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1.
Woodland encroachment on United States rangelands has altered the structure and function of shrub steppe ecosystems. The potential community structure is one where trees dominate, shrub and herbaceous species decline, and rock cover and bare soil area increase and become more interconnected. Research from the Desert Southwest United States has demonstrated areas under tree canopies effectively store water and soil resources, whereas areas between canopies (intercanopy) generate significantly more runoff and erosion. We investigated these relationships and the impacts of tree encroachment on runoff and erosion processes at two woodland sites in the Intermountain West, USA. Rainfall simulation and concentrated flow methodologies were employed to measure infiltration, runoff, and erosion from intercanopy and canopy areas at small-plot (0.5 m2) and large-plot (13 m2) scales. Soil water repellency and vegetative and ground cover factors that influence runoff and erosion were quantified. Runoff and erosion from rainsplash, sheet flow, and concentrated flow processes were significantly greater from intercanopy than canopy areas across small- and large-plot scales, and site-specific erodibility differences were observed. Runoff and erosion were primarily dictated by the type and quantity of ground cover. Litter offered protection from rainsplash effects, provided rainfall storage, mitigated soil water repellency impacts on infiltration, and contributed to aggregate stability. Runoff and erosion increased exponentially (r2 = 0.75 and 0.64) where bare soil and rock cover exceeded 50%. Sediment yield was strongly correlated (r2 = 0.87) with runoff and increased linearly where runoff exceeded 20 mm·h?1. Measured runoff and erosion rates suggest tree canopies represent areas of hydrologic stability, whereas intercanopy areas are vulnerable to runoff and erosion. Results indicate the overall hydrologic vulnerability of sagebrush steppe following woodland encroachment depends on the potential influence of tree dominance on bare intercanopy expanse and connectivity and the potential erodibility of intercanopy areas.  相似文献   

2.
Land managers across the western United States are faced with selecting and applying tree-removal treatments on pinyon (Pinus spp.) and juniper (Juniperus spp.) woodland-encroached sagebrush (Artemisia spp.) rangelands, but current understanding of long-term vegetation and hydrological responses of sagebrush sites to tree removal is inadequate for guiding management. This study applied a suite of vegetation and soil measures (0.5 ? 990 m2), small-plot rainfall simulations (0.5 m2), and overland flow experiments (9 m2) to quantify the effects of mechanical tree removal (tree cutting and mastication) on vegetation, runoff, and erosion at two mid- to late-succession woodland-encroached sagebrush sites in the Great Basin, United States, 9 yr after treatment. Low amounts of hillslope-scale shrub (3 ? 15%) and grass (7 ? 12%) canopy cover and extensive intercanopy (area between tree canopies) bare ground (69 ? 88% bare, 75% of area) in untreated areas at both sites facilitated high levels of runoff and sediment from high-intensity (102 mm ? h? 1, 45 min) rainfall simulations in interspaces (~ 45 mm runoff, 59 ? 381 g ? m? 2 sediment) between trees and shrubs and from concentrated overland flow experiments (15, 30, and 45 L ? min? 1, 8 min each) in the intercanopy (371 ? 501 L runoff, 2 342 ? 3 015 g sediment). Tree cutting increased hillslope-scale density of sagebrush by 5% and perennial grass cover by twofold at one site while tree cutting and mastication increased hillslope-scale sagebrush density by 36% and 16%, respectively, and perennial grass cover by threefold at a second more-degraded (initially more sparsely vegetated) site over nine growing seasons. Cover of cheatgrass (Bromus tectorum L.) was < 1% at the sites pretreatment and 1 ? 7% 9 yr after treatment. Bare ground remained high across both sites 9 yr after tree removal and was reduced by treatments solely at the more degraded site. Increases in hillslope-scale vegetation following tree removal had limited impact on runoff and erosion for rainfall simulations and concentrated flow experiments at both sites due to persistent high bare ground. The one exception was reduced runoff and erosion within the cut treatments for intercanopy plots with cut-downed-trees. The cut-downed-trees provided ample litter cover and tree debris at the ground surface to reduce the amount and erosive energy of concentrated overland flow. Trends in hillslope-scale vegetation responses to tree removal in this study demonstrate the effectiveness of mechanical treatments to reestablish sagebrush steppe vegetation without increasing cheatgrass for mid- to late-succession woodland-encroached sites along the warm-dry to cool-moist soil temperature ? moisture threshold in the Great Basin. Our results indicate improved hydrologic function through sagebrush steppe vegetation recruitment after mechanical tree removal on mid- to late-succession woodlands can require more than 9 yr. We anticipate intercanopy runoff and erosion rates will decrease over time at both sites as shrub and grass cover continue to increase, but follow-up tree removal will be needed to prevent pinyon and juniper recolonization. The low intercanopy runoff and erosion measured underneath isolated cut-downed-trees in this study clearly demonstrate that tree debris following mechanical treatments can effectively limit microsite-scale runoff and erosion over time where tree debris settles in good contact with the soil surface.  相似文献   

3.
Land owners and managers across the western United States are increasingly searching for methods to evaluate and mitigate the effects of woodland encroachment on sagebrush steppe ecosystems. We used small-plot scale (0.5 m2) rainfall simulations and measures of vegetation, ground cover, and soils to investigate woodland response to tree removal (prescribed fire and mastication) at two late-succession woodlands. We also evaluated the effects of burning on soil water repellency and effectiveness of aggregate stability indices to detect changes in erosion potential. Plots were located in interspaces between tree and shrub canopies and on undercanopy tree and shrub microsites. Erosion from untreated interspaces in the two woodlands differed more than 6-fold, and erosion responses to prescribed burning differed by woodland site. High-intensity rainfall (102 mm · h-1) on the less erodible woodland generated amplified runoff and erosion from tree microsites postfire, but erosion (45–75 g · m-2) was minor relative to the 3–13-fold fire-induced increase in erosion on tree microsites at the highly erodible site (240–295 g · m-2). Burning the highly erodible woodland also generated a 7-fold increase in erosion from shrub microsites (220–230 g · m-2) and 280–350 g · m-2 erosion from interspaces. High levels of runoff (40–45 mm) and soil erosion (230–275 g · m-2) on unburned interspaces at the more erodible site were reduced 4–5-fold (10 mm and 50 g · m-2) by masticated tree material. The results demonstrate that similarly degraded conditions at woodland-encroached sites may elicit differing hydrologic and erosion responses to treatment and that treatment decisions should consider inherent site-specific erodibility when evaluating tree-removal alternatives. Strong soil water repellency was detected from 0 cm to 3 cm soil depth underneath unburned tree canopies at both woodlands and its strength was not altered by burning. However, fire removal of litter exacerbated repellency effects on infiltration, runoff generation, and erosion. The aggregate stability index method detected differences in relative soil stability between areas underneath trees and in the intercanopy at both sites, but failed to provide any indication of between-site differences in erodibility or the effects of burning on soil erosion potential.  相似文献   

4.
Infiltration was measured in a western juniper (Juniperus occidentalis Hook.) watershed to characterize the hydrologic processes associated with landscape position. Infiltration rate, runoff, and sediment content were measured with the use of a small-plot rainfall simulator. Study sites were located in each of the four primary aspects (north, south, east, and west). Research sites were located in two ecological sites—South Slopes 12–16 PZ and North Slopes 12–16 PZ. Within aspect, plots were located in three juniper cover levels: high (> 22%), moderate (13%–16%), and low (<3%) juniper canopy cover. During rainfall simulation, water was applied at a 10.2-cm · h−1 rate, levels comparable to an infrequent, short-duration, high-intensity precipitation event. Runoff was measured at 5-min intervals for 60 min. Comparing canopy cover levels, steady-state infiltration rates on control plots (9.0 cm · h−1) was 68% greater than high juniper cover sites (2.87 cm · h−1) and 34% greater than moderate juniper cover sites (5.97 cm · h−1) on south-facing slopes. On north-facing slopes, no differences in infiltration rates were observed between juniper cover levels, demonstrating differential hydrologic responses associated with ecological site. Generally, all water applied to control plots infiltrated. Highest infiltration rates were positively associated with increased surface litter and shrub cover. In addition, depth of water within the soil profile was lowest in high juniper cover plots. This suggests that less water is available to sustain understory and intercanopy plant growth in areas with high juniper cover. Accelerated runoff and erosion in juniper dominated sites (high level) across east-, west-, and south- facing slopes can lead to extensive degradation to the hydrology of those sites. These data suggest that sustained hydrologic processes are achieved with reduced western juniper canopy cover.  相似文献   

5.
6.
We investigated soil compaction and hydrologic responses from mechanically shredding Utah juniper (Juniperus ostesperma [Torr.] Little) to control fuels in a sagebrush/bunchgrass plant community (Artemisia nova A. Nelson, Artemisia tridentata Nutt. subsp. wyomingensis Beetle & Young/Pseudoroegneria spicata [Pursh] A. Löve, Poa secunda J. Presl) on a gravelly loam soil with a 15% slope in the Onaqui Mountains of Utah. Rain simulations were applied on 0.5-m2 runoff plots at 64 mm · h?1 (dry run: soil initially dry) and 102 mm · h?1 (wet run: soil initially wet). Runoff and sediment were collected from runoff plots placed in five blocks, each containing four microsites (juniper mound, shrub mound, vegetation-free or bare interspace, and grass interspace) with undisturbed or tracked treatments for each microsite type and a residue-covered treatment for grass and bare interspace microsites. Soil penetration resistance was measured at the hill slope scale, and canopy and ground cover were measured at the hill slope and runoff plot scale. Although shredding trees at a density of 453 trees · ha?1 reduced perennial foliar cover by 20.5%, shredded tree residue covered 40% of the ground surface and reduced non–foliar-covered bare ground and rock by 17%. Tire tracks from the shredding operation covered 15% of the hill slope and increased penetration resistance. For the wet run, infiltration rates of grass interspaces were significantly decreased (39.8 vs. 66.1 mm · h?1) by tire tracks, but infiltration rates on juniper mounds and bare interspaces were unchanged. Bare interspace plots covered with residue had significantly higher infiltration rates (81.9 vs. 26.7 mm · h?1) and lower sediment yields (38.6 vs. 313 g · m?2) than those without residue. Because hydrologic responses to treatments are site- and scale-dependent, determination of shredding effects on other sites and at hill slope or larger scales will best guide management actions.  相似文献   

7.
This study evaluated the efficacy of prescribed fire applied within landscape-scale rotational grazing treatments to reduce mesquite (Prosopis glandulosa Torr.) encroachment and restore herbaceous productivity and cover. One-herd, multiple-paddock rotational grazing was used to accumulate herbaceous fine fuel for fires via prefire deferment and to provide periodic postfire deferment for grass recovery. Treatments were an unburned continuous-grazed control, a four-paddock-1 herd system with fire (4:1F), and an eight-paddock-1 herd system with fire (8:1F), with two replicates per treatment (1 294–2 130 ha per replicate). The management plan was to burn 25% of each system (one paddock in the 4:1F; two paddocks in the 8:1F treatments) and defer grazing during all or portions of the 9 mo (May to January) prior to burning. Deferral was “internalized” by grazing on the remaining 75% of each treatment without reducing stocking rate determined for the entire system. Mesquite cover increased on clay-loam soils from 22% to 40% in unburned paddocks over 7 yr (1995–2001). This increase, coupled with extended drought, reduced fine fuel amounts for fire and limited the number and intensity of fires that were applied. It was possible to burn one paddock in the 8:1F treatment (12.5% of total area), but not in the 4:1F treatment (25% of total area) during drought. Fires reduced mesquite and cactus (Opuntia spp.) cover by 25–79% and 24–56%, respectively, but cover of these species increased to prefire levels within 6 yr. All fires reduced (P ≤ 0.05) total herbaceous biomass for 1 yr postfire. The 8:1F treatment increased (P ≤ 0.05) grass biomass on loamy-bottom soils and reduced (P ≤ 0.05) bare ground on clay-loam and loamy-bottom soils in unburned paddocks compared to the unburned continuously grazed control. The 8:1F treatment, through internalized grazing deferment, facilitated the application of fire to reduce woody cover during extended drought without degrading the herbaceous understory.  相似文献   

8.
Thurber’s needlegrass (Achnatherum thurberianum [Piper] Barkworth) is an important component of many sagebrush communities in the Intermountain West. Prescribed fall burning is often implemented in sagebrush plant communities to mimic historic wildfires, improve wildlife habitat, and increase livestock forage production. Burning is used because it shifts dominance from sagebrush to herbaceous vegetation. The effects of prescribed fall burning on Thurber’s needlegrass are largely unexplored. The purpose of this study was to determine the response of Thurber’s needlegrass to prescribed fall burning. A randomized block design was used, and each block consisted of a fall burned and unburned (control) Wyoming big sagebrush (Artemisia tridentata subsp. wyomingensis [Beetle & A. Young] S. L. Welsh)–bunchgrass communities. Response variables measured in the first and second years after burns were Thurber’s needlegrass community foliar cover and density, vegetative and reproductive biomass, photosynthetic rates, tissue carbon (C) and nitrogen (N) content, and N (15N:14N) and C (13C:12C) isotope ratios. Density of Thurber’s needlegrass in both postburn years and cover in the second postburn year were not different between treatments (P > 0.05), but cover was less in the burned than control treatment in the first postburn year (P = 0.008). Carbon isotope ratios in Thurber’s needlegrass differed between the burn (-25.9 ±  SE) and control (-26.3 ±  SE) treatments in the first postburn year (P = 0.019). Nitrogen isotope ratios indicated nitrogen was more available in the burned than control treatment in both years (P < 0.05). Photosynthetic rates of Thurber’s needlegrass were also generally greater in the burned than control treatment (P = 0.045). Our results suggest burning altered the availability of resources to Thurber’s needlegrass plants. Our results also suggest that prescribed fall burning is not detrimental to Thurber’s needlegrass and, thus, can be used as a method to shift dominance from sagebrush to herbaceous vegetation.  相似文献   

9.
Postfire succession in mountain big sagebrush (Artemisia tridentata Nutt. subsp. vaseyana [Rydb.] Beetle) ecosystems results in a gradual shift from herbaceous dominance to dominance by shrubs. Determining the quality, quantity, and distribution of carbon (C) in rangelands at all stages of succession provides critical baseline data for improving predictions about how C cycling will change at all stages of succession under altered climate conditions. This study quantified the mass and distribution of above- and belowground (to 1.8-m depth) biomass at four successional stages (2, 6, 20, and 39 yr since fire) in Wyoming to estimate rates of C pool accumulation and to quantify changes in ecosystem carbon to nitrogen (C∶N) ratios of the pools during recovery after fire. We hypothesized that biomass C pools would increase over time after fire and that C∶N ratios would vary more between pools than during succession. Aboveground and live coarse roots (CR) biomass increased to 310 and 17 g C · m?2, but live fine roots (FR) mass was static at about 225 g C · m?2. Fine litter (≤ 1-cm diameter) accounted for about 70% of ecosystem C accumulation rate, suggesting that sagebrush leaves decompose slowly and contribute to a substantial soil organic carbon (SOC) pool that did not change during the 40 yr studied. Total ecosystem C (not including SOC) increased 16 g · m?2 · yr?1 over 39 yr to a maximum of 1 100 g · m?2; the fastest accumulation occurred during the first 20 yr. C∶N ratios ranged from 11 for forb leaves to 110 for large sagebrush wood and from 85 for live CR to 12 for bulk soil and were constant across growth stages. These systems may be resilient to grazing after fire because of vigorous regrowth of persistent bunchgrasses and stable pools of live FR and SOC.  相似文献   

10.
Degradation of shrublands around the world from altered fire regimes, overutilization, and anthropogenic disturbance has resulted in a widespread need for shrub restoration. In western North America, reestablishment of mountain big sagebrush (Artemisia tridentata Nutt. ssp. vaseyana [Rydb.] Beetle) is needed to restore ecosystem services and function. Western juniper (Juniperus occidentalis ssp. occidentalis Hook) encroachment is a serious threat to mountain big sagebrush communities in the northern Great Basin and Columbia Plateau. Juniper trees can be controlled with fire; however, sagebrush recovery may be slow, especially if encroachment largely eliminated sagebrush before juniper control. Short-term studies have suggested that seeding mountain big sagebrush after juniper control may accelerate sagebrush recovery. Longer-term information is lacking on how sagebrush recovery progresses and if there are trade-offs with herbaceous vegetation. We compared seeding and not seeding mountain big sagebrush after juniper control (partial cutting followed with burning) in fully developed juniper woodlands (i.e., sagebrush had been largely excluded) at five sites, 7 and 8 yr after seeding. Sagebrush cover averaged ~ 30% in sagebrush seeded plots compared with ~ 1% in unseeded plots 8 yr after seeding, thus suggesting that sagebrush recovery may be slow without seeding after juniper control. Total herbaceous vegetation, perennial grass, and annual forb cover was less where sagebrush was seeded. Thus, there is a trade-off with herbaceous vegetation with seeding sagebrush. Our results suggest that seeding sagebrush after juniper control can accelerate the recovery of sagebrush habitat characteristics, which is important for sagebrush-associated wildlife. We suggest land manager and restoration practitioners consider seeding sagebrush and possibly other shrubs after controlling encroaching trees where residual shrubs are lacking after control.  相似文献   

11.
Western juniper (Juniperus occidentalis spp. occidentalis Hook.) has encroached on and now dominates millions of acres of sagebrush/bunchgrass rangeland in the Great Basin and interior Pacific Northwest. On many sites western juniper has significantly increased exposure of the soil surface by reducing density of understory species and surface litter. We used rainfall and rill simulation techniques to evaluate infiltration, runoff, and erosion on cut and uncut field treatments 10 years after juniper removal. Juniper-dominated hillslopes had significantly lower surface soil cover of herbaceous plants and litter and produced rapid runoff from low-intensity rainfall events of the type that would be expected to occur every 2 years. Direct exposure of the soil to rainfall impacts resulted in high levels of sheet erosion (295 kg · ha-1) in juniper-dominated plots. Large interconnected patches of bare ground concentrated runoff into rills with much higher flow velocity and erosive force resulting in rill erosion rates that were over 15 times higher on juniper-dominated plots. Cutting juniper stimulated herbaceous plant recovery, improved infiltration capacity, and protected the soil surface from even large thunderstorms. Juniper-free plots could only be induced to produce runoff from high-intensity events that would be expected to occur once every 50 years. Runoff events from these higher-intensity simulations produced negligible levels of both sheet and rill erosion. While specific inferences drawn from the current study are limited to juniper-affected sites in the Intermountain sagebrush steppe, the scope of ecosystem impacts are consistent with woody-plant invasion in other ecosystems around the world.  相似文献   

12.
The role of fire in restoration of sagebrush plant communities remains controversial mainly because of paucity of information from long-term studies. Here, we examine 15-year post-fire responses of big sagebrush (Artemisia tridentata ssp wyomingensis) and broom snakeweed (Gutierrezia sarothrae), the two most abundant native shrubs at the John Day Fossil Beds National Monument, a protected area in north-central Oregon, USA. Fire effects were studied along gradients of topography and community type through time post-burn. Community types were distinguished as brush, plots dominated by big sagebrush and woodland, plots with a significant presence of Western juniper (Juniperus occidentalis) trees. Fire reduced big sagebrush cover in brush plots up to 100% and in woodland plots up to 86%. Broom snakeweed cover declined by 92% and 73% in brush plots and woodland plots, respectively. Big sagebrush did not show signs of recovery 15 years after burning regardless of topography and community type while broom snakeweed populations were clearly rebounding and prospering beyond pre-burn levels. Our results showed that an area initially dominated by big sagebrush (cover of big sagebrush 10-20%, cover of broom snakeweed 2-4%) dramatically shifted to an area dominated by broom snakeweed (cover of big sagebrush < 1%, cover of broom snakeweed 5%) in brush-dominated plots. Our results indicated that brush-dominated plots at lower elevation and southern exposures are the least post-fire resilient. We also observed a declining population of big sagebrush on unburned areas, suggesting the lack of post-fire recovery on burned areas was perhaps a result of low seeding potential by extant populations. Although more years of observation are required, these data indicate that recovery time, the encroachment of opportunistic competing shrubs, and the initial condition of vegetation are essential considerations by land managers when prescribing fire in big sagebrush communities.  相似文献   

13.
Runoff and sediment yield from rangelands are extremely important variables that affect productivity, but are difficult to quantify. Studies have been conducted to assess erosion on rangelands, but very little has been done to determine if microchannels (rills) affect runoff and sediment yield. Rainfall simulations were used to quantify the effects of microchannels on runoff and sediment loss on a shortgrass prairie with two types of range conditions (good and fair). Natural flow paths within plots in the two range conditions were defined and then enhanced with an ellipse-shaped hoe to create microchannels. Soil from plots was removed at two rates (11.2 t· ha−1 and 22.4 t· ha−1) to create three soil surface configurations. Soil was removed by vacuuming to create either a single microchannel or multiple microchannels down the plot, and the third treatment was uniform soil removal over the entire plot (sheet). Results showed significantly greater total runoff from both single and multiple microchannel treatments compared with sheet soil removal. The microchannels resulted in significantly less sediment yield per unit of runoff compared with the sheet soil removal treatment. Both runoff and sediment yield were affected by range condition. Plots that were in fair range condition, dominated by buffalo grass (Buchloe dactyloides [Nutt.] Engleman), had a greater amount of total runoff (double) but less sediment yield (75%) than plots in good range condition that were dominated by blue grama (Bouteloua gracilis [H.B.K.] Lag. ex Stued.). The dense buffalo grass sod protected the soil surface from erosion, but water flowed freely across the sod. This study has provided a greater understanding of how microchannels affect runoff and sediment yield under different rangeland conditions, and has illustrated how plant species composition and soil surface features relate to several hydrologic functions.  相似文献   

14.
Much interest lies in long-term recovery rates of sagebrush communities after fire in the western United States, as sagebrush communities comprise millions of hectares of rangelands and are an important wildlife habitat. Little is known about postfire changes in sagebrush canopy cover over time, especially at a landscape scale. We studied postfire recovery of shrub canopy cover in sagebrush-steppe communities with the use of spectral mixture analysis. Our study included 16 different fires that burned between 1937 and 2005 and one unburned site at the US Sheep Experiment Station in eastern Idaho. Spectral mixture analysis was used with September 2006 Systeme Pour l’Observation de la Terre-5 (SPOT-5) satellite imagery to estimate percent shrub canopy cover within pixels. Very large-scale aerial (VLSA) imagery with 24-mm resolution was used for training and validation. SPOT-5 image classification was successful and the spectral mixture analysis estimates of percent shrub canopy cover were highly correlated with the shrub canopy cover estimates in the VLSA imagery (R2 = 0.82; P < 0.0001). Additional accuracy assessment of shrub classification produced 85% overall accuracy, 98% user’s accuracy, and 78% producer’s accuracy. This successful application of spectral mixture analysis has important implications for the monitoring and assessment of sagebrush-steppe communities. With the use of the percent shrub canopy cover estimates from the classified SPOT-5 imagery, we examined shrub canopy recovery rates since different burn years. With the use of linear-plateau regression, it was determined that shrub cover in mountain big sagebrush (Artemisia tridentata Nutt. subsp. vaseyana [Rydb.] Beetle) communities recovered approximately 27 yr after fire, with an average shrub cover of 38%. These results are consistent with other field-based studies in mountain big sagebrush communities.  相似文献   

15.
Prescribed burning of aboveground biomass in tallgrass prairie is common and may influence dynamics and magnitudes of carbon (C) movement between the surface and atmosphere. Carbon dioxide (CO2) fluxes were measured for 2 yr using conditional sampling systems on two adjacent watersheds in an ungrazed tallgrass prairie near Manhattan, Kansas. One watershed was burned annually (BA) and the other biennially (BB). Leaf and soil CO2 fluxes were measured in the source area. Net ecosystem exchange (NEE) of CO2 reached a maximum daily gain of 26.4 g CO2·m?2·d?1 (flux toward surface is positive) in July 1998 (year when both sites were burned and precipitation was above normal); gains were similar between sites in 1998. The maximum daily NEE loss of CO2 was ?21.8 g CO2·m?2·d?1 from BA in September 1997 (year when only BA was burned and precipitation was below normal). When data were integrated over the two years, both sites were net sources of atmospheric CO2; NEE was ?389 g C·m?2·2 yr?1 on BA and ?195 g C·m?2·2 yr?1 on BB. Burning increased canopy size and photosynthesis, but the greater photosynthesis was offset by corresponding increases in respiration (from canopy and soil). Carbon losses from fire represented 6–10% of annual CO2 emissions (bulk came from soil and canopy respiration). Data suggest that annual burning promotes C loss compared to less-frequently burned tallgrass prairie where prairie is not grazed by ungulates. Greater precipitation in 1998 caused large increases in biomass and a more positive growing season NEE, indicating that C sequestration appears more likely when precipitation is high. Because C inputs (photosynthesis) and losses (canopy and soil respiration) were large, small measurement or modeling errors could confound attempts to determine if the ecosystems are long-term CO2 sources or sinks.  相似文献   

16.
Treatments to reduce shrub cover are commonly implemented with the objective of shifting community structure away from shrub dominance and toward shrub and perennial grass codominance. In sagebrush (Artemisia L.) ecosystems, shrub reduction treatments have had variable effects on target shrubs, herbaceous perennials, and non-native annual plants. The factors mediating this variability are not well understood. We used long-term data from Utah’s Watershed Restoration Initiative project to assess short-term (1  4 yr post-treatment) and long-term (5  12 yr post-treatment) responses of sagebrush plant communities to five shrub reduction treatments at 94 sites that span a range of abiotic conditions and sagebrush community types. Treatments were pipe harrow with one or two passes, aerator, and fire with and without postfire seeding. We analyzed effect sizes (log of response ratio) to assess responses of sagebrush, perennial and annual grasses and forbs, and ground cover to treatments. Most treatments successfully reduced sagebrush cover over the short and long term. All treatments increased long-term perennial grass cover in Wyoming big sagebrush (A. tridentata Nutt. ssp. wyomingensis Beetle & Young) communities, but in mountain big sagebrush (ssp. vaseyana [Rydb.] Beetle) communities, perennial grasses increased only when seeded after fire. In both sagebrush communities, treatments generally resulted in short-term, but not long-term, increases in perennial forb cover. Annual grasses (largely invasive cheatgrass, Bromus tectorum L.) increased in all treatments on sites dominated by mountain big sagebrush but stayed constant or decreased on sites dominated by Wyoming big sagebrush. This result was unexpected because sites dominated by Wyoming big sagebrush are typically thought to be less resilient to disturbance and less resistant to invasion than sites dominated by mountain big sagebrush. Together, these results indicate some of the benefits, risks, and contingent outcomes of sagebrush reduction treatments that should be considered carefully in any future decisions about applying such treatments.  相似文献   

17.
Within the sagebrush steppe ecosystem, sagebrush plants influence a number of ecosystem properties, including nutrient distribution, plant species diversity, soil moisture, and temperature, and provide habitat for a wide variety of wildlife species. Recent increases in frequency and size of wildfires and associated annual grass expansion within the Wyoming big sagebrush alliance have increased the need for effective sagebrush restoration tools and protocols. Our objectives were to quay the success of Wyoming big sagebrush transplants relative to transplant stock (nursery seedlings vs. wildlings) across different ecological sites and vegetation types and to test the hypothesis that reduction of herbaceous vegetation would increase survival of transplanted sagebrush. We used a randomized block (reps = 5) design at each of three sites—1) cheatgrass dominated, 2) native plant dominated, and 3) crested wheatgrass dominated—near Elko, Nevada. Treatments included plant stock (nursery stock or locally harvested wildlings) and herbicide (glyphosate) to reduce competition from herbaceous vegetation. Transplants were planted in the spring of 2009 and 2010 and monitored for survival. Data were analyzed for site and treatment effects using mixed-model ANOVA. Surviving plant density at and 2 yr postplanting was generally highest (up to 3-fold) on the native site (P < 0.05). Density of surviving transplants was almost 3-fold higher for nursery stock on most sites for the 2009 planting, but differences in survival by planting stock were minimal for the 2010 planting. Glyphosate application increased surviving plant density up to 300% (depending on site) for both years of planting. High labor and plant material investments (relative to traditional drilling or broadcasting) may limit the size of projects for which sagebrush transplants are practical, but these costs may be partially offset by high success relative to traditional methods. Our data indicate that sagebrush transplants can be effective for establishing sagebrush on depleted sites.  相似文献   

18.
Western juniper (Juniperus occidentalis Hook.) has expanded into sagebrush steppe plant communities the past 130 ? 150 yr in the northern Great Basin. The increase in juniper reduces herbage and browse for livestock and big game. Information on herbaceous yield response to juniper control with fire is limited. We measured herbaceous standing crop and yield by life form in two mountain big sagebrush communities (MTN1, MTN2) and a Wyoming/basin big sagebrush (WYOBAS) community for 6 yrs following prescribed fire treatments to control western juniper. MTN1 and WYOBAS communities were early-successional (phase 1) and MTN2 communities were midsuccessional (phase 2) woodlands before treatment. Prescribed fires killed all juniper and sagebrush in the burn units. Total herbaceous and perennial bunchgrass yields increased 2 to 2.5-fold in burn treatments compared with unburned controls. Total perennial forb yield did not differ between burns and controls in all three plant communities. However, tall perennial forb yield was 1.6- and 2.5-fold greater in the WYOBAS and MTN2 burned sites than controls. Mat-forming perennial forb yields declined by 80 ? 90% after burning compared with controls. Cheatgrass yield increased in burned WYOBAS and MTN2 communities and at the end of the study represented 10% and 22% of total yield, respectively. Annual forbs increased with burning and were mainly composed of native species in MTN1 and MTN2 communities and non-natives in WYOBAS communities. Forage availability for livestock and wild ungulates more than doubled after burning. The additional forage provided on burned areas affords managers greater flexibility to rest and treat additional sagebrush steppe where juniper is expanding, as well as rest or defer critical seasonal habitat for wildlife.  相似文献   

19.
Western juniper (Juniperus occidentalis Hook.) encroachment and exotic annual grass (medusahead [Taeniatherum caput-medusae L. Nevski] and cheatgrass [Bromus tectorum L.]) invasion of sagebrush (Artemisia L.) communities decrease ecosystem services and degrade ecosystem function. Traditionally, these compositional changes were largely confined to separate areas, but more sagebrush communities are now simultaneously being altered by juniper and exotic annual grasses. Few efforts have evaluated attempts to restore these sagebrush communities. The Crooked River National Grassland initiated a project to restore juniper-encroached and annual grass-invaded sagebrush steppe using summer (mid-July) applied prescribed fires and postfire seeding. Treatments were unburned, burned, burned and seeded with a native seed mix, and burned and seeded with an introduced seed mix. Prescribed burning removed all juniper and initially reduced medusahead cover but did not influence cheatgrass cover. Neither the native nor introduced seed mix were successful at increasing large bunchgrass cover, and 6 yr post fire, medusahead cover was greater in burned treatments compared with the unburned treatment. Large bunchgrass cover and biological soil crusts were less in treatments that included burning. Exotic forbs and bulbous bluegrass (Poa bulbosa L.), an exotic grass, were greater in burned treatments compared with the unburned treatment. Sagebrush communities that are both juniper encroached and exotic annual grass invaded will need specific management of both juniper and annual grasses. We suggest that additional treatments, such as pre-emergent herbicide control of annuals and possibly multiple seeding events, are necessary to restore these communities. We recommend an adaptive management approach in which additional treatments are applied on the basis of monitoring data.  相似文献   

20.
Disturbances and their interactions play major roles in sagebrush (Artemisia spp. L.) community dynamics. Although impacts of some disturbances, most notably fire, have been quantified at the landscape level, some have been ignored and rarely are interactions between disturbances evaluated. We developed conceptual state-and-transition models for each of two broad sagebrush groups—a warm-dry group characterized by Wyoming big sagebrush (Artemisia tridentata Nutt. subsp. wyomingensis Beetle & Young) communities and a cool-moist group characterized by mountain big sagebrush (Artemisia tridentata Nutt. subsp. vaseyana [Rydb.] Beetle) communities. We used the Vegetation Dynamics Development Tool to explore how the abundance of community phases and states in each conceptual model might be affected by fire, insect outbreak, drought, snow mold, voles, sudden drops in winter temperatures (freeze-kill), livestock grazing, juniper (Juniperus occidentalis var. occidentalis Hook.) expansion, nonnative annual grasses such as cheatgrass (Bromus tectorum L.), and vegetation treatments. Changes in fuel continuity and loading resulted in average fire rotations of 12 yr in the warm-dry sagebrush group and 81 yr in the cool-moist sagebrush group. Model results in the warm-dry sagebrush group indicated postfire seeding success alone was not sufficient to limit the area of cheatgrass domination. The frequency of episodes of very high utilization by domestic livestock during severe drought was a key influence on community phase abundance in our models. In the cool-moist sagebrush group, model results indicated at least 10% of the juniper expansion area should be treated annually to keep juniper in check. Regardless, juniper seedlings and saplings would remain abundant.  相似文献   

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