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1.
Ediacaran assemblages immediately predate the Cambrian explosion of metazoans and should have played a crucial role in this radiation. Their wider relationships, however, have remained refractory and difficult to integrate with early metazoan phylogeny. Here, we describe a frondlike fossil, Stromatoveris (S. psygmoglena sp. nov.), from the Lower Cambrian Chengjiang Lagerst?tte (Yunnan, China) that is strikingly similar to Ediacaran vendobionts. The exquisite preservation reveals closely spaced branches, probably ciliated, that appear to represent precursors of the diagnostic comb rows of ctenophores. Therefore, this finding has important implications for the early evolution of this phylum and related diploblasts, some of which independently evolved a frondose habit.  相似文献   

2.
Halkieriids and wiwaxiids are cosmopolitan sclerite-bearing metazoans from the Lower and Middle Cambrian. Although they have similar scleritomes, their phylogenetic position is contested. A new scleritomous fossil from the Burgess Shale has the prominent anterior shell of the halkieriids but also bears wiwaxiid-like sclerites. This new fossil defines the monophyletic halwaxiids and indicates that they have a key place in early lophotrochozoan history.  相似文献   

3.
Soft-bodied marine faunas from the Lower and Middle Cambrian, exemplified by the Burgess Shale of British Columbia, are a key component in understanding the major adaptive radiations at the beginning of the Phanerozoic ("Cambrian explosion"). These faunas have a widespread distribution, and many taxa have pronounced longevity. Among the components appear to be survivors of the preceding Ediacaran assemblages and a suite of bizarre forms that give unexpected insights into morphological diversification. Microevolutionary processes, however, seem adequate to account for this radiation, and the macroevolutionary patterns that set the seal on Phanerozoic life are contingent on random extinctions. They weeded out the morphological spectrum and permitted rediversification among surviving clades. Although the predictability of which clades will play in successive acts of the Phanerozoic theater is low, at least the outlines of the underlying ecological plot are already clear from the opening of the drama.  相似文献   

4.
Phosphatized univalves, recovered from the Lower Cambrian ( approximately 530 million years old) Qiongzhusi Formation in southern China, are recognized as early instars belonging to bradoriid ostracodes whose later instars are bivalved. The univalved form is the primitive larval character for shell-secreting crustaceans, although most post-Cambrian ostracodes bypassed this developmental phase. The univalved-bivalved transition during early on-togeny represents an important evolutionary event in ostracodes, with implications for crustacean classification, and implies that the ostracode ancestor achieved this bivalved capacity before the appearance of mineralized skeletons during the "Cambrian explosion."  相似文献   

5.
In a problematical fauna discovered in Australia, 152 meters below diagnostic Lower Cambrian species, are fossilized remains belonging in the phylum Coelenterata. These jellyfish-like fossils were defined as a new class(Dipleurozoa) by Harrington and Moore because of their strong bilateral symmetry and differentiated extremities. The class was not reported elsewhere before many specimens were discovered recently in the Shawangunk (Tuscarora) formation at Delaware Water Gap, Pennsylvania. Three new species belonging to a new genus can be recognized, the indication being that the class ranges stratigraphically from the infra-Cambrian to at least the Lower Silurian, and geographically from Australia to eastern North America.  相似文献   

6.
Shu D  Morris SC  Zhang ZF  Liu JN  Han J  Chen L  Zhang XL  Yasui K  Li Y 《Science (New York, N.Y.)》2003,299(5611):1380-1384
Yunnanozoans are a distinctive clade of Lower Cambrian metazoans. Although widely accepted as deuterostomes, their exact placement within this superphylum is controversial. Here we describe a new species of Haikouella (H. jianshanensis) from the Chengjiang Lagerst?tte (Yunnan, China) with exceptional preservation of a number of features. These include external gills, which suggest that the origin of the pharyngeal clefts was independent of the gills. The diagnostic branchial arches of chordates may, therefore, be composite structures. No evidence was found for the chordate-like structures that have been described in other yunnanozoans. We propose that yunnanozoans are stem-group deuterostomes, allied to the vetulicolians.  相似文献   

7.
Fossil medusoids identified as Cyclomedusa davidi?, Beltanelliformis brunsae, and cf. Sekwia excentrica are reported from Late Precambrian strata in the Wernecke Mountains. They are representatives of the Ediacaran fauna, the oldest assemblage of cosmopolitan metazoans, and are only the third such occurrence in Canada. In addition, specimens broadly resembling the problematic structure Rugoinfractus ovruchensis, previously known only from the Precambrian of the Ukraine, are reported from the Lower Cambrian of the nearby Mackenzie Mountains.  相似文献   

8.
A fusiform, spirally coiled and pleated, free-living, heavily placate echinoderm with an expansible test has been discovered in the Lower Cambrian Olenellus zone of California. It is characterized by 10 "interambulacral" areas, a single principal endothecal ambulacrum with a short branch, and oral and apical regions at opposite poles. A new class, the Helicoplacoidea is proposed for the new genus Helicoplacus, with two new species, H. gilberti (type species) and H. curtisi.  相似文献   

9.
Phosphatic dermal fragments of Anatolepis, interpreted as a heterostracan fish (class Agnatha), have been discovered in the Deadwood Formation of Late Cambrian age in northeastern Wyoming. This discovery extends back the age of the earliest known vertebrate fossils by approximately 40 million years. Other occurrences of Anatolepis in North America, Greenland, and Spitzbergen show that these fish had a widespread geographic distribution in Late Cambrian and Early Ordovician marine environments.  相似文献   

10.
Fluvial sediments in paleovaileys cut in the Ashburton surface of the Davenport province of central Australia form terrace remnants that appear to retain their original depositional tops and have probably existed as subaerial landforms since their inception. Marine fossils in sediments conformable with the fluvial sediments near the southeast margin of the province give a Cambrian age for the terraces; the Ashburton surface forming the ridgetops between the paleovalleys is Cambrian or older.  相似文献   

11.
Cambrian arthropods are now well known, but there has been little agreement on how they contribute to an understanding of arthropod phylogeny. Fossils have either been lumped together as "trilobitomorphs" or, more recently, have been the subject of speculation invoking a multiple polyphyletic origin of arthropods. Cladistic analysis of characters of Cambrian and living representatives (excluding Uniramia) shows that trilobites and chelicerates are relatively advanced compared with "crustaceans," and there are doubts whether the latter constitute a national group. An undue emphasis on singular autapomorphies of problematic fossils has obscured these relationships in the past. "Trilobitomorphs" were simply an artificial taxon based on shared primitive characters. The arthropods that evolved during the Cambrian radiation show no more apparent morphological diversity than do the living groups. The evidence of wellpreserved problematica is critical to understanding the nature of this radiation and the affinities of the groups that remain today.  相似文献   

12.
Small bilaterian fossils from 40 to 55 million years before the cambrian   总被引:1,自引:0,他引:1  
Ten phosphatized specimens of a small (<180 micrometers) animal displaying clear bilaterian features have been recovered from the Doushantuo Formation, China, dating from 40 to 55 million years before the Cambrian. Seen in sections, this animal (Vernanimalcula guizhouena gen. et sp. nov.) had paired coeloms extending the length of the gut; paired external pits that could be sense organs; bilateral, anterior-posterior organization; a ventrally directed anterior mouth with thick walled pharynx; and a triploblastic structure. The structural complexity is that of an adult rather than a larval form. These fossils provide the first evidence confirming the phylogenetic inference that Bilateria arose well before the Cambrian.  相似文献   

13.
The shallow-water limestones and dolostones of the Conococheague Limestone (Upper Cambrian) of western Maryland contain large amounts of authigenic potassium feldspar. The presence of halite daughter crystals in breached fluid inclusions, low whole-rock ratios of chlorine to bromine, and thermochemical data suggest that the potassium feldspar formed at low temperature by the reaction of connate brines with intercalated siliciclastic debris. Analyses of argon age spectra indicate that the authigenic feldspar probably formed during Late Pennsylvanian to Early Permian time. These results may indicate mobilization and migration of connate brines brought about by Alleghanian folding. The widespread occurrence of authigenic potassium feldspar in Cambrian and Ordovician carbonate rocks throughout the Appalachians suggests that this may have occurred throughout the entire basin.  相似文献   

14.
The Chinese Early Cambrian Chengjiang fauna includes three different anomalocaridids, a globally spread, extinct marine group including the largest known Cambrian animals. Anomalocaridids were active predators, and their presence implies that a complex ecosystem appeared abruptly in the earliest Phanerozoic. Complete specimens display several sets of characters shared only with some other exclusively Cambrian forms. This evidence indicates that anomalocaridids, Opabinia, and Kerygmachela form a monophyletic clade. Certain features indicate arthropod affinities of the lade, and for this group an unnamed (sub)phylum-level taxon within an arthropod (super)phylum is proposed.  相似文献   

15.
Diverse bilaterian clades emerged apparently within a few million years during the early Cambrian, and various environmental, developmental, and ecological causes have been proposed to explain this abrupt appearance. A compilation of the patterns of fossil and molecular diversification, comparative developmental data, and information on ecological feeding strategies indicate that the major animal clades diverged many tens of millions of years before their first appearance in the fossil record, demonstrating a macroevolutionary lag between the establishment of their developmental toolkits during the Cryogenian [(850 to 635 million years ago (Ma)], and the later ecological success of metazoans during the Ediacaran (635 to 541 Ma) and Cambrian (541 to 488 Ma) periods. We argue that this diversification involved new forms of developmental regulation, as well as innovations in networks of ecological interaction within the context of permissive environmental circumstances.  相似文献   

16.
The fossil Pteridinium, a distinctive component of a worldwide early metazoan (Ediacaran)assemblage, is provisionally recorded from probable early Cambrian strata in eastern California. In context with other evidence, this finding implies a Cambrian age for the Ediacaran fauna and approximate coincidence of limits between Phanerozoic-Cryptozoic, Paleozoic-Precambrian, and Cambrian-Precambrian.  相似文献   

17.
Conodont pearls?     
Conodonts are zoologically enigmatic, toothlike phosphatic microfossils occurring in marine sedimentary rocks ranging in age from Cambrian to Triassic. Dimpled spheres of less than 1 millimeter in diameter are sporadic associates of conodonts and have identical chemical composition and microstructure. Mineralogy, morphology, and occurrence of these spheres suggest that they are pearls secreted by the conodont-bearing animal.  相似文献   

18.
The oldest known "carpoids" of the echinoderm subphylum Homalozoa have been discovered in lower Middle Cambrian rocks of northern Utah. They were free-living benthonic animals characterized by a unique ctenoid feeding apparatus and flattened flexible theca with near bilateral symmetry. A new class, Ctenocystoidea, is proposed to include the new genus and new species, Ctenocystis utahensis.  相似文献   

19.
Agglutinated foraminifera have been recovered from siltstones in the Walidiala Valley, Taoudeni Basin, West Africa. Associated faunas suggest an Early Cambrian age for these strata. These now earliest known unequivocal foraminifera help constrain hypotheses concerning the origin of skeletalization at the beginning of the Phanerozoic.  相似文献   

20.
Fossiliferous Cambrian, Ordovician, Mississippian, and Pennsylvanian rocks, never before found in southern Mexico, have been discovered in the Nochixtlán region. Superjacent unfossiliferous sedimentary rocks may be Permian in age. Early Paleozoic and late Paleozoic intervals of marine sedimentation were bounded by intervals of positive tectonism and erosion.  相似文献   

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