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1.
Abstract

The success of natural regeneration on patch clear-cuts in Norway spruce stands in Southern Finland was examined in terms of stocking and tree height. The experiment was established in mature spruce stands in the submesic Myrtillus (MT) site type. In each of the eight study sites, three plots were treated with clear-cutting and planting, clear-cutting with partial tree retention and planting, and patch clear-cutting without site preparation, respectively. In three of the study sites, one plot with patch clear-cutting and site preparation was established. Each plot was 1 ha, out of which three patches of 40×40 m were clear-cut in the patch treatments. 10–11 years after cutting, the patches had on average 1316 crop trees ha–1, of which 91% were spruces. Some 27% of the stands were up to the target stocking level (≥1600 ha?1), and 36% were at least satisfactory (≥1300 ha?1). Site preparation did not yield greater stocking levels on patches, but that result is ambiguous due to a difference in initial stocking. The average spruce tree height in the patches (0.76 m) was much smaller than in the case of clear-cutting and planting (2.42 m). In conclusion, the patches had been restocked tolerably well with spruce and birch for practical purposes during the 10- to 11-year period, but the regeneration process had been very slow compared to clear-cutting and planting.  相似文献   

2.
Johansson  Tord 《New Forests》2004,27(1):13-24
Plantings with pendula (Betula pendula Roth) and pubescent (Betula pubescens Ehrh.) birches were studied on five localities (Lat. 57–60 ° N) in Sweden. Seedlings were planted in 1988 or 1989. The experiment contained 50 plants per parcel and five replications per treatment. Among treatments harrowed area and birches covered with tree shelter were included. In spring 1993 five plants growing on harrowed area, with and without tree shelter, respectively were randomly chosen on each parcel. A total of 25 birches per treatment, species and locality were used. The trial was replicated in 1994 and 1995. The plant height was estimated and diameter at 20 and 60% of tree height was measured each year for 4 yr. The stem taper was calculated by the quotient D60/D20. The stem taper for birches without tree shelter was 0.5 for both species on all localities. The quotients for birches, which have grown in tree shelter, were 0.6–0.75 the first years of the trial. Then the quotients decreased and 2–3 years later they were on the same level as for birches planted without tree shelters. The results of the studies indicate that tree shelters should be removed two to three years after the plant has reached the top of the shelter. The height of the tree shelter might be shortened to 0.8–1.0 m to stabilise the plant earlier than with the conventional shelter height of 1.2 m.  相似文献   

3.
Abstract

This study compares the results of the prediction of crown height characteristics using airborne laser scanner (ALS) data and intensive field measurements in boreal forests. The data consisted of 31 sample plots located in Kalkkinen, southern Finland. Crown height models were constructed at both the tree and plot level. Scots pine, Norway spruce and birches were used. The models included independent variables of tree levels, such as tree height, crown area and independent plot-level variables, i.e. canopy height and density quantiles and proportion of vegetation hits. Field measurement-based models used tree height and diameter at breast height as the independent tree-level variables, whereas basal area, mean diameter and height were used as the plot-level variables. The results indicated that the ALS-based crown height models were more accurate than the field measurement-based models when plot-level information was used as independent variables. However, the field measurement-based tree-level models for Scots pine and Norway spruce were more accurate than the ALS-based models. Even so, the accuracy of the different models was very similar and the study data set was quite small. The results of this study can be used for different tree growth studies and for the assessment of tree stock quality in boreal forests.  相似文献   

4.
The single-tree selection system is an important option for management of Norway spruce (Picea abies (L.) Karst.) and silver fir (Abies alba Mill.) forests because it provides continuous cover, requires low investments for tending, and promotes natural regeneration as well as high stand resistance and elasticity. It is often regarded as a very conservative system that usually results in only minor spatiotemporal changes in forest structure and composition. We studied management history, structural changes, regeneration dynamics, and light climate of a traditional single-tree farmer selection silver fir-Norway spruce forest (site typology Bazzanio-Abietetum). Stand structure was analyzed on five 0.25 ha permanent plots in 1994, 2001, and 2008. Regeneration density and height growth, forest floor vegetation, and light climate were also assessed on 1.5 × 1.5 m regeneration subplots in 2001 and 2008. Tree cores extracted from dominant trees from both species in two plots were used for reconstructing stand history and age structure of the canopy layer. We documented the forest response to three types of selection management regimes: excessive, normal, and conservative. Excessive management with harvest intensity significantly above the increment was documented until the late 1950s, including two peaks of heavy fellings (diameter limit cut) in the 1880s and 1930s, which favoured establishment of Norway spruce and released regeneration. The period that followed was characterized by normal selection management, but was nevertheless marked by a decline of silver fir as a result of air pollution and several droughts. This led to sanitary fellings that were carried out from the late 1970s to the early 1990s. In the last two decades conservative management followed, which led to suppression and decline of regeneration, especially of Norway spruce, and loss of selection structure. Although we recorded lower regeneration potential of silver fir compared with Norway spruce within the seedling category, silver fir outcompeted Norway spruce within the small-sized tree category (1 cm < dbh ? 10 cm) because of its superior height growth in low light levels (diffuse light <6%) and occupied a greater share of the canopy. Nevertheless, we anticipate that over the long-term the low light regime will also cause regeneration decline of silver fir and broadleaves. Our research revealed significant structural changes in a single-tree farmer selection forest during the last 150 years. These were a result of variable management regime and environment. A farmer single-tree selection system could better mimic the natural disturbance regime if spatiotemporal combinations of diverse felling regimes would be used.  相似文献   

5.
Natural regeneration measurements are the main silvicultural objective in overaged protective forests of the Bavarian Limestone Alps. While manifold problems with these stands, especially the impact of browsing, are widely recognised, the regeneration niches of Norway spruce (Picea abies [L.] Karst.) are insufficiently known. The purpose of this study was to determine favourable combinations of site factors for the development of spruce in small, unfenced canopy gaps, located on Aposerido-Fagetum caricetosum albae forest sites. We recorded the occurrence of spruce saplings (as dependent variable) and of six site factors (as independent variables) on 480 0.5 m2-subplots. In addition, we estimated the coverage of six acid adapted plant species to determine correlations with the humus depth. A binary logistic regression analysis was used to predict the probability of the occurrence of a spruce sapling in dependency of the different site factors. Supported by other studies, we assumed that the supply of solar radiation was adequate for the sufficient regeneration of spruce within the canopy gaps. Other site factors significantly determined the regeneration niches of spruce saplings. More spruce saplings were found near hindrances and on rough surfaces than would be expected from a random occurrence of saplings. These microsite types may have characteristics, especially protection against snow gliding that promotes spruce establishment. A calculated “hindrance index”, which accounted for the number, and the distance of surrounding hindrances might be a good specific value to describe the influence of hindrances on steep slopes. The sapling establishment decreased on thin humus layers. Our assumption for the sites was that thick organic layers might represent a good seedbed for spruce. Decayed dead wood was scarce, but was exceedingly favoured by spruce saplings. Results obtained suggest that the natural regeneration establishment of spruce on steep slopes can be successfully influenced by site factors which inhibit the influence of snow gliding. According to a “positive microsite” concept, we recommend for artificial regeneration measurements with spruce, microsites close to hindrances (e.g. stumps, downed trees) and Vaccinium myrtillus as a predictor for thick, acid humus layers.  相似文献   

6.
In single‐tree selection, trees removed by harvest or lost through mortality are replaced by ingrowth from the seedling/sapling bank. Because the level of ingrowth is governed not only by the recruitment rate of new seedlings, but also by mortality and growth rates within the seedling/sapling stratum, knowledge of how these processes are related to the tree stratum is important for successful application of single‐tree selection. Therefore, Norway spruce (Picea abies (L.) Karst.) regeneration (0.1 m ≤ height ≤ 2.0 m) was measured on seventy 100 m2 circular subplots at each of two sites in central and northern Sweden. Both sites had previously been selectively logged, but the time elapsed since the last harvest was at least 30 yrs. Basal area of trees of at least 2 m height within three different radii from the subplot centres was measured. Measurements on regeneration included height and leader length. Influence of local stand basal area on density and growth of regeneration was analysed by ordinary least‐squares linear regression for each of four height intervals. Regeneration was significantly aggregated at both sites, but number of Norway spruce seedlings and saplings per circular subplot was not correlated to local stand basal area. Out of 36 regressions, only eight showed a significantly (p < 0.05) negative correlation between seedling and sapling height growth and local stand basal area. In five of these cases, removal of one point (subplot) made p > 0.05. The results suggest that density and growth of Norway spruce regeneration in selectively logged uneven‐aged Norway spruce stands is affected more by ground conditions than by local stand basal area.  相似文献   

7.
A bark beetle (Ips typographus) infestation caused the death of almost all Norway spruce (Picea abies) trees in a mountain forest in the Swiss Alps. We developed a tree regeneration model, ‘RegSnag’ (=REGeneration in a SNAG stand), to project the future amount and height of tree regeneration in these snag stands. The model combines a height-class structured tree module with a microsite-based module of snag decay and ground-vegetation succession. Microsite-specific rates of germination, mortality and height growth were modelled for four tree species (Picea abies, Sorbus aucuparia, Acer pseudoplatanus and Betula pendula) in eight height classes (from seedlings to saplings 5 m tall) and on 26 microsite types (e.g. moss, grass). Model tests with independent field data from 8 years after the Picea die-back demonstrated that microsites had a considerable effect on the development of tree regeneration on both the montane and the subalpine level. With microsite-specific parameters, the height and frequency of Picea in each microsite could be simulated more accurately than without considering microsite effects (e.g. bias of 8 vs. 119 saplings ha−1 on the montane level). Results of simulations 40 years into the future suggest that about 330–930 Picea saplings per ha out of those that germinated in 1994 and 1996 will reach a height of 5 m within 30–35 years after Picea die-back. This is due to differences in seed inflow and browsing intensities. Picea and not Betula or Sorbus trees will replace the current herbaceous vegetation in these snag stands.  相似文献   

8.
Growth data were collected from 157 Norway spruce (Picea abies (L.) Karst.) stands planted on farm land in Sweden from 55 to 66° N. The mean age of the stands was 41 years (range, 25–91), the mean stand density 1 640 stems ha–1 (range, 400–3 722), and the mean diameter at breast height (outside bark) 25 cm (range, 12–48). The height growth was measured in 56 stands during the initial five years after plantation and followed systematically until the stands were 30 years old. Early height growth for spruces growing on clay soils was lower than for trees growing on sand, till and peat soils. The height increment for 5-year-old spruces predicted the height increment for mature spruces (30–50 years old).Site index curves were constructed for total age. Curves for H40 (dominant height at 40 years total age) were made for both northern (50 stands, 61–66° N) and southern (107 stands, 55–61° N) Sweden. Site index curves for H50 at total age are presented for Sweden as a whole (Lat. 55–66° N) and southern Sweden only. Curves based on breast height age were also constructed for H40 and H50. Curves fitted for H40 at breast height were well in accordance with the curves presented in the past for spruce on forest lands by Nordic studies. The curves from the present study have slower growth for young spruces than curves for forest land. For 40–90-year-old spruce stands, curves from the present study indicate taller heights than from forest land curves.  相似文献   

9.
Spatial distribution of Heterobasidion genets over a period of ca 50 years in two successive generations of Norway spruce (Picea abies) was unravelled. The genets were first identified in 1993 in a naturally regenerated 43‐year‐old spruce stand that had been thinned the previous winter. Heterobasidion parviporum was found in 17.5% of the old stumps of the previous spruce generation. Nine genets were identified on the study plot; seven of them were present in old stumps of the previous tree generation and two only in the new spruce generation. Eighteen spruce trees of the new generation were infected, 15 of them by vegetative growth of genets originating from the old stumps. The study plot was investigated again in 2005. No new genets had been established after thinning, and three old genets had died out. The remaining genets had infected five new trees, most likely from the thinning stumps of diseased trees. At the age of 56 years, 16.1% of the residual spruces were infected by Heterobasidion. The results of this study suggest that if spore infection to stumps of spruce can be prevented, the decay frequency caused by H. parviporum will not necessarily increase in successive generations.  相似文献   

10.
Growth of regenerating trees in different light environments was studied for the mountainous, mixed-species forests in the Carpathian Mountains of Romania. The primary species in these mixtures were silver fir (Abies alba Mill.), European beech (Fagus sylvatica L.) and Norway spruce (Picea abies (L.) Karst). Seedlings/saplings of these species were selected and measured in different stands from two different geographical locations. Regenerating trees were measured for height and diameter growth during the summer of 2002. For each seedling/sapling, percentage of above canopy light (PACL) and stand basal area (BA) were used to assess available and occupied growing space respectively. Regeneration growth was compared against these two variables and regression relationships were developed. Using these models, we predicted the dynamics of regeneration as both growth and species composition. Our results showed that in low-light environments (PACL<20–35%; BA>30 m2/ha), shade tolerant fir and beech clearly outcompeted the spruce. Therefore, in dense stands, spruce could be eliminated by the shade tolerant species. For intermediate levels of cover (PACL=35–70%; BA=15–35 m2/ha) the spruce grew at comparable rates as the beech and fir. All three species showed similar growth rates in open conditions (PACL>80–90%; BA<15–20 m2/ha) with the spruce having a tendency to outgrow the others. However, in terms of establishment, such conditions favor spruce and inhibit fir and beech.  相似文献   

11.
Prévost  Marcel  Charette  Lise 《New Forests》2019,50(4):677-698

Vast areas of hardwood and mixedwood forests of eastern North America have been high-graded in the past and need silvicultural treatments to increase their value and productivity. To rehabilitate a high-graded temperate mixedwood stand, in Quebec, Canada, we used a split–split-split plot design with three replicates to assess different seed-tree and strip cutting systems, in combination with scarification and planting. The experiment consisted of three regeneration cuts in main plots: 10 seed-trees/ha, 40 seed-trees/ha and a 40-m wide strip clearcut (0 seed-tree/ha) with 60 seed-trees/ha in leave strips, thereby resulting in four levels of tree retention, and all included understory brushing. We applied two types of scarification (patch scarification or disk-trenching) to subplots, two regeneration modes (natural regeneration or planting with white spruce [Picea glauca]) to sub-subplots and two mechanical release treatments (softwood or mixedwood production) to planted sub–sub-subplots. Density of seed-trees did not affect the natural regeneration dynamics after 5 years, but disk-trenching was more efficient for the establishment of yellow birch (Betula alleghaniensis) and sugar maple (Acer saccharum). Few seed-trees of desirable white spruce were present and most died standing, confirming the importance of supplemental planting. Height growth of planted seedlings was 15% higher in the 0 and 10 (26–27 cm/year) than in the 40 and 60 (23 cm/year) seed-trees/ha treatments, and release doubled mean height growth (33.1 vs. 16.6 cm/year). Despite intensive site preparation, pre-established beaked hazel (Corylus cornuta) and mountain maple (Acer spicatum) were present at high densities in the regeneration stratum. Controlling this recalcitrant layer might be the greatest challenge for rehabilitating degraded stands of the mixedwood forest, especially since the use of herbicides is prohibited on Quebec’s public lands.

  相似文献   

12.
Natural regeneration of windthrow areas is an important issue when planning forestry measures after forest disturbances. Seedling recruitment was investigated in storm-damaged hemiboreal mixed forests in eastern Estonia. The establishment and growth of seedlings from natural regeneration was registered for tree species in soil pits and in mounds of uprooted trees in stands that were either heavily or moderately damaged. Seedling growth is expected to be better in large but shallow soil pits created by uprooted Norway spruce [Picea abies (L.) Karst.] and poorer in small but deep pits created by the hardwoods in the area, silver birch (Betula pendula Roth.) and European aspen (Populus tremula L.). The most abundant regenerating species was birch. Pits hosted larger seedling numbers than mounds, due to soil instability in mounds. Rowan (Sorbus aucuparia L.) showed significantly faster growth than the other seedling species. Norway spruce pits were preferred to pits of other species by both birch and spruce seedlings. Black alder [Alnus glutinosa (L.) J. Gaertn.] did not show a preference for pits of a certain species of uprooted tree. Both spruce and rowan preferred hardwood mounds over spruce mounds. Storm severity also affected species composition: birch predominantly occurred on pits and mounds in heavily disturbed areas, while spruce was more abundant in the moderately damaged areas. The effects of advance regeneration and surrounding stands on seedling microsite preferences should be considered in future research and subsequent management recommendations.  相似文献   

13.
In the absence of reliable and representative data on the frequency of seed years, seed amounts, germination and survival of seeds, the data of the observation period 1992–1996 of the permanent national forest inventory of Austria are used to develop a model describing the probability for the occurrence, density and height of Norway spruce (Picea abies (L.) Karst) natural regeneration in the forests of Austria. The forest inventory provided data on natural regeneration (saplings between 10 and 130 cm height) on clusters within a 3.89 km square sampling grid. A logistic equation is used to predict the probability for natural regeneration occurrence. Input parameters for this equation are the variables that describe the site, with slope and azimuth as continuous variables, and growth districts and vegetation types as discrete variables; the crown competition factor describes density and the quadratic mean diameter describes the stage of development of the stand. The same equation type is used to predict the probability for the occurrence of Norway spruce, conditional on the occurrence of some regeneration. An additional variable in this species specific model is a dummy variable which is set to 1 if Norway spruce occurs in the overstory and otherwise zero. Additional site variables entering this model are elevation and vegetation types characterizing soil fertility and moisture. Because the density and height of spruce regeneration depends on the stand's susceptibility to browsing, the probability for browsing is also modelled as a logistic equation depending on elevation, vegetation type, and stand density. Finally the probability distribution for height and density of the regeneration is described by two bivariate Weibull-distributions, each one describing browsed and unbrowsed Norway spruce regeneration respectively.  相似文献   

14.
Average tree height and basal area growth for Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) in Sweden were studied as functions of species, age, stand density, location and year of inventory in the period 1953–1992, on the basis of sample tree data from the National Forest Inventory. A highly significant annual increase of both height and basal area growth was found, of the magnitude 0.5–0.8%, during the 40 yr period. Possible reasons for the trend are discussed. The altered way of cutting in the early 1950s, from selective cutting to clear felling and thinning from below, has had a large impact. Also, improved regeneration methods, nitrogen fertilization and ditching have increased growth. The increasing atmospheric deposition of nitrogen is another possible factor.  相似文献   

15.
Abstract

The effect of different cleaning treatments on moose (Alces alces L.) damage to Scots pines (Pinus sylvestris L.) was examined using a generalized linear mixed model (GLMM) with a binomial response (logit-link function). The treatments were the following: no cleaning, point-cleaning of broadleaves (mainly birch, Betula pendula Roth and B. pubescens Ehrh.) within a radius of 1 m from the pine, point-cleaning delayed 2 years, topping of competing broadleaves (main stem cut at a height of 1.3 m) and total cleaning of broadleaves. A randomized complete block design with three replicates was established in three planted pine stands (mean height of pines 1.5 m). Measurements taken four growing seasons later (mean height of pines 3.5 m) showed that all cleaning treatments reduced moose damage, although the difference was not significant (p=0.078) in the case of topping. The proportion of damaged pines was predicted as a function of the characteristics of pines and broadleaves in two untreated, planted pine stands with a height of 3 m. Within these untreated stands the number of damaged pines increased in plots with a higher number of broadleaves and taller broadleaves. In conclusion, the results show that on relatively fertile growing sites of pine cleaning treatments reduced moose damage on pine. It is recommended that cleaning is performed before broadleaved trees occur as overgrowth above pines.  相似文献   

16.
The individual tree growth in stands of mixed Norway spruce (Picea abies (L.) Karst.) and birch (Betula pendula Roth & Betula pubescens Ehrh.) is estimated using basal area and height growth functions for each species separately. The individual tree growth models are distant dependent and the number, size and proximity of neighbours are expressed as size‐ratio competition indices. The competition indices were calculated using a basal area factor gauge to define competitors. The tree growth functions are based on data from nine mixed stands of Norway spruce and birch. The recursive multivariate regression approach is used. The growth functions have standard deviation about the function/standard deviation about the mean (sf/sm) values between 31 and 61% and the evaluations made with root mean square error (rmse) give estimates which vary between 8 and 45 % of the observed mean value. These values are comparable with the precision reported in other studies. In the present investigation the distant dependent indices are important independent variables. It seems suitable to describe the change in growth conditions from retarded to released growth by means of size ratio competition indices. For birch, a positive effect on growth is obtained the more the total competition consists of Norway spruce. For Norway spruce a negative effect on growth is obtained the more the total competition consists of Norway spruce. The lower competitors have a positive effect on the growth of the spruce trees. The interpretation should be that it is better to have a small competitor than a larger one, not that small competitors as such have a positive effect on growth of Norway spruce.  相似文献   

17.
闽楠天然次生林自然更新的影响因子研究   总被引:3,自引:0,他引:3       下载免费PDF全文
[目的]分析闽楠天然次生林自然更新与环境因子之间的关系,为其科学经营及保护提供参考依据。[方法]以江西省吉安市闽楠天然次生林为研究对象,通过标准地调查获取数据,运用多元数量化模型I建立闽楠幼树幼苗重要值与环境因子的关系模型,采用单因素方差分析单个环境因子对闽楠幼树幼苗更新的影响。[结果]研究表明:密度(闽楠下种母树株数、郁闭度、株数密度)、坡位、腐殖质层厚度、坡向、林下植被盖度、凋落物层厚度是影响闽楠天然次生林自然更新的主要因素,偏相关系数在0.325 7 0.715 7之间,t检验结果为极显著或显著;模型复相关系数为0.966,F检验结果为极显著(F=30.96~(**))。[结论]闽楠下种母树株数对其幼树幼苗的更新起着最主要的作用,而郁闭度与株数密度过高或过低、凋落物层越厚、腐殖质层越薄、林下植被盖度越大均不利于其自然更新,同时,半阴坡、下坡位条件下的闽楠幼树幼苗的更新好于其它坡向与坡位。为促进闽楠幼树幼苗的自然更新,林分中闽楠下种母树应保留200株·hm~(-2)以上,以及对郁闭度、林下植被盖度及凋落物层厚度等实施相应的调控措施。  相似文献   

18.
《Southern Forests》2013,75(2):77-88
Estimating tree volume and biomass constitutes an essential part of the forest resources assessment and the evaluation of the climate change mitigation potential of forests through biomass accumulation and carbon sequestration. This research article provides stem volume and biomass equations applicable to five tree species, namely Afzelia africana Sm. (Caesalpiniaceae), Anogeissus leiocarpa (DC.) Guill. and Perr. (Combretaceae), Ceiba pentandra (L.) Gaertn. (Bombacaceae), Dialium guineense Willd. (Caesalpiniaceae), Diospyros mespiliformis Hochst. ex A.DC. (Ebenaceae) in natural protected tropical forests and, in addition, Tectona grandis L.f. (Verbenaceae) in plantations. In addition to the tree species specific equations, basic wood density, as well as carbon, nitrogen, organic matter and ash content were determined for these tree species in tropical conditions in West Africa. One hundred and sixty-two sample trees were measured through non-destructive sampling and analysed for volume and biomass. Stem biomass and stem volume were modelled as a function of diameter (at breast height; Dbh) and stem height (height to the crown base). Logarithmic models are presented that utilise Dbh and height data to predict tree component biomass and stem volumes. Alternative models are given that afford prediction based on Dbh data alone, assuming height data to be unavailable. Models that include height are preferred, having better predictive capabilities. Ranges in carbon, nitrogen and ash contents are given as well. The successful development of predictive models through the use of non-destructive methods in this study provide valuable data and tools for use in determining the contribution of these major African rainforest tree species to global carbon stocks, while ensuring the preservation of this valued African resource. This study needs to be expanded to further regions and tree species to complete a full inventory of all tree species, emphasising the relevance of African trees to carbon stocks at a global scale.  相似文献   

19.
Abstract

An experiment was established in 1978 in two Norway spruce [Picea abies (L.) Karst.] plantations in southern Sweden to study yield after mortality in patches with and without supplementarily planted (SP) seedlings. Gaps of different sizes were created by removing the originally planted seedlings. The gaps were either left unplanted or a supplementary planting was performed with one of four species [Norway spruce, Scots pine (Pinus sylvestris L.), lodgepole pine (Pinus contorta Dougl.) or hybrid larch (Larix deciduas Mill×L. Leptolepis Gord.)] 2 (at Knäred) or 6 years (at Ullasjö) after the original plantation. In 2002, most of the SP Scots pine, lodgepole pine and hybrid larch seedlings were dead or severely damaged by roe deer and moose. Survival was high among SP Norway spruces, but they had slower growth than the originally planted spruces. Growth was lower at Ullasjö than at Knäred. In Ullasjö, growth was lower in small gaps than in large gaps. Trees in original regeneration in areas surrounding unplanted gaps were larger than trees surrounding gaps with SP seedlings, which in turn were larger than originally planted trees in plots without gaps. In conclusion, because the original plantation surrounding unplanted gaps used a large part of the open space and growth of SP seedlings was slow, supplementary planting resulted in an insignificant growth increase. However, supplementary planting may increase the timber quality of trees surrounding the gaps, although this effect remains to be quantified.  相似文献   

20.
It is believed that European beech (Fagus sylvatica L.) will increase its competitive ability at its northern range margin in Scandinavia due to climate change. In mixed old-growth forests of beech and Norway spruce (Picea abies (L.) Karst.) at Siggaboda nature reserve (southern Sweden), stand structure characteristics were sequentially recorded in the years 2004, 2005 and 2007 as well as growth in stem diameter using tree-coring analyses. Using these measurements, we studied the effects on stand dynamics of an extreme storm event (2005 “Gudrun” hurricane), drought and heat (mid-summer 2006, spring 2007) and subsequent bark beetle attacks on spruce (growing season 2007), overlaid with warming tendencies. The storm, which caused disastrous damage in many stands nearby, had comparatively little impact on the structure of the spruce–beech stand. All together, only 32 trees (19 spruces, 10 beeches, 3 other species) per hectare were thrown or broken mainly in the leeward direction (NE) or impacted by secondary damage by uprooted neighbour trees; this represents 7% of the total tree number and 11% of the growing stock. Diameter and height structure did not change significantly. However, the 2006 drought and the 2007 attack of biotic agents changed the stand structure and composition strongly due to the death of about 19% of the dominating older spruce trees that accounted for 35% of total stand volume. This resulted in a considerable increase in beech’s contribution to stem number (4% increase) and wood volume of the living stand (7% increase). A comparison of diameter growth of beech and spruce during the periods 1894–1949 and 1950–2005 showed a distinct decrease in growth superiority of spruce during the last 50 years. These results support the idea of a northward migration of European beech as a nemoral tree species in Sweden, due to a higher tolerance to the abiotic and biotic threats accompanying climate change and an increased competitive ability compared to boreal tree species Norway spruce.  相似文献   

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