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1.
2.

Context

There is strong interest in sustainable forest management systems that preserve characteristics of forests close to naturalness. Assessing the effectiveness of these systems is difficult because defining “natural” baselines from which impacts are estimated is challenging and because the influence of harvesting can have complex interactions with major natural disturbances.

Aims

We used SORTIE/NZ, an individual tree-based forest dynamics model, to understand how harvesting and earthquake disturbance affect the dynamics of a New Zealand podocarp–angiosperm forest.

Methods

Having parameterized SORTIE/NZ with extensive field data, we ran simulations for three natural dynamics scenarios (no disturbance and two earthquake scenarios) and then added podocarp harvesting scenario to each of these.

Results

Simulations suggest that this forest is experiencing transient dynamics, with a natural rise in the dominance of one species of slow-growing podocarp with and without earthquake. Harvesting podocarps strongly affected its increase in basal area.

Conclusion

Our results indicate that transient dynamics may occur in mixed podocarp forests and major disturbances may have complex interactions with management. Evaluating management impacts without accounting for these complex dynamics may be misleading. Models make predictions about transient trajectories that may help to evaluate these impacts.  相似文献   

3.

? Context

Modification of stand density by thinning may buffer the response of tree growth and vigor to changes in climate by enhancing soil water availability.

? Aims

We tested the impact of thinning intensity on cambial growth of Aleppo pine (Pinus halepensis L.) under semi-arid, Mediterranean conditions.

? Methods

A multiple thinning experiment was established on an Aleppo pine plantation in Spain. We analysed the stem growth dynamics of two different crown classes under four different thinning intensities (15 %, 30 %, and 45 % removal of the basal area) for 2 years, based on biweekly band dendrometer recordings. Local relative extractable soil water was derived from the use of a water balance model Biljou© (available at https://appgeodb.nancy.inra.fr/biljou/) and used as an explanatory variable.

? Results

Radial growth was mainly controlled by soil water availability during the growing season, and differed by crown class. The growth rates of dominant trees were significantly higher than the growth rates of suppressed trees. Removal of 30 % and 45 % of the initial basal area produced a growth release in both dominant and suppressed trees that did not occur under less intense thinning treatments.

? Conclusions

Soil water availability was the main driver of radial growth during the growing season. Forest management confirmed its value for ameliorating the effects of water limitations on individual tree growth. These results may help managers understand how altering stand density will differentially affect diameter growth responses of Aleppo pine to short-term climatic fluctuations, promoting forests that are resilient to future climatic conditions.  相似文献   

4.

Context

Reliable estimates of wood density (WD) within individual trees could maximize the value of Pinus massoniana for specific end-use.

Aim

We examined and quantified the axial patterns of WD in trees with different social status in the stands.

Methods

Wood disks were sampled at the bottom, breast height, and middle of each 1-m sections from 108 stems, harvesting from three social classes in subtropical forests. A mixed-effects model was designed to quantify axial variation.

Results

The WD at different height was significantly different from the whole-stem WD (WWD) except the relative height of 0.1. An overall decrease of 133.8 kg m?3 in WD was found from stem base to top. WD was significantly influenced by relative heights, tree age, and social class. WD of each relative height in mature trees was significantly higher than that of younger trees. Tree social class can affect WD development in the axial direction at age classes 2 and 3. Combining the fixed plus random effects, the final model explained 91 % of the observed variation in WD.

Conclusion

The WD development patterns in the axial direction vary considerably among tree age, diameter at breast height, and social class. To distinguish and supply timber for specific end-uses, we should use the axial variation in disk WD (DWD) instead of WWD directly. The accurate predictions of WD provided by the model could be used to optimally classify logs into different product classes and maximize economic benefits. We can use DWD at the relative height of 0.1 instead of WWD of a single tree.  相似文献   

5.

Context

Climate change is expected to increase forest vulnerability through disturbances such as windstorms and droughts. Forest managers are therefore investigating strategies to increase forest resistance and resilience, especially by promoting uneven-aged and mixed forests through group selection, and by reducing stand stocking and large trees proportion. However, there is little information on the long-term impacts of these two practices.

Aims

The objectives of this study were (1) to develop an original silviculture algorithm designed for uneven-aged management and (2) to use it to assess the effects of the above-mentioned management methods in long-term simulations.

Methods

We simulated individual and group selection techniques in order to study the effects of group size, harvesting intensity and their interactions on wood production, stand heterogeneity, and regeneration in mountain spruce–fir forests. We used the spatially explicit individual-based forest model Samsara2 to simulate forest dynamics.

Results

Our simulation results confirmed the positive effect of group selection practices on structure diversity and regeneration but not on spruce maintenance. Increasing harvesting intensity enabled forest destocking but decreased structure diversity and led to non-sustained yields for the most intensive scenarios.

Conclusion

As adaptation measure, we thus recommend moderate group selection harvesting creating 500 m2 gaps.  相似文献   

6.

Context

Natural regeneration with broadleaved species and reforestation with coniferous trees are two widely practiced forest regeneration strategies after timber harvesting. They lead to different tree species composition and may cause different understory biodiversity, but the effects on ground bryophyte composition and diversity are not well-known.

Aims

We tested whether natural regeneration with broadleaved species and reforestation with spruce induced different diversities of the ground bryophyte populations 20–40 years after old-growth spruce forest clearcutting in the subalpine regions of southwestern China.

Methods

Differences between natural stands and plantations were compared through the analysis of 13 paired stands, with 78 plots, 390 shrub/herb quadrats, and a total of 1,560 bryophyte quadrats.

Results

Naturally regenerated forests were characterized by lower density and cover and lower tree height but higher herbaceous plant height, shrub cover, and bryophyte diversity. They also harbored many more ground bryophytes. The species richness of pleurocarpous mosses and fans, mats, and turfs were significantly higher in naturally regenerated forests. Frequency difference analysis demonstrated that more bryophyte species preferred ground habitats in naturally regenerated forests than in plantations (116 vs. 48 species). The canonical correspondence analysis indicated that stand structure attributes were more important determinants of ground bryophyte diversity and abundance.

Conclusion

Natural regeneration and reforestation resulted in large differences in ground bryophyte populations. A larger diversity was observed in the former case, and natural regeneration practices can be an effective measure for the protection of ground bryophyte diversity after clearcutting.  相似文献   

7.

Context

Managing forests under climate change requires adaptation. The adaptive capacity of forest tree populations is huge but not limitless. Integrating evolutionary considerations into adaptive forestry practice will enhance the capacity of managed forests to respond to climate-driven changes.

Aims

Focusing on natural regeneration systems, we propose a general framework that can be used in various and complex local situations by forest managers, in combination with their own expertise, to integrate evolutionary considerations into decision making for the emergence of an evolution-oriented forestry.

Methods

We develop a simple process-based analytical grid, using few processes and parameters, to analyse the impact of forestry practice on the evolution and evolvability of tree populations.

Results

We review qualitative and, whenever possible, quantitative expectations on the intensity of evolutionary drivers in forest trees. Then, we review the effects of actual and potential forestry practice on the evolutionary processes. We illustrate the complexity of interactions in two study cases: the evolutionary consequences for forest trees of biotic interactions and of highly heterogeneous environment.

Conclusion

Evolution-oriented forestry may contribute adapting forests to climate change. It requires combining short-term and long-term objectives. We propose future lines of research and experimentation.  相似文献   

8.

? Context

The rising demand of energy wood for heating purposes in Germany leads to concerns regarding the overexploitation of forests. A major aspect is the impact of whole-tree harvesting on long-term productivity of forest soils.

? Aims

This study aimed to analyze the effects of nutrient removal on productivity using the historically prevalent practice of litter raking. Since there is a lack of controlled whole-tree harvesting experiments in Germany, we used litter raking as a surrogate management practice entailing the removal of nutrients from forest stands.

? Methods

We used three sites with documented litter raking to analyze the effects of nutrient removal on productivity using dendroecological methods: two recent litter removal experiments in two Scots pine stands (Siegenburg and Burglengenfeld) and one oak stand (Eichhall) with documented historic litter raking. Basal area increment (BAI) and tree-ring characteristics were compared between periods with litter raking and the preceding periods for both treatment and control plots.

? Results

For the two Scots pine sites with a relatively short litter raking period, no effects of litter raking on BAI could be ascribed to nutrient removal. On the oak site with a longer history of litter utilization, the loss in BAI due to litter raking amounts to 22 % during the period with active raking and to still 17 % in the recovery period.

? Conclusions

These results contribute to the still very limited understanding about the impact of whole-tree harvesting on forest productivity in Germany by laying down an upper limit of possible effects due to nutrient removal, as nutrient loss by litter raking tends to be higher than nutrient loss by whole-tree harvesting.  相似文献   

9.

Key message

Radial growth of silver and red maples was investigated across three forests in northwest Ohio following the outbreak of the invasive emerald ash borer. The growth response of maples was driven by an advancement in canopy class and disturbance severity.

Context

Forest disturbances resulting in species-specific diffuse mortality cause shifts in aboveground and belowground competition. This competition may differentially affect non-impacted trees, depending on crown class, disturbance severity, and species-specific responses.

Aims

The purpose of this study is to elucidate the primary drivers of silver and red maple (Acer saccharinum and A. rubrum) growth following emerald ash borer (EAB, Agrilus planipennis)-induced ash tree (Fraxinus spp.) mortality in riparian forests of northwest Ohio.

Methods

Using dendroecological approaches, we analyzed the pattern of radial growth in red and silver maples in conjunction with the EAB outbreak.

Results

This study revealed growth rates of maples increased 72% following EAB arrival and trees advancing in crown class grew 41% faster than those not advancing. The growth response varied by initial crown class, with trees in the intermediate class responding most dramatically. Furthermore, the positive correlation between relative basal area of ash and the radial growth response of maples indicates the important role of disturbance severity in post-disturbance dynamics.

Conclusion

These findings suggest that, although advancement in crown class may allow predictions of “winners” in forest succession post-disturbance, even trees not changing crown class benefit from decreased competition. Results from this study provide a detailed account of radial growth responses in maples following EAB-induced ash mortality and lend insight into the future canopy composition of ash-dominated riparian forests.
  相似文献   

10.

Context

Old-growth trees play a very important role in the maintenance of biodiversity in forests. However, no clear definition is yet available to help identify them since tree age is usually not recorded in National Forest Inventories.

Aims

To develop and test a new method to identify old-growth trees using a species-specific threshold for tree diameter in National Forest Inventories.

Methods

Different nonlinear mixed models for diameter–age were generated using data from the Spanish Forest Inventory in order to identify the most appropriate one for Aleppo pine in its South-western distribution area. The asymptote of the optimal model indicates the threshold diameter for defining an old-growth tree. Additionally, five site index curves were examined to analyze the influence of site quality on these models.

Results

The Hossfled III mixed model was found to be the most appropriate to fit diameter–age curves for Aleppo pine trees. The overall diameter at breast height threshold for old-growth trees was 40.6 cm, although over a range of sites with increasing site quality, the threshold figure was 36.0, 38.0, 40.4, 43.1, and 46.3 cm, respectively.

Conclusions

This method allows the identification of old-growth trees and therefore of biodiversity hotspots, thus providing decision makers with a useful tool for management purposes.  相似文献   

11.

? Context

The knowledge of how shrub–seedling interactions vary with summer drought, canopy opening, and tree species is crucial for adapting forest management to climate change.

? Aims

The aim of this study was to assess variation in shrub–oak recruitment associations along a south–north drought climate gradient and between two levels of canopy cover in coastal dune forest communities in a climate change-adapted forest management perspective.

? Material and methods

Mapped data of associational patterns of seedlings of three oak species with interspecific pooled shrubs were analyzed using a bivariate pair correlation function in 10 (0.315 ha) regeneration plots located in forest and recent gap sites along the climate gradient. An index of association strength was calculated in each plot and plotted against a summer moisture index.

? Results

The association strength increased with increasing summer drought from wet south to dry north and from closed forests to gaps.

? Conclusion

Consistent with facilitation theory, our results suggest that climate change may shift associational patterns in coastal dune forest communities towards more positive associations, in particular in canopy gaps. In a perspective of climate change, foresters may need to conserve understory shrubs in gaps in order to promote oak species regeneration.  相似文献   

12.

? Context

Biomass expansion factors (BEFs, defined as the ratios of tree component biomass (branch, leaf, aboveground section, root, and whole) to stem biomass) are important parameters for quantifying forest biomass and carbon stock. However, little information is available about possible causes of the variability in BEFs at large scales.

? Aims

We examined whether and how BEFs vary with forest types, climate (mean annual temperature, MAT; mean annual precipitation, MAP), and stand development (stand age and size) at the national scale for China.

? Method

Using our compiled biomass dataset, we calculated values for BEFs and explored their relationships to forest types, climate, and stand development.

? Results

BEFs varied greatly across forest types and functional groups. They were significantly related to climate and stand development (especially tree height). However, the relationships between BEFs and MAT and MAP were generally different in deciduous forests and evergreen forests, and BEF–climate relationships were weaker in deciduous forests than in evergreen forests and pine forests.

? Conclusion

To reduce uncertainties induced by BEFs in estimates of forest biomass and carbon stock, values for BEFs should be applied for a specified forest, and BEF functions with influencing factors (e.g., tree height and climate) should be developed as predictor variables for the specified forest.  相似文献   

13.

??Context

It is assumed that climate change will favour European beech (Fagus sylvatica L.) to Norway spruce (Picea abies [L.] Karst.) at its northern range margins due to climate change and induced disturbance events.

??Aims

An old-growth mixed forest of spruce and beech, situated near the northern beech margin, was studied to reveal effects of disturbances and response processes on natural forest dynamics, focussing on the understory.

??Methods

We carried out analyses on understory dynamics of beech and spruce in relation to overstory release. This was done based on a sequence of stand and tree vitality inventories after a series of abiotic and biotic disturbances.

??Results

It became apparent that beech (understory) has a larger adaptive capacity to disturbance impacts and overstory release (68 % standing volume loss) than spruce. Understory dynamics can play a key role for forest succession from spruce to beech-dominated forests. Disturbances display an acceleration effect on forest succession in the face of climate change.

??Conclusion

Beech is poised strategically to replace spruce as the dominant tree species at the study area. Due to an increasing productivity and a lower risk of stand failure, beech may raise into the focus of forestry in southern Sweden.  相似文献   

14.

?Context

Selective logging followed by natural regeneration is rarely employed for restocking subtropical evergreen broad-leaved forests in East Asia compared with the use of clear-cutting.

?Aims

To clarify the succession of these forests, the effects of selective logging on stand structure, species diversity, and community similarity were studied in a mature and regenerating forest in Okinawa, Japan.

?Methods

Four study plots were established, and trees ≥1.2 m height were identified by species name, tree height, and diameter at breast height.

?Results

The results showed that the species composition of regenerating forest was similar to mature forest; however, the former had a greater species density and Shannon–Wiener index than the latter. Castanopsis sieboldii and Distylium racemosum, the predominant trees in the mature forest, continued to dominate the regenerating forest, with a broad layer distribution. High Sørensen and Jaccard community similarity indices for mature and regenerating forest indicated that the regeneration occurred in a progressive succession.

?Conclusion

The similar species composition and stand structure for both mature and regenerating forest, and the higher species diversity for the latter, provided no evidence of forest degeneration and suggested that the regenerating forest may develop into a stand similar to preselective logging forest.  相似文献   

15.

? Context

Snow gliding is a downhill motion of snow on the ground; observations have shown gliding to be possible not only on open slopes but also in forest stands. Larch stands, with their low canopy density and open forest structure with clearings and gaps, are particularly prone to high glide rates. Snow gliding may have negative effects on juvenescent trees which can be damaged by extraction from the ground.

? Aim

The goal of this study was to determine whether snow gliding depends on forest cover (canopy) and size of clearings.

? Methods

Snow gliding was measured during eight winter periods at six measuring positions (ranging from ‘dense forest’ to ‘open slope’) in and beside a larch stand in the Stubai Valley, Tyrol, Austria.

? Results

The results showed that gliding is strongly influenced by forest cover. Snow gliding increases with decreasing canopy density. The difference between the six measuring positions was highly significant (p?<?0.005).

? Conclusion

The identified glide cracks on at least two measuring positions, indicating extreme glide rates and, therefore, strong negative effects on juvenescent trees. To prevent glide rates of a magnitude such as this requires a mature forest with at least 300 stems/ha.  相似文献   

16.

Context

After wildfire, surviving trees are of major ecological importance as they can help in the post-fire regeneration process. Although these trees may be damaged, they may also benefit from reduced fuel hazard and competition. However, little is known about the long-term growth response of surviving trees.

Aims

This study aims to explain short- to long-term variations in the postfire growth of surviving black pines in an area burnt in 1994, focusing on levels of fire severity and tree sizes.

Methods

Relative basal area increments were used to detect time-course variations in postfire radial tree growth depending on fire severity. Linear mixed-effects models were used to describe the factors affecting postfire ring growth.

Results

In the short term, fire caused stronger reduction in growth in small trees with increasing bole char height. However, as time since fire increased, a positive effect of fire on growth due to reduced competition counteracted the short-term fire impacts. Indeed, small surviving trees demonstrated a surge in growth 15 years after the fire.

Conclusion

It was concluded that reduced competition might offset the short-term negative effects of fire in surviving black pines.  相似文献   

17.

Context

The current fire regime threatens black pine (Pinus nigra Arn.) persistence in the Mediterranean Basin, which recommends larger-scale fuel treatments. Prescribed burning is an option for stand protection but its use in young stands (which are particularly at risk) is hindered by the scarce knowledge on post-fire tree survival.

Aims

The objectives were to characterize bark thickness as a fire-resistance trait in P. nigra and to describe how post-fire tree survival responds to tree size and fire effects in a 16-year-old plantation.

Methods

Bark thickness was related to diameter at breast height and height in the stem. Metrics describing tree size and stem and crown damage were measured 1?year after prescribed burning in 259 trees. Tree survival was modeled with logistic regression and Classification and Regression Tree analysis.

Results

Bark thickness increased linearly with diameter at breast height (dbh) and decreased with height in the stem. Tree survival was primarily a function of crown injury. Stem damage was an influent factor in small trees.

Conclusion

Due to thinner bark and lower tolerance to crown damage, young P. nigra trees are less fire-resistant than other Mediterranean pines, e.g., Pinus pinaster. Prescribed fire should not be attempted if dbh <10?cm. Mechanical clearing is the treatment of choice in young stands with a significant shrub layer.  相似文献   

18.

?Context

Understanding tree interactions requires an insight into their spatial distribution.

?Aims

We looked for presence and extent of tree intraspecific spatial point pattern (random, aggregated, or overdispersed) and interspecific spatial point pattern (independent, aggregated, or segregated).

?Methods

We established twelve 0.64-ha plots in natural bottomland hardwood stands in the southeastern USA.

?Results

Spatial point pattern analyses (Ripley’s K, L, and L 12) indicated that, when species were combined, trees were frequently aggregated and less commonly overdispersed. Plots with larger trees were more likely to exhibit overdispersion, confirming a shift to this pattern as trees grow. The intraspecific pattern of cherrybark oak and water oak was either aggregated or random. Sweetgum was aggregated on all plots and always at smaller distances (less than 5 m) than the two oak species. Intraspecific overdispersion was very rare. Interspecific segregation among the two oak species was more commonly observed (six plots) than aggregation (one plot). Cherrybark oak and sweetgum were segregated at some scale on seven of the 12 plots and aggregated on only two plots.

?Conclusion

The results from the analyses suggest that strong interspecific competition may result in segregation of trees from different species, while weaker intraspecific competition may lead to aggregations of conspecifics.  相似文献   

19.

Context

Edible stone pine (Pinus pinea L.) nut is a forest product which provides the highest incomes to the owners of stone pine forests.

Aim

The objective of this work is to evaluate the effect of first thinning on growth and cone production in an artificially regenerated stand in order to determine optimum intensity.

Methods

A thinning trial was installed in 2004 to compare two thinning regimes (heavy and moderate) and a control treatment. From 2004 to 2012, six inventories of forest attributes were carried out, and the cone crop was harvested annually. We evaluated the effect of thinnings on growth using repeated measures analysis of variance with a mixed model approach. With regards to cone production, we first estimated the probability of finding cones in a tree by applying a generalized mixed model and then estimated cone production by using a mixed model, including climatic variables.

Results

We found that thinning had a positive influence on tree diameter increment. Thinning increased the probability of finding cones and cone production. However, significant differences between heavy and moderate thinnings were not found.

Conclusion

We recommend early silvicultural treatments in stone pine stands to favor the development of trees and larger edible pine nut production.  相似文献   

20.

? Context

Copaifera species produce an oleoresin of commercial importance that is widely extracted in Amazon communities.

? Aims

This paper addresses two questions. (1) What are the morphological characteristics of Copaifera species that influence oleoresin production? (2) How do different Copaifera species respond to repeated harvests?

? Methods

We performed a large-scale experiment in the Brazilian Amazon. We tapped 110 Copaifera trees belonging to four species, and several morphological tree characteristics were measured to determine their effect on oleoresin production.

? Results

The proportion of Copaifera reticulata and Copaifera paupera trees that yielded more than 1 ml oleoresin was higher than the other species. The oleoresin volumes from yielding Copaifera pubiflora trees were significantly higher than those from C. reticulata and C. paupera, with Copaifera multijuga yielding intermediate values. Interestingly, none of the studied morphological tree characteristics had a significant effect on the proportion of yielding trees. Hollowed trees yielded significantly smaller volumes than non-hollowed trees. Both the proportion of yielding trees and oleoresin volumes decreased from the first to the second harvests for C. reticulata and C. paupera; however, the opposite pattern was observed for C. pubiflora.

? Conclusions

Oleoresin production capacity varies by species, and management protocols should account for these differences.  相似文献   

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