height at 43 months. Female and male variances were generally insignificant (P > 0.05) and in some cases, the estimates were 0. This was not unexpected, as the parents for the hybrids were preselected for superior height, diameter at breast height (DBH) and volume production as pure species. Ratio of dominance to phenotypic variance ( $ \hat{d}^{2} $ ) was significant for all traits for all hybrids (P < 0.05). Trait–trait dominance correlations (r D ) at 43 months between height and DBH were generally large and positive, above 0.6. Type-B dominance correlation (r BD ) suggested evidence of genotype-by-environment interaction (G × E), but one high elevation site contributed most to the observed G × E. If these results are confirmed in larger mating designs with progeny established on multiple sites, it seems that the most appropriate breeding strategy for E. grandis × E. tereticornis and E. grandis × E. camaldulensis hybrids may be one that exploits both additive and dominance genetic variance, such as the reciprocal recurrent selection (RRS) scheme or RRS with forward selection. 相似文献
5.
In South Africa, there is currently a phenomenon where a Eucalyptus grandis × E. urophylla clone is dying on a large scale, whereas co-occurring clones in the same plantation area remain healthy. No primary biotic pathogen has been isolated from these dying trees. However, the plantation region has endured severe drought conditions. We hypothesised that the specific E. grandis × E. urophylla clone dieback is due in part to the affected trees having an elevated drought vulnerability. We tested this hypothesis retrospectively by examining how the dying (branch dieback started) as compared with two healthy and co-occurring E. grandis × E. urophylla clones vary in branch xylem anatomy (water transport) and leaf stable carbon isotopes (gas exchange) across two levels of mean annual precipitation in the planted landscape. There was significant intra-hybrid variation in the range of studied hydraulic traits. It appeared that the dying clone is maladapted to drier field conditions relative to the two healthy growing clones. Individuals of the drought-susceptible clone had increased hydraulic conductivity, having both wider vessel lumens and larger lumen fractions, and also did not regulate leaf-level transpiration in the drier landscape as efficiently as the two unaffected clones (lower δ13C ratios). In turn, one of the healthy growing clones had a completely different hydraulic strategy, having higher wood density, lower lumen fractions, and higher δ13C ratios that are widely considered as hydraulically resilient during water stress. The other clone appeared to also gain drought tolerance via higher leaf-level water-use efficiency, although drought tolerance was less clear regarding xylem anatomy. We highlight how variation in relative drought vulnerability can be highly significant even between clones within a single hybrid group. Knowledge of this variation in plant hydraulics between commercially planted and closely related Eucalyptus trees would complement clone deployment programmes in a target landscape. 相似文献
6.
Izette Greyling Michael J Wingfield Martin PA Coetzee Seonju Marincowitz 《Southern Forests》2016,78(2):123-129
Species of Teratosphaeria include some of the most important fungal pathogens of plantation-grown eucalypt trees. During routine disease surveys, symptoms and signs of leaf spot and blight were observed on the foliage of one-year-old E. grandis × E. urophylla hybrids in the Zululand region of KwaZulu-Natal, South Africa. These were distinct from those caused by the well-known and leaf-infecting fungus Teratosphaeria suttonii, which is not considered an important pathogen in the country. Culture and morphological characteristics as well as DNA sequences for three gene regions were used to compare the fungus isolated from the newly emerging symptoms with those for known Teratosphaeria species. DNA sequences were the same as those for T. destructans and this was consistent with the distinctive morphology of the asexual spores and the symptoms on leaves. Teratosphaeria destructans is an aggressive pathogen and actions will be needed to ensure that it does not impart serious losses to the local forestry industry. 相似文献
7.
Gert J van den Berg Steven D Verryn Paxie W Chirwa Francois van Deventer 《Southern Forests》2018,80(1):9-19
Conventionally, Eucalyptus grandis × E. urophylla (GU) hybrid material is first tested as seedlings in progeny trials for at least four years before ortets are selected and ramets of the selected ortets are propagated to test in clonal trials. The primary constraint with this ‘conventional hybrid breeding strategy’ (CHBS) is the time required to first test the hybrid material as seedlings. In order to address this, an ‘accelerated hybrid breeding strategy’ (AHBS) was investigated to reduce the time spent on testing GU hybrid material as seedlings. However, it is of utmost importance to quantify the impact the AHBS might have on genetic gains and genetic information. With this in mind, two clonal populations have been established with genetic material that derived from the CHBS and the AHBS. The main purpose of this study was to do a comparative study between the CHBS and AHBS, firstly to quantify the genetic gains per unit time for GU hybrid clonal populations that have been derived from the CHBS and AHBS respectively, and secondly to obtain genetic parameters such as heritabilities, the ratio of dominance, clonal within family variance, and the proportion of additive and non-additive genetic variance. The results indicated that the percentage realised volume gains per year was higher for the AHBS (3.7%) than for the CHBS (1.9%) when compared with the GU commercial clone. Thus, shortening the testing time of GU seedlings had a positive impact on volume gains per year. With regard to genetic parameters, both the AHBS and CHBS clonal populations indicated that non-additive genetic variation explained the majority (88% and 71%, respectively) of the genetic variation. Due to the pre-eminence of non-additive genetic variation, the narrow-sense heritabilties for the female and male effects were negligible for both clonal populations. Overall, the majority of the non-additive genetic variation was explained by the proportion of dominance variance, and less by the clone within family effect. These results suggest that, firstly, the time spent on testing GU hybrid material as seedlings should be minimised and, secondly, a hybrid breeding strategy to capture non-additive genetic variation should be adopted. 相似文献
8.
[目的]评估Resistograph钻刺法间接测定尾叶桉×细叶桉木材密度的可靠性,检测杂交亲本对子代表型的效应以及生长与木材密度的相关,评选速生、优质的尾细桉杂种。[方法]基于10株尾叶桉与10株细叶桉不完全析因交配产生的56个杂交组合的7.5年生试验林,利用79株分析容积法与Resistograph钻刺法测定的木材密度的相关,通过方差分析检测亲本对杂种生长和木材密度的效应,结合多重比较和独立淘汰法进行材积和木材密度的联合选择。[结果]容积法与Resistograph钻刺法测定的木材密度的表型相关系数为0.52(P0.001),遗传相关系数为0.55(P0.05);树高、胸径及材积的母本间和父本间均呈极显著差异(P0.001或0.01),但母本×父本互作的效应不显著;对钻刺木材密度,父本间呈极显著差异(P0.001),母本×父本互作显著(P0.05),但母本间差异不显著;树高、胸径和材积间的表型相关和遗传相关均极显著(P0.001),其与钻刺木材密度的表型相关极显著(P0.001),但遗传相关不显著;评选出速生、木材密度较高的杂交组合14个、单株17株。[结论]Resistograph钻刺法是一种间接测定尾细桉木材密度的简便、经济和可靠的方法;母本和父本选择以及母本与父本的组配对培育速生、材质优良的尾细桉杂种均较重要;尾细桉生长与木材密度的遗传相关不显著,需要对这两类性状分别进行选择;评选的尾细桉杂交组合和单株为培育速生、优质的桉树良种提供了有用的材料。 相似文献
9.
Garel Makouanzi Gilles Chaix Sophie Nourissier Philippe Vigneron 《Southern Forests》2018,80(2):151-158
Thirty-four Eucalyptus urophylla × Eucalyptus grandis hybrids were evaluated with a view to selecting for improved growth and wood-quality traits for plantations in the Congo. Height, circumference at breast height and volume were measured at 12, 27, 37, 49 and 60 months. Lignin content, the syringyl/guaiacyl ratio and total extractives content were predicted by near-infrared spectroscopy using wood powder samples collected from trees at breast height. While wood chemical properties were stable and under strong genetic control, growth traits were not. The genetic correlation between lignin content and growth was weak and negative, whereas the environmental correlation was also weak but positive. The genetic improvement of E. urophylla × E. grandis clones, based on growth features, leads to a limited decrease in lignin content and syringyl content and to a limited increase in extractives content. 相似文献
10.
《Southern Forests》2013,75(4):185-188
Eucalyptus grandis × Eucalyptus nitens (G×N) hybrid clones are selected to combine complimentary characteristics of E. nitens and E. grandis. G×N hybrid clones also have the potential to increase growth rates and provide adaptability to a changing climate. A series of three trials planted across a range of high-productivity, mid-altitude sites in the KwaZulu-Natal midlands was established to test the suitability of a suite of G×N hybrid clones. Seven commercially available G×N clones together with two widely planted pure species controls, Eucalyptus dunnii and Eucalyptus smithii, were chosen for this study. The trials were set out in random complete block designs with nine treatments and four replications set out in square plots of 25 trees. The trials were measured for diameter at breast height at 23 months and monitored for snow damage over two winter seasons. Snow was noted at the Baynesfield site and subsequent snow damage assessments were performed. Basal area per hectare was calculated for each plot as a function of diameter at breast height and survival. Significant differences were observed between G×N hybrid clones in both growth and snow tolerance. The top-performing clones significantly outperformed both pure species controls in terms of growth and snow tolerance. Early results indicate that G×N hybrids may be better suited to high-potential, mid-altitude sites exposed to light snow risk than the currently recommended pure species. 相似文献
11.
[目的]通过尾叶桉与赤桉杂种F1材质性状的遗传参数估算及其分析,为桉树杂交育种材性改良亲本选配和交配设计提供理论依据。[方法]以尾叶桉与赤桉6×6正反析因交配杂种F1测定林为材料,并以相应亲本自由授粉的半同胞子代作对照。9年生时,对各小区的杂种和家系(5株小区,6次重复)选取2株平均木,在胸高1. 3 m处沿南北向钻取木芯,共984个,测定木材基本密度、纤维长、纤维宽和纤维长宽比,利用ASReml-R估算4个材质指标的特殊配合力、杂种优势、父母本单株狭义遗传力、遗传相关和表型相关系数。[结果]表明:尾叶桉与赤桉正反交的杂种木材基本密度优于对照,呈明显的杂种优势;反交组合杂种其木材基本密度优于正交组合杂种,正交组合杂种的纤维特性优于反交组合的;在正交组合杂种中,木材基本密度、纤维长和纤维长宽比的母本效应低于父本效应;而反交组合杂种中,4个材质指标的母本效应均高于父本效应。木材基本密度和纤维特性指标受中至低遗传控制。正反交组合杂种中,除纤维长宽比外,其余3个材质指标的显性效应大于加性效应。对于遗传相关,杂种的木材基本密度与纤维宽呈极显著的负相关,与纤维长宽比呈极显著的正相关,与纤维长呈不显著负相关,纤维长与纤维宽呈显著正相关。对于表型相关,除纤维宽与纤维长宽比呈不显著正相关外,其余性状间均呈极显著正相关。[结论]尾叶桉与赤桉杂种的材质性状遗传差异因亲本和交配方式而异,表明通过种间杂交和正向选择进行材质性状的遗传改良具有潜力。 相似文献
12.
Lack of association between allozyme heterozygosity and juvenile traits in Eucalyptus 总被引：2，自引：0，他引：2
Genetic variability for juvenile traits, which included basal diameter, height, biomass accumulation, and growth increment, was studied in eight provenances involving four species, Eucalyptus grandis, E. saligna, E. camaldulensis and E. urophylla, under uniform greenhouse conditions. The species differed significantly for all juvenile traits including individual mean allozyme heterozygosity, and high levels of variability were observed within and among species, which indicated that scope for selection exists for these traits. There was considerable reduction in genetic variability for all traits at 6 months of age when compared to that observed at 4 months. Eucalyptus saligna recorded the highest mean for basal diameter and biomass accumulation, while E. camaldulensis exhibited the highest mean for height. The two E. camaldulensis provenances exhibited the highest individual mean heterozygosities (0.425 and 0.417), followed by E. saligna (0.296 and 0.258), E. urophylla (0.254 and 0.233), and E. grandis (0.300 and 0.163). The juvenile traits were correlated with individual mean allozyme heterozygosity to examine the possibility of developing a genotypic basis for early selection of potentially superior individuals in a tree improvement program. 相似文献
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《Southern Forests》2013,75(2):141-146
Tree breeders attempt to predict the genetic gains that are likely to be achieved through selection and breeding of new generations, using stochastic or deterministic modelling. There are many factors that may cause a discrepancy between the predicted and realised genetic gains. Often the predictions for genetic gains are based on single trait selection, whereas in reality the breeding tends to be multitrait in nature. The violation of Hardy-Weinberg conditions, assumptions regarding outcrossing and relatedness, assumptions regarding the effect of the interaction between the environment and the genotype, and numerous possible errors in the process of breeding, all could result in unexpected discrepancies between the realised and predicted genetic gains. A series of genetic gains trials containing representatives of three generations of Eucalyptus grandis selections were compared with the view to verifying the effectiveness of the E. grandis breeding program. Genetic gains of the F3 (third generation of pedigreed progeny) over the F2 generation (second generation of pedigreed progeny) were 15% for tree growth (volume). A comparison between F2 and P0 revealed an improvement of between 20% and 33% for growth. This exercise highlighted complexities of modelling the predicted genetic gains of assimilated genetic breeding trials. The predictions of genetic gains did deviate (in both directions) from those realised, although these deviations may be explained as functions of imperfect modelling. On average, however, the predicted genetic gains for tree volume over three generations was 13% between generations, whereas the average realised genetic gain in the genetic gains trial was 14%. It is therefore assumed that the E. grandis breeding population is indeed performing as expected, following classical tree breeding assumptions. 相似文献
15.
《Southern Forests》2013,75(3):105-111
This study describes the stomatal response occurring during water stress and subsequent recovery of three Eucalyptus grandis clonal hybrids. The aim was to investigate the degree to which stomatal conductance (g s) and stomatal density differ between the clonal hybrids across seasons and in response to water stress. Plants from one E. grandis × E. camaldulensis (GC) and two E. grandis × E. urophylla (GU1 and GU2) clones were grown for 18 months in 80 l planting bags. Plants were subjected to three watering treatments: control (100% field capacity), chronic water stress (maintained at 15% of field capacity) and acute water stress (cyclic water stress, where water was withheld until leaf wilting point, and a subsequent period of recovery followed). Stomatal conductance was measured after 6, 12 and 18 months growth. At 12 months of age, the recovery of g s 1, 2 and 7 d after rewatering (following acute water stress) was further investigated. The GC hybrid showed consistently higher g s than the GU clones at each measurement period. Stomatal conductance was 24–66% higher during winter (after 12 months growth) than during summer. The recovery of stomatal conductance from acute water stress was more rapid in the GC clone than the GU clones. Chronic water stress was shown to decrease g s in GU clones by up to 70%, but not in the GC clone. Water stress did not affect stomatal density or size. Remarkably, stomata were absent from the adaxial leaf surface of clone GU1 leaves, but not from the leaves of the other E. urophylla hybrid cross (GU2). Total biomass of the GC clone was significantly greater at 9 months growth, but after 18 months growth the GU1 clone had attained greater biomass accumulation (although not significantly). Measurement of g s, transpiration, stomatal density and total biomass in the GU1 clone indicated stomatal sensitivity to water stress, a favourable trait during periods of drought. The differing growth strategies of the GU and GC clones could be partially explained by their differences in stomatal sensitivity in response to water stress. 相似文献
16.
试论世界桉树栽培现状和我国桉树人工林发展策略 总被引：4，自引：0，他引：4
桉树是世界上热带、亚热带地区最重要的人工林树种之一,其面积已超过600万ha。文中着重介绍了巴西发展按树的经验,并提出了我国应采取的对策。 相似文献
17.
Guoqing Jin Guofeng Qin Weihong Liu Deyu Chu Suzhou Hong Zhichun Zhou 《Frontiers of Forestry in China》2009,4(1):101-106
Two groups of filial generations derived from two different Pinus massoniana complete-diallel crosses were analyzed. Results show that the general combining ability (GCA), specific combining ability
(SCA) and reciprocal effects were significant for some growth traits, including height, DBH and volume index. The heredity
of these growth traits was controlled by additive and non-additive genes, of which the additive genes played a dominant role.
The epistatic effect was greater for group I (cross in 1992) than group II (cross in 1993). The SCA of P. massoniana growth traits was significantly greater than GCA, which may be related to different geographical provenance for parents and
the indirect selection by GCA. Inbreeding depression was commonly observed for P. massoniana growth traits. The extent of
inbreeding depression was −17.8%–−18.4%, −23.3%–−27.7% and −44.3%–−50.6% for height, DBH and volume index, respectively. It
was observed that parents with small GCA values exhibited a greater extent of inbreeding depression. Large differences in
hybrid vigor of different crosses were observed and the difference between original cross and reciprocal cross was not significant.
Based on the volume index, 10 fine crosses were selected for two groups respectively, and the average increment of volume
index was 59.41% and 41.76%, respectively, in comparison with the average of the testing groups, and was 100.58% and 74.61%
in comparison with the local commercial variety.
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Translated from Scientia Silvae Sinicae, 2008, 44(6): 28–33 [译自: 林业科学] 相似文献
18.
桉蝙蛾是近年来在广西、广东新发现严重危害桉树的蛀干害虫,本文记述了该害虫的形态特征和生物学特性,该虫在广西分布于12个市49个县,10个自治区直属国有林场,232个乡镇,危害桉树面积达2 894.42 hm2,多数个体1年完成1代,以幼虫于12月中下旬开始在受害树干中越冬,次年2-3月化蛹,4月羽化,5月上旬初孵幼虫出现,6-8月幼虫上树转移危害,1年只有1次羽化;该虫主要分布在丘陵山地,以沟谷和坡底危害较严重,主要危害1～2年生巨尾桉和尾巨桉无性系.1年生和2年生受害桉树单株材积生长量下降率分别为16.70％和24.78％.沟谷和坡底1～2年生速生桉无性系是防治的主要对象. 相似文献
19.
[目的]为了解雷州半岛尾巨桉速生人工林生态系统的C、N、P分配格局及化学计量特征。[方法]采用空间换时间的方法,选取雷州半岛4种不同林龄(1、3、5、7 a)的尾巨桉人工林为研究对象,对尾巨桉叶片、凋落物及土壤的C、N、P含量及化学计量特征进行测定分析。[结果]表明:C、N含量表现为叶片凋落物土壤,P含量表现为叶片土壤凋落物,且3个库间差异显著;土壤的C含量随林龄增加而增加,N、P含量差异不显著,土壤C∶N随林龄的增加而增加,说明土壤有机质分解速率逐渐下降;凋落物的C∶N为54. 07 92. 18 ( 25),表明尾巨桉林下凋落物分解速率较慢,N元素成为主要限制凋落物分解的元素,凋落物的C∶N随林龄的增加先增加后下降,凋落物分解速度先降低后升高;叶片的N∶P为10. 80 12. 98,说明中幼林龄尾巨桉受N限制较明显。相关性分析表明:凋落物养分元素含量受叶片限制,土壤养分含量受凋落物限制,表明生态系统内部C、N、P元素在植物、凋落物与土壤之间实现了运输和转换。[结论]雷州半岛尾巨桉中、幼林龄时期土壤有机质及凋落物分解速率较慢;随林龄的增加,土壤有机质、凋落物分解速率下降,N元素成为其主要分解限制性元素,林分生长受N限制明显。 相似文献
20.
《Southern Forests》2013,75(3-4):131-135
Hybrid clones with Eucalyptus grandis as one parent and, generally, E. urophylla as the other parent, are deployed commercially as monoclonal blocks by Sappi in the subtropical region of coastal Zululand, South Africa. In monoclonal blocks genotypes are grown in self-competition. Performance under this scenario may not necessarily correlate well with performance where genotypes are competing for resources with a number of genetically dissimilar genotypes. There is an uncertainty whether plot configuration or size influences clonal ranking and therefore selection of clones for commercialisation. Final results at age eight years of a trial testing eight selected Eucalyptus hybrid clones in single-tree, eight-tree line, and eight x eight tree square plots are reported in this paper. Ranking of the eight clones between the half rotation (44 months) and full rotation (94 months) measurement have remained stable in the single-tree and line plots, but there have been some changes in the square plots. At 94 months, clonal ranking between the single-tree and line plots was consistent (rp = 0.98) with only two clones changing ranks. When comparing the clonal ranking of the single-tree plots to the large square plots rank changes have been significant (rp = 0.65). The overall means for the three different plot types are very similar. The range of the clonal means for tree volume in the single-tree plots (0.4042 m3) is more than five times the range for the square plots (0.0709 m3). Gain predictions using data from the single-tree plots overestimated the realised gains measured in the square plots. The top-performing clone in the single-tree plots had a predicted gain of 74% over the trial mean, but only yielded 7% more in the square plots. Similarly, the worst-ranked clone in the single-tree plots did not perform as poorly as what was predicted (?59%) and produced only 13% less than the trial mean on the square plots. 相似文献