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1.
The aim of this study was to determine how soil chemistry and the distribution of fine roots (<1 mm) in the organic and upper mineral soil horizons were affected by an admixture of birch ( Betula pendula Roth and B. pubescens Ehrh.) in Norway spruce ( Picea abies (L.) Karst) stands. The surface organic horizons (LF and H) and mineral soil were characterized to a depth of 10 cm on three sites in southern and central Sweden. On these sites, replicated plots had been established that contained either ca. 30-year-old birch growing as a shelter over similar-aged spruce (mixed plots) or spruce only. The treatments had been created 8–11 years before this study was done. A fourth site, with plots containing ca. 90-year-old spruce or birch/spruce, and a fifth site, with 30-year-old spruce and a low admixture (12% by basal area) of birch, were also included in the study. Concentrations of Ca and Mg and pH in the LF layer were significantly higher in plots with a birch admixture. In the H-horizon, concentrations of K, Ca and Mg were significantly higher in mixed plots than in plots with pure spruce. Consequently, base saturation was higher in mixed plots than in pure spruce plots. A shelter of birch decreased the total amount of spruce fine roots (<1 mm), as revealed at one of the sites. Total fine root biomass (birch + spruce) in the organic and mineral soil horizons (to 10 cm) did not differ significantly between the pure spruce stands and the spruce stands with a birch shelter. 相似文献
2.
Stand structure, height and diameter growth, above- and below-ground biomass, and nutrient concentrations and content were determined for a 35-year-old fire-origin paper birch ( Betula papyrifera) and subalpine fir ( Abies lasiocarpa) mixed-species stand in the Sub-Boreal Spruce (SBS) zone of British Columbia. Paper birch, which formed the dominant overstory following the 1961 fire, had normal distributions of height and diameter classes. Subalpine fir, which dominated the understory, had the reverse J-shaped height and diameter distributions that are expected of a shade tolerant, climax species. Paper birch grew more than three times the height of subalpine fir. Growing in the summer shade of the birch, subalpine fir had slow but steady height growth during the first 10–15 years, after which height growth declined somewhat. Allometric equations, relating dry weight of foliage, branches, stemwood, stembark, roots, and total biomass to diameter at breast height (DBH), were developed to estimate above- and below-ground biomass. Total biomass of paper birch reached 83.2 t ha −1, while subalpine fir biomass was 26.7 t ha −1. Subalpine fir allocated more biomass to foliage and branches compared to paper birch. Foliage of paper birch had higher nutrient concentrations of N, P, K, Ca, and Mg than subalpine fir foliage. Branches and stembark of subalpine fir had higher P, and Ca concentrations than paper birch. Subalpine fir branches contained more of all examined nutrients than paper birch branches. This is a significant component in nutrient cycling of the mixed-species forest. 相似文献
3.
For reasons of aesthetics, sheltering, biodiversity, and children’s need of playgrounds, selective coppicing could be a useful and effective silvicultural method in urban areas. A randomised block experiment was carried out in three broad-leaved coppices situated within the fenced area of Arlanda Airport to compare a selective coppice regime (SCR), where all trees above a certain height limit were cut, with a traditional coppice regime, where all trees were felled. The effects of the treatments of the dominating tree species European aspen ( Populus tremula L.) and birch ( Betula pendula Roth and B. pubescens Ehrh.) were studied. During three growing seasons after performed treatment, the proportion of sprouting stools and root shooting of European aspen were reduced in SCR. Birch stools had fewer sprouts stool −1, and sprouts of both birch and European aspen tended to be shorter in SCR. There was a tendency that the sprout mortality was lower in SCR for both birch and European aspen. Residual stands of SCR consisted of proportionally less birch, and more rowan, bird cherry, oak and other species. Coppicing with single tree selection from above suppressed light demanding tree species and promoted the more shade-tolerant species compared to traditional coppicing. 相似文献
4.
Carbon (C) sequestration was studied in managed boreal forest stands and in wood products under current and changing climate in Finland. The C flows were simulated with a gap-type forest model interfaced with a wood product model. Sites in the simulations represented medium fertile southern and northern Finland sites, and stands were pure Scots pine and Norway spruce stands or mixtures of silver and pubescent birch. Changing climate increased C sequestration clearly in northern Finland, but in southern Finland sequestration even decreased. Temperature is currently the major factor limiting tree growth in northern Finland. In southern Finland, the total average C balance over the 150 year period increased slightly in Scots pine stands and wood products, from 0.78 Mg C ha−1 per year to 0.84 Mg C ha−1 per year, while in birch stands and wood products the increase was larger, from 0.64 Mg C ha−1 per year to 0.92 Mg C ha−1 per year. In Norway spruce stands and wood products, the total average balance decreased substantially, from 0.96 Mg C ha−1 per year to 0.32 Mg C ha−1 per year. In northern Finland, the total average C balance of the 150 year period increased under changing climate, regardless of tree species: in Scots pine stands and wood products from 1.10 Mg C ha−1 per year to 1.42 Mg C ha−1 per year, in Norway spruce stands and wood products from 0.69 Mg C ha−1 per year to 0.99 Mg C ha−1 per year, and in birch stands and wood products from 0.43 Mg C ha−1 per year to 0.60 Mg C ha−1 per year. C sequestration in unmanaged stands was larger than in managed systems, regardless of climate. However, wood products should be included in C sequestration assessments since 12–55% of the total 45–214 Mg C ha−1 after 150 years' simulation was in products, depending on tree species, climate and location. The largest C flow from managed system back into the atmosphere was from litter, 36–47% of the total flow, from vegetation 22–32%, from soil organic matter 25–30%. Emissions from the production process and burning of discarded products were 1–6% of the total flow, and emissions from landfills less than 1%. 相似文献
5.
The aim of this study was to quantify 5-year growth, yield and mortality responses of 9- to 13-year-old naturally regenerated, even-aged paper birch ( Betula papyrifera Marsh.) stands to pre-commercial thinning in interior British Columbia. The study included four residual densities (9902–21,807 stems ha −1 (unthinned control), 3000, 1000 and 400 stems ha −1) and four sites with 3-fold within-site replication in a randomised block design. The largest, straightest, undamaged trees were selected to leave during thinning. Thinning reduced stand basal area from 5.90 m 2 ha −1 in the control to 2.50, 1.53 and 0.85 m 2 ha −1 in the three thinning treatments, representing 42, 26 and 15% of control basal area, respectively. After 5 years, total stand volume per plot remained lower in the three thinning treatments than the control (50.20, 30.07, 18.99 and 11.86 m 3 in the control, 3000, 1000 and 400 stems ha −1 treatments), whereas mean stand diameter, diameter increment, height, and height increment were increased by thinning, and top height (tallest 100 trees ha −1) was unaffected. When a select group of crop trees (largest 250 trees ha −1) in the thinning treatments was compared with the equivalent group in the control, there was a significant increase in mean diameter, diameter increment, basal area, basal area increment, and volume increment. Mean height, height increment, top height, and total volume were unaffected by thinning. Crop tree diameter increment was the greatest following thinning to 400 stems ha −1 for all diameter classes. Thinning to 1000 stems ha −1 resulted in lower diameter increment than thinning to 400 stems ha −1 but tended to have higher volume increment. Dominant trees responded similarly to subdominant trees at 400 stems ha −1, but showed the greatest response at 3000 stems ha −1. Results suggest that pre-commercial thinning of 9–13-year-old stands to 1000 stems ha −1 would improve growth of individual trees without seriously under-utilising site resources. 相似文献
6.
Both incoming shortwave radiation ( Rg) and photosynthetically active radiation (PAR) in percentage of full daylight were measured at the same time by point and strip sampling in four plots (0.1 ha) of Picea abies (L.) Karst. The standard deviations (%) of Rg and PAR were, respectively, 11.1 and 9.8 at 64 points, 15.7 and 13.9 at 32 points, and 24.7 and 23.8 at 16 points per plot. A period of at least 40 s per strip (30 m min−1) gives a CV (coefficient of variation) of 30%. There is no significant difference between relative irradiance (RI) estimated by the point method (64 points) and by the strip method (8 strips). Curves of RI (Rg and PAR) and basal area (m2 ha−1), diameter sum (m ha−1) and density (stems ha−1) of fifteen trials with different thinning programmes are presented. Irradiance (Rg) in heavily thinned stands was 3–14% of irradiance on an open place. The irradiance, Rg, in extra-heavily thinned stands is 12–27%, and in unthinned stands, 1–3% that of an open place. The Rg curve lies above the PAR curve in all cases. Some practical implications of the study are presented. Heavy thinning of Norway spruce stands gives RI (Rg) values 10% at basal area of 25m2 ha−1 which is necessary to minimize development of suckers of broadleaved trees. 相似文献
7.
Two thinning and fertilization studies, the first in 1969 and the second in 1971, were established to evaluate the question of nutrient limitation to tree growth and the consequences of stand manipulation of soil moisture supply. Fertilizer was applied yearly for the first 5 years in both studies; growth response has been measured through 1987. Results indicate that thinning is necessary to obtain a growth response to fertilizer applied at the rate of 111 kg nitrogen ha −1. The response to fertilization after fertilization ended lasted for 4 years in plots thinned to 800 stems ha −1, while a significant response continued for only 2 years in plots thinned to 1600 stems ha −1. A soil water-balance model was calibrated for the control and treatment plots of these two studies. Soil water-deficits were estimated and correlated with yearly average basal-area growth per tree. Results indicated that there is a correlation between seasonal soil-moisture deficit and growth during the years when soil moisture was measured for the unthinned control plots (r2 = −0.787, P = 0.002) but not for the thinned and fertilized plots (r2 = −0.652, P = 0.057). 相似文献
8.
Pre-commercial thinning (PCT) is a silvicultural practice that can provide diverse understory and overstory vegetation conditions. We tested the hypothesis that relative habitat use by snowshoe hare ( Lepus americanus), mule deer ( Odocoileus hemionus), and moose ( Alces alces) would increase in response to enhanced abundance of herbs and shrubs, and species diversity and structural diversity of conifers, in heavily thinned (≤1000 stems/ha) stands, at 12–15 years post-thinning. Replicate study areas were located near Penticton, Kamloops, and Prince George in south-central British Columbia, Canada. Each study area had three young pine stands thinned to densities of 500 stems/ha (low), 1000 stems/ha (medium), and 2000 stems/ha (high), with an unthinned young pine and old-growth pine stand for comparison. Relative habitat use, based on counts of fecal pellets and pellet-groups, was similar among the five treatment stands for hares (P = 0.24), deer (P = 0.23), and moose (P = 0.16). However, low-density stands (500 stems/ha) had ca. 3–20 times as many deer pellet-groups, and ca. 2–4 times as many moose pellet-groups, than other stands. Low-density stands had significantly greater canopy openness, volume of shrubs <2 m, and horizontal hiding cover <1.6 m than other treatments. Relative habitat use by deer and moose was positively related to understory characteristics such as enhanced abundance of forage and security cover. These results support our hypothesis that deer and moose responded positively to enhanced volume of herbs and shrubs as well as to species diversity and structural diversity of conifers and overall vegetation in heavily thinned (≤1000 stems/ha) stands at 12–15 years post-thinning. Our results suggest that ungulate management would be enhanced if greater emphasis was placed on forage enhancement throughout the year, which differs from current management recommendations which tend to focus on winter range and snow-interception cover. 相似文献
9.
In 1989, the first recorded outbreak of hemlock looper ( Lambdina fiscellaria fiscellaria (Guen.)) occurred in New Brunswick, Canada. Data were collected from ten plots established in an area infested from 1992–1994, to assess impacts of hemlock looper. Ocular and branch sample assessments of current defoliation and ocular assessments of total defoliation (all age classes of foliage) were conducted for balsam fir ( Abies balsamea [L.] Mill.), white spruce ( Picea glauca [Moench] Voss), and black spruce ( Picea mariana (Mill.) B.S.P.). Stand response was assessed and related to cumulative defoliation. Ocular assessments were found to accurately estimate defoliation, which was significantly related to tree mortality. Ninety-two percent of balsam fir trees that had cumulative defoliation >90% died. Mortality of balsam fir was significantly ( p < 0.05) related to tree size, in both lightly and severely defoliated plots; trees with DBH <11 cm sustained 22–48% higher mortality than larger trees. Mortality of balsam fir, in terms of both percent stems/ha and m 3/ha merchantable volume, increased exponentially in relation to three estimates of cumulative (summed) plot mean defoliation. The strongest relationships ( r2 = 0.75–0.79) were between mortality and the ocular defoliation assessment for 1990–1993 foliage. Tree mortality caused by the looper outbreak ranged from 4–14% stems/ha in lightly defoliated and from 32–100% in severely defoliated plots; merchantable volume killed was 3–14 m 3/ha and 51–119 m 3/ha, respectively. Relationships between mortality and defoliation were similar when defoliation was assessed for 1987–1993 and 1990–1993 foliage age classes. 相似文献
10.
Tree vegetation and size structure was sampled in a miombo woodland area in E Tanzania and related to environmental factors, particularly soil and disturbance history. A total of 86 tree species was found. Four plant communities were distinguished through multivariate classification. Community 1 was dominated by Brachystegia boehmii, Brachystegia bussei and Julbernardia globiflora, and community 2 by B. boehmii and Brachystegia spiciformis. Community 1 was found on grey, eroded soil and community 2 on red, residual soil, a fact that opens up possibilities to use soil signals of satellite data for vegetation mapping. Community 3 is heavily disturbed miombo woodland near villages and community 4 was found on more clayey soil where miombo woodland is not expected. At our 42 sampled sites, density ranged from 74 to 1041 individuals ha−1 and basal area from 3.9 to 16.7 m2 ha−1. Regeneration is generally good but large sized trees are less prominent in communities 3 and 4 due to harvesting. With reduced disturbance miombo species may rapidly resume dominance in community 3. A higher than expected representation by the size class 30–40 (−50) cm dbh in community 2 is probably related to disturbance history. Prevalence of certain species (Pseudolachnostylis maprouneifolia, Pterocarpus angolensis and Diplorhynchos condylocarpon) may be related to frequent fires. Selective logging will soon cause extinction of Dalbergia melanoxylon, whereas Pterocarpus angolensis still has good regeneration, possibly because individuals below logging size have a good seed set. A way to get an easy overview of size classes in all species in an area using PCA is discussed. 相似文献
11.
For 20 years, there has been 42,000 ha estate of clonal Eucalyptus plantations around Pointe-Noire in Congo on sandy soils that have very low reserves of available nutrients. These plantations have been based on a natural hybrid ( E. PF1). This hybrid is being replaced by E. urophylla × E. grandis (UG), a more productive hybrid developed by the breeding program of UR2PI. A study of biogeochemical cycles showed that nutrient removal by harvesting is the main nutrient output in the E. PF1 ecosystem. It is therefore important to quantify the nutrient content (NC) in both hybrids to compare corresponding nutrient removal values. The work dealt with four UG clones and the most planted clone of E. PF1. Twelve trees per clone were sampled at the logging age (8 years) in a clonal test for UG clones and in a nearby stand for E. PF1. Tables were established to predict, from girth at breast height (C1.30 m), the biomass and nutrient content of stemwood, bark, dead and living branches, leaves, and were applied to the inventory of the different stands to evaluate corresponding biomass, NC and nutrient use efficiency (NUE) on a per-hectare basis. Total biomass differed between the two hybrids and among UG clones: 109 t ha−1 for E. PF1 and 108–155 t ha−1 for UG clones. In E. PF1 trees, total NC was globally lower for N, K, and Mg, but greater for P and Ca. In stemwood, nitrogen content was similar for both hybrids. By contrast, in UG clones, NC was much lower for P (−72%) and Ca (−40% to −55%). The same trends were observed for NUE: equivalent for both hybrids for N, but higher in UG clones for P (+72%) and Ca (+43% to +59%). A marked variability among clones was observed for K and Mg. UG clones allocated proportionally more nutrients in leaves than E. PF1. These results show that clones should not be selected only on growth traits but also on NUE and on the concentration of nutrients in tree components removed by harvesting. It will be then possible to limit the cost of fertilising needed to maintain stand growth and soil fertility. 相似文献
12.
The influence of above-ground insect herbivory and other agents of damage to seedlings was studied in a field experiment. Nine different tree species were planted in each of five randomised blocks. The species were: Alnus glutinosa (L.) Gaertner, Betula pendula Roth., Fagus sylvatica L., Fraxinus excelsior L., Larix eurolepis Henry, Picea abies (L.) Karst., Prunus avium L., Quercus robur L. and Tilia cordata Mill. The seedlings were monitored through the 2000–2002 growing seasons. Pine weevils ( Hylobius abietis L.) fed significantly more on Norway spruce seedlings than on all other tree species. The following rough preference order was obtained: Norway sprucelarch>birch, beech, oak>cherry>alder, ash, lime. Thus, during the conversion of single-species coniferous plantations to mixed-species broadleaved stands, the use of costly insecticides against pine weevil is not required. Short-snouted weevils (e.g. Strophosoma melanogrammum Forst. and Otiorhynchus scaber L.) and other insects caused leaf and needle area loss primarily during the first growing season. 相似文献
13.
Early management of the regenerated seedlings shapes the future stand properties. To address these issues, pre-commercial thinning (PCT) and control treatments were applied to planted Norway spruce ( Picea abies L. Karst) and naturally regenerated birch ( Betula pendula Roth., Betula pubescens Ehrh.) stands in forest experiments in southern Sweden (lat. 56–57?N) containing 1.1–5.5?m tall saplings. The treatments were retention of 1000 or 2000?stems?ha ?1 of Norway spruce, with no birch or birch at 1000?stems?ha ?1. Treatments were replicated with and without annual removal of birch sprouts from stumps. The periodic annual increment (PAI) over five years was calculated for total stand volume and individual trees. The mean PAI of dominant trees was significantly higher both following all PCT treatments than controls, and following low rather than high-density PCT. Birch retention did not affect growth of the dominant trees but PAI was lower in plots with uncontrolled sprouting. The PAI of birch was significantly higher in low-density Norway spruce plots than in control plots and the high-density plots. The treatment response was significant even in stands with initial heights of only 1–2?m. 相似文献
15.
Annual net primary production (NPP) and N uptake were estimated for lysimeter-grown basket willows ( Salix viminalis L.) during 3 years after planting. The willows were grown in a stand structure and continuously supplied with water and liquid fertilizer through drip tubes. The lysimeters contained either clay from the site or washed quartz sand. Shoot growth and leaf litter were measured and fine-root dynamics observed in minirhizotrons. Destructive samples were taken annually in late autumn and entire root systems were washed out. Dry mass and N content of all plant parts were determined. Fine-root production was estimated by two methods, based on destructive samplings and observations in minirhizotrons. The proportion of biomass allocated below ground increased considerably when estimates based on accumulated NPP were compared with those based on standing dry mass. In the first year, 49 and 58% of annual NPP in willows grown in clay and sand, respectively, was belowground. In subsequent years the proportions were 36–38% and 33–40%. Most belowground production was fine roots. Relatively more N was used belowground in the first year than subsequently, but no substrate-induced differences were observed in the allocation pattern. Both annual NPP and N uptake was always higher in plants in clay than in those in sand: in the final 2 years, 21–22 tonnes DM ha−1 year−1 and 190 kg N ha−1 year−1 in clay, and 9–10 tonnes DM ha−1 year−1 and 100 kg N ha−1 year−1 in sand. 相似文献
16.
Three stand types on drained wetlands, all 31 years old, were studied. The stands were: (1) Scots pine, unfertilized; (2) Scots pine, fertilized; and (3) Norway spruce, fertilized. Amounts of nutrients (N, K, Ca, Mg, P, S, B, Fe, Mn, Zn, Cu) in above-ground biomass for all three stand types could be simulated precisely by a curvilinear regression model, with stand volume on bark as regressor. Net H + production of the fertilized pine was estimated to be 661 mol H + ha −1 year −1 from establishment to 31 years of age. The corresponding value for spruce was 1232 mol H + ha −1 year −1. Atmospheric inputs to the pine and spruce sites were 695 and 516 mol H + ha −1 year −1, respectively. Atmospheric input of N was 2.3 and 1.3 times the accumulation in the biomass of unfertilized and fertilized pine, whereas the value for spruce was 0.7. The corresponding ratios for S were 43, 19, and 11. 相似文献
17.
Management scenarios with rotation lengths of 20 and 30 years were developed for different site qualities (high, medium and low) under two different management options (high individual tree growth versus high stand growth) for teak ( Tectona grandis L.f.) in Costa Rica. The scenarios are based on data collected in different regions in Costa Rica, representing different site conditions, offering a variety of possible management options for high-quality teak yield. Three competition indices were used for modeling the competition and for the definition of intensities and the plantation age at thinning. The maximum site occupation (MSO) and the Reineke density index (RDI) provide conservative stand density management limits, resulting in the need to execute several thinning frequently. The competition factor (CF) matches the field observations and seems to be more appropriate for the growth characteristics of the species. Final stand densities varied between 120 and 447 trees ha−1, with mean diameter at breast height (dbh) of 24.9–47.8 cm, and mean total heights between 23.0 and 32.4 m, depending on rotation length and site quality. The mean annual increment of total volume (MAIVol) at the end of the rotation varied from 11.3 to 24.9 m3 ha−1 year−1, accumulating a total volume over rotation of 268–524 m3 ha−1. The most suitable scenario for teak plantations for high-quality sites is the 30-year-rotation scenario with five thinnings of intensities between 20 and 50% (of the standing trees) at the ages of 4, 8, 12, 18 and 24 years. After the sectioning of the merchantable stem in 4-m length logs, the merchantable volume varied between 145 and 386 m3 ha−1, with an estimated heartwood volume of 45–195 m3 ha−1, both depending on rotation length and site quality. 相似文献
18.
To assess the effects of clearcutting on snowshoe hare ( Lepus americanus) relative abundance, we surveyed pellets in 1 m 2 circular plots and, vegetation and browse surveys in 4.5 m 2 circular plots among four different aged clearcut (30, 20, 10, 5 years post-harvest) and mature forests (>150 years old) in central Labrador, Canada. Data were modelled at three grain sizes: transect (4400 m 2), plot (314 m 2) and subplot scales (4.5 m 2). Betula papyrifera, distance from mature forest edge, tree and herb cover as well as remotely sensed forest inventory data were used as predictors for hare pellets. We found pellet abundance was 5 and 37 times greater (new and old pellets, respectively) in clearcut stands 30 years old than the next highest in 20 year old cuts. There were few hare pellets in the remaining stand ages. B. papyrifera was the most proportionately used browse species and most important of our fine-detailed vegetation in predicting hare pellets. The coarse-detailed, forest inventory and topographic data better predicted hare pellets than the fine-detailed vegetation data. 相似文献
19.
Stand composition and structure utilizing stem analysis was studied in two hardwood stands in Vermont. In a mixed hardwood stand with some white pine and hemlock, a major entry of new trees in the main canopy seems associated with harvesting coincident with land exchange. More recent partial cuttings have promoted establishment of new seedlings or development of suppressed advanced-growth shade-tolerant beech, ( Fagus grandifolia, Ehrh.), hophornbeam ( Ostrya virginiana, (Mill) K. Koch), and striped maple ( Acer pensylvanicum, L.). Very few sugar maple and red oak seedlings and saplings are present. In a northern hardwood stand some red spruce (Picea rubens, Sarg.), that were 240 to 306 years old, became established before any known harvest, and exhibited release following harvests of the mid-1800s. This major harvest, coupled with the differential growth between spruce and hardwoods, and seed/seedling availability, resulted in a major change in stand composition. Trees now in the main canopy of sampled stands appear to have either been released or newly established following various harvests. Harvests have been of such frequency that natural disturbances seem insignificant. Many of the competitive understory species have become abundant following harvests of the 1960s and 1980s and may have been present as advanced growth and responded to the release. Following the harvest of 1981–1982, abundant yellow birch became established on skid trails. Elsewhere in the stand, yellow birch seedlings and saplings are only in great abundance in areas that were possibly sizable gaps following earlier harvesting. The dynamics of tree entry and growth in gaps of small or large size probably occur in a similar way in many other stands of the region. Though the sampling of this study is limited, there is no suggestion of continuous tree establishment at any particular location, the new age classes seem associated with either a gap or stand replacing disturbance attributed to harvesting. 相似文献
20.
We quantified structural features and the aboveground biomass of the deciduous conifer, Metasequoia glyptostroboides (Hu and Cheng) in six plantations in central Japan. In order to derive biomass estimates we dissected 14 M. glyptostroboides trees into three structural components (stem wood, branch wood and foliage) to develop allometric equations relating the mass of these components and of the whole tree to diameter at breast height (DBH). We found robust relationships at the branch and whole tree level that allow accurate prediction of component and whole tree biomass. Dominant tree height was similar within five older (>40 years) plantations (27–33 m) and shorter in a 20-year-old plantation (18 m). Average stem diameter varied from 12.8 cm in the youngest stand to greater than 35 cm in the oldest stand. Metasequoia have relatively compact crowns distributed over the top 30% of the tree although the youngest stand had the deepest crown relative to tree height (up to 38%). At the individual tree level in older stands, 87% of the aboveground biomass was allocated to the stem, 9% to branch wood and 4% to foliage. We found little difference in the relative distribution of above ground biomass among the stands with the exception of lower foliage biomass in larger diameter trees. Total aboveground biomass of the older stands varied twofold, ranging from a maximum of 450 Mg ha−1 in a 42-year-old stand to a minimum of 196 Mg ha−1 in a 48-year-old stand. Total above ground biomass of the 20-year-old stand was 176 Mg ha−1. 相似文献
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